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This description covers characteristics that may be relevant to fire ecology and is not meant for identification. Keys for identification are available (e.g., [35,41,74,105,155,160]).
Aboveground description: Crownvetch is a mostly hairless, perennial forb with weak spreading stems from a short branching caudex [8,41,105,109,155]. Crownvetch lacks tendrils that are characteristic of true vetches (Vicia spp.) [139]. In flower, crownvetch plants may be 3 or more feet (1 m) tall, but plants are generally just 1 foot (0.3 m) tall in their vegetative state [8,162]. Crownvetch produces trailing stems that may reach 6 feet (2 m) long [139] and often appears as a heavy mat of stems [3]. Crownvetch is often described as "long-lived" [56,162], but precise life expectancy was rarely reported. One report indicates that crownvetch plants live up to 20 years [31]. Another report indicates that crownvetch has survived in the same location for over 50 years [3].
Crownvetch has odd-pinnate compound leaves that are 2 to 6 inches (4-16 cm) long. Leaves have 7 to 25 leaflets that measure 0.4 to 1 inch (1-3 cm) long [8,49,67,133]. In the winter, crownvetch's basal, and sometimes upper, leaves remain green [129]. Crownvetch produces pea-like flowers in a spreading umbel that resembles a crown [49,155]. Individual flowers are 9 to 12 mm long, have 10 stamens, and are white-pink to rose colored [8,109,160]. Umbels contain 6 to 25 flowers [8]. Pubescence, flower color, and leaflet size and shape can vary considerably [41].
Crownvetch generally produces seeds in a slender, linear, jointed pod [49,155]. Pods may reach 2 inches (6 cm) long and have up to 12 joints. Each joint contains a single seed [8,35,105,133]. Pods are described as indehiscent by Stubbendieck and Conard [133], but Stevens and Monsen report that pods break open when dry [129]. Crownvetch seeds are cylindrical and measure 3 to 4 mm by 1 to 1.2 mm [7,133].
For descriptions of the morphological similarities and differences between the 3 most common crownvetch cultivars: 'Penngift', 'Chemung', and 'Emerald', see [92].
Belowground description: Anecdotal reports about crownvetch's underground growth are common, but details and measurements are generally lacking. Crownvetch produces both roots and rhizomes. The taproot is deep and multibranched (review by [120]). Rhizomes are strong and fleshy and may grow to 10 feet (3 m) or longer (reviews by [119,120,139]). From the lower surface, rhizomes produce roots and from the upper surface, shoots (review by [22]). In a cemetery near Cincinnati, Ohio, crownvetch roots occurred more than 6.5 feet (2 m) deep in heavy clay soils [39]. On cut slopes with moist, loose soil and "ample" phosphorus in West Virginia, an 8-month-old crownvetch plant produced roots over 6 feet (2 m) long. On a steep, dry slope, 47- to 77-day-old crownvetch seedlings had 3- to 6-inch-long (8-16 cm) taproots. Within 100 days, most grass seedlings on the site had died, while crownvetch established well [100].
Crownvetch is a nonnative species found throughout the contiguous United States and southern Canada [8,96,139,160,169]. Crownvetch also occurs in Hawaii [155] but does not occur in Alaska [148]. Most reports indicate that crownvetch is native to Eurasia [8,96,160,169], and it may also be native to northern Africa [46,139]. Although crownvetch is widely distributed in North America, it is most common in areas near sites where it was planted [155]. In the northern and central parts of the United States, crownvetch is most common [92] and can be invasive (see Impacts). For a map of crownvetch's North American distribution, see Plants Database.
Introduction: In the reviewed literature, the earliest report of crownvetch in the United States occurred in 1869 in New York. As of 1872, this population near the Hudson River was described as "thoroughly established and naturalized, spreading in open spaces of the wood" [163]. In 1874, crownvetch was "well naturalized" in fields in the Pine Plains region of Dutchess County, New York [57]. Reviews report that crownvetch was available commercially in the United States by 1890 [92,120].
After 1935, when crownvetch plants were found covering over 10 acres (4 ha) of very poor quality soils in Pennsylvania, research into crownvetch as an erosion control and revegetation plant began. By the 1950s, crownvetch was planted extensively in North America [139]. The crownvetch population in Pennsylvania developed from a single plant that emerged in an alfalfa field planted in 1905 [39]. Crownvetch persisted in soils with "fertility levels so low that only poverty grass would normally survive" [162]. Crownvetch was commonly used to revegetate roadside cuts, mining sites, and railroad embankments. It was also planted as an ornamental and used as a cover crop or as green fertilizer [8,35,67,109,119,132,159,160]. By the 1960s, crownvetch was abundant in the eastern United States [60]. Although widely planted to control erosion, in some areas crownvetch has only camouflaged erosion (Grover and Harper-Lore 2001 personal communications cited in [139], review by [147]). For more information, see Other Uses.
On abandoned mine sites, crownvetch was commonly recommended for revegetation [152], and throughout the Appalachian coal region, crownvetch was seeded on mine sites [73,137]. In 1977, the US federal government established guidelines that required establishment of 70% perennial ground cover on abandoned mine sites. Crownvetch provided rapid perennial cover and was used extensively [58]. In some areas, crownvetch has escaped from the revegetation site and is growing in adjacent native vegetation. On an 18-year-old coal surface-mined site in southeastern Kentucky, crownvetch was "adapting to the point of becoming naturalized". In the mid 1960s, crownvetch was used to revegetate coal spoils, but in 1984, researchers described crownvetch as "especially aggressive" in the area and found plants in adjacent vegetation excluding native plants. Although seeding crownvetch on coal spoils provided rapid cover as required by federal and state reclamation laws, dense crownvetch growth may limit establishment of native herbs, shrubs, and trees [137].
Local distribution changes: Escaped and invasive crownvetch populations are common in the Midwest and occur sporadically in surrounding areas, but in other areas escapees are rare [133,155] or restricted to disturbed sites [27,65,126]. Although not known to escape and persist throughout its nonnative range, reviews indicate rapid spread by seed [22,121], and long-distance seed dispersal by deer is likely (see Seed dispersal).
In the Midwest and eastern United States, studies have documented crownvetch spread. In Pennsylvania, crownvetch was seeded on highway slopes at more than 100 sites in 30 counties. Even with years of below normal precipitation, crownvetch cover was dense on 75% of seeded slopes within 2 years [92]. Within ten years of seeding a 10-foot (3-m) length of roadside in Indiana, crownvetch spread to occupy a 75-foot (23 m) length of roadside [46]. In Boone County, Iowa, a crownvetch patch increased in size from 32,900 ft² (3,060 m²) in 2004 to 39,100 ft² (3,630 m²) in 2005, through the establishment of new patches and growth of existing patches [84]. During surveys conducted along railroads in St. Louis, Missouri, crownvetch colony was found in 1971 but was not found in earlier surveys that began in 1954 [97]. As of 2002, crownvetch was invading and/or disrupting native plant communities in more than 10 Missouri counties [95]. In 2003, crownvetch occurred on 4 sites and 0.4 acres (0.2 ha) of Michigan's Ottawa National Forest [145]. Two years later, crownvetch occurred on 14 sites and 11 acres (4.5 ha) of the Forest [143]. Crownvetch occurred in oak openings in northwestern Ohio by 1928. In a 1978 survey of the same study area, crownvetch was found fewer than 5 times [28]. Crownvetch was absent from Ohio's Athens State Forest flora in 1957, but in 2005 it occurred in disturbed areas of the Forest [45]. As of 2001, Ohio's Department of Natural Resources considered crownvetch well established in the state (Windus and Kromer 2001 cited in [61]). On cut slopes in West Virginia, crownvetch spread beyond seeded areas on at least 2 sites. On one site, crownvetch spread 33 feet (10 m) outside of the planting area within 6 years of seeding. On an "extremely harsh" site, crownvetch spread to double the size of the seeded area. Vegetation adjacent to these sites was not described [100].
Fuel characteristics of crownvetch were only noted in a few references. Crownvetch generally remains green in very dry conditions [40] when most associated grasses are "drought-browned" [39]. Fire spread may be limited by dense patches of crownvetch in invaded habitats, although no studies have documented this. In a review, Tu [139] reported that fire spreads poorly in crownvetch-dominated stands and often fails to burn into the center of dense patches. Fire studies on sites with dense crownvetch populations are needed.
See the Fire Regime Table for more information on the FIRE REGIMES in communities that may include crownvetch. Find further fire regime information for the plant communities in which this species may occur by entering the species name in the FEIS home page under "Find FIRE REGIMES".
Generally, crownvetch seed germination is best when seeds are scarified and shallowly buried in moist, nonsaline, warm soils (air temperatures 68-77 °F (20-25 °C)). The percentage of crownvetch seed that is hard or impermeable to water varies. Reports indicate that 20% [111] to 80% of seed is hard [1]. Once scarified, crownvetch seed germination percentages are usually high (85-90%) [40,111].
In his review of species that produce water-impermeable seeds, Rolston [110] indicated that hard seed coats can be broken down or punctured naturally by fire or digestive tract passage. In the laboratory, 29% of unscarified crownvetch seeds germinated. When seeds were kept in 190 °F (90 °C) water for 4 hours, germination was 42%, and when mechanically scarified, germination was 63%. Regardless of the method, germination was significantly greater for scarified than unscarified seeds (P<0.05) [26]. Researchers found that most hard crownvetch seeds had to be pierced to a depth of at least 98 µm before they absorbed water and germinated [91]. In other controlled experiments, very short-duration heating softened crownvetch seeds, but heating for more than 1 minute killed many seeds. Hard seeds heated to 190 °F (85 °C) or 210 °F (100 °C) in less than 1 minute beneath heat lamps germinated twice as well as untreated seeds. About 60% of hard seeds rotted when heating occurred over a 2-minute period. Oven heat treatments of 210 °F (100 °C) or 260 °F(125 °C) resulted in "nearly complete (seed) death" [13]. Experiments conducted on hard seeds of 2 crownvetch cultivars suggested that abrupt changes in temperature produced greater germination than long exposure to high temperatures. Germination of hard seeds increased after 5 seconds in boiling water, and for seeds that were still hard after this treatment, germination was increased by a second boiling water treatment (<30 seconds). After 2 short-duration boiling water treatments, crownvetch seed germination was greater than after a single long-duration boiling water treatment [33].
Based on 2 studies, shallow burial in nonsaline soils is best for crownvetch seed germination. In the greenhouse, crownvetch emergence was significantly greater from 2-inch (4-cm) depths than from the soil surface or 3-inch (8-cm) depths (P<0.05) [85]. A Utah forage and conservation planting guide recommends seeding crownvetch in the fall at 0.25 to 0.5 inch (0.5-1 cm) deep [56]. Another guide recommends a seeding depth of 0.5 cm or less (review by [129]). According to a review [147], frost heaving can force crownvetch seeds into the soil. In soils with a salt content of 5 g/kg, crownvetch seed germination was delayed, and germination percentages were "markedly" decreased compared to seeds in nonsaline conditions [82].
Warm temperatures (up to 75 °F (24 °C)) and increased available moisture generally increase crownvetch seed germination. In controlled conditions, increasing temperatures typically lead to increased germination of scarified crownvetch seed. Germination was about 20% at 54/43 °F (12/6 °C), 60% at 59/50 °F (15/10 °C), and 80% at 75/64 °F (24/18 °C) day/night temperature regimes [48]. In soil samples collected from a highway road cut, crownvetch seedling emergence was 57%, 10%, and almost 0% from samples with moisture tensions of -1/3, -3, and -6 bars, respectively. After 3 or 4 days of monitoring, total emergence was greater at 70 °F (21 °C) than at 82 °F (28 °C), regardless of moisture levels [168].
Throughout its North American range, crownvetch is most common in disturbed areas including fields, former dwellings, roadsides, rights-of-way, and stream banks [7,89,105,113,119,133,166,169]. For a discussion of areas where crownvetch is invasive, see Impacts.
Climate: Crownvetch tolerates a wide range of climates in North America. Reviews report that crownvetch grows well in USDA hardiness zones 3 to 7 [123] and tolerates "long" periods of drought, precipitation of up to 65 inches (1,650 mm)/year, and temperatures as cold as -27 °F (-33 °C) [139]. Revegetation guides for the Intermountain West suggest that crownvetch grows well on sites receiving at least 21 inches (530 mm) of annual precipitation and thrives on sites receiving more than 30 inches (760 mm) of annual precipitation [56,129]. Although tolerant of many climate regimes and present nearly throughout the United States, crownvetch introductions were not very successful in central Alaska, Palo Alto, California, southern Georgia, or Florida. Crownvetch establishment failed on saline shorelines, and plants were winter-killed in central Alaska [92]. For information on the cold tolerance of crownvetch seedlings, see Temperature and soils.
Elevation: Only in Nevada and parts of California are elevation ranges reported for crownvetch. In Nevada, crownvetch occupies a narrow elevational range of 4,500 to 4,600 feet (1,400 m) [67]. In California's Butte and Nevada counties, crownvetch populations were reported at 2,341 feet (714 m) and 7,100 feet (2,160 m), respectively [24].
Soils: Crownvetch establishes and persists on sands, loams, clays, and gravelly and rocky soils (reviews by [83,139,147]). Although crownvetch has "excellent" drought tolerance [83] and in several areas occurs in dry rocky soils ([24], review by [123]), plants are unlikely to persist on sites with less than 15% very fine sand, silt, and/or clay [147]. Highly compacted soils can also limit crownvetch establishment and growth [111]. Moderately acidic and low fertility soils are tolerated [83], but saline and poorly drained soils are not [92,139]. In soils with a salt content of 5 g/kg, crownvetch germination was delayed and reduced as compared to seeds in nonsaline soils [82].
Moisture: Although tolerant of both moist and dry climates [31], crownvetch requires well drained soils [92]. Crownvetch rarely grows well or survives long in poorly drained soils [56,92]. Along the shores of Cave Run Lake in northeastern Kentucky, crownvetch was absent from mud flats but did occur on frequently and infrequently flooded shorelines. Mud flats occurred below the summer pool shoreline, frequently flooded sites were up to 7.9 feet (2.4 m) above summer shoreline, and infrequently flooded sites occurred at 7.9 to 24 feet (2.4-7.3 m) above summer shoreline [87]. During field surveys in Pennsylvania, crownvetch was absent from areas with evidence of soil mottling or water pooling [93].
pH and fertility: Crownvetch may grow best on neutral or alkaline soils [56,59] but also establishes on acidic soils (pH 5) and spreads vegetatively into highly acidic soils [152]. In Europe, crownvetch often occurs on calcareous soils [59], and in Utah, crownvetch grows well in calcareous soils [56]. In a revegetation guide for eastern coal mine spoils, Vogel [152] reported that crownvetch seedling establishment was best on soils with a pH of 5.5 or greater, but established plants usually spread into soils with a pH of 4.5 or less. Crownvetch stand development was considered a "failure" 2 to 4 years after seeding a strip mine spoil with a pH of 4.5 [111]. One to two years after seeding in Davis County, southeastern Iowa, crownvetch cover was greatest (7%) in moderately acidic (5.2-5.9 pH) soils. Crownvetch plants were largest in acidic (4.8-5.4 pH) silt loams. Plant size and cover (2%) were lowest on acidic (5.4-6.2 pH) soils with the shallowest top soil layer [14].
Low fertility soils are tolerated, but crownvetch growth is improved on fertile soils (Barnes and others 1995 cited in [31]). Based on field and greenhouse studies conducted in Pennsylvania, researchers concluded that rapid seedling growth and persistence of crownvetch occurs on soils with good or excellent drainage, high levels of phosphorus, potassium, calcium, and magnesium, and pH levels of 6.5 to 7. In the greenhouse, crownvetch failed to establish in coal mine soils with a pH of 2.8. Seedlings in soils with a pH of 6.3 to 7.7 and high calcium content (1,900-4,800 kg/ha) were larger than those in soils with lower pH and calcium content. In the field, dense crownvetch stands occurred on soils with pH ranging from 4.6 to 7.7 [93].
Although crownvetch growth may be best on fertile and moist but well drained sites, sources indicate decent growth and persistence on dry, low fertility sites as well. During droughts on dry, low fertility soils in West Virginia, crownvetch grew "vigorously" [111]. In Pennsylvania, researchers recognized crownvetch as a potentially useful plant for the revegetation of harsh sites, when it occurred "at fertility levels so low that only poverty grass would normally survive" [162].
Although its seeds and foliage may be poisonous to nonruminants [41,133], elk, deer, most classes of livestock, and rabbits feed on crownvetch. Crownvetch is considered high quality livestock, deer, and elk forage, and in the winter, these large mammals paw through the snow to feed on crownvetch. Ground-nesting birds, meadow voles, and rabbits utilize crownvetch as cover (reviews by [129,139,147]).
Livestock: Reviews indicate that crownvetch is good, palatable livestock forage [129,139,147] but contains nitroglycosides that are toxic to horses (review by [123]). Sheep consume crownvetch leaves but typically avoid stems, and cattle may avoid crownvetch in early spring when foliage is less palatable (review by [120]). In Pennsylvania, Hereford steers gained 1.5 to 2 lbs/head/day on crownvetch pasture [92].
Deer: Along an Interstate in central Pennsylvania, researchers found white-tailed deer abundance increased in late fall and winter in crownvetch-dominated vegetation. Other vegetation in the study area included cultivated fields and mixed-deciduous forest with conifer clumps [19].
Small mammals: Studies indicate that crownvetch can be important meadow vole cover but may not be important forage. In a commercial crownvetch field in Centre County, Pennsylvania, 4 of 5 meadow vole brood nests occurred in dense crownvetch patches (94% cover) [117]. In this same area, meadow voles did not consume crownvetch when presented with other foods [63], and when provided only crownvetch and water, meadow voles died within 3 days (Jones 1978 cited in [63]). In newly planted apple orchards in New York, meadow vole population densities were high in plots where crownvetch was the ground cover. Researchers observed meadow voles feeding on crownvetch leaves, and long crownvetch stems allowed meadow voles to climb inside the mesh guards installed to protect tree trunks. Cumulative tree mortality over a 4-year period was much higher in plots with crownvetch cover than in plots with different or lacking ground cover [94].
Insects: Crownvetch flowers are visited by butterflies and moths in North America. During surveys of Pennsylvania's Allegheny Portage Railroad National Historic Site and Johnstown Flood National Memorial, butterflies feeding on crownvetch made up 7% of 200 feeding observations. Crownvetch was utilized by 2 butterfly species [68]. In southern New Jersey, crownvetch was a food source for orange sulfur butterflies in mid-winter and was also a "reliable" summer food [112]. In the early 1990s, researchers found Coleophora colutella on crownvetch plants on Cape Cod. This European moth was likely introduced with imported plant material. Early surveys indicated that this moth was using only crownvetch as a host, but in Europe, the moth uses several legume hosts [53]. For more information, see the discussion on crownvetch's impacts on insects.
Like most nonnative invasive plants, successful control of crownvetch is most likely in the early invasion stages (review by [139]). Because crownvetch is often restricted to disturbed sites and other habitats primarily dominated by nonnative species [32,70], nontarget effects of control may be reduced, but a lack of native plants and seeds may require active revegetation on control sites. Control of biotic invasions is most effective when it employs a long-term, ecosystem-wide strategy rather than a tactical approach focused on battling individual invaders [88].
In all cases where invasive species are targeted for control, no matter what method is employed, the potential for other invasive species to fill their void must be considered [16]. In the Lost Mound Unit of the Upper Mississippi River National Wildlife and Fish Refuge in northwestern Illinois, Kentucky bluegrass (Poa pratensis) increased dramatically when crownvetch cover was reduced by herbicide treatments. The study area was dominated by little bluestem and sand dropseed (Sporobolus cryptandrus) with monoculture patches of crownvetch. One year after herbicide treatments, crownvetch cover was less than 1% on treated sites, significantly less than that of untreated sites (P<0.0001). Available soil nitrogen was significantly (P<0.0001) greater inside than outside crownvetch patches but was not significantly affected by the herbicide treatment. Kentucky bluegrass cover was greater than 50% on treated plots and averaged 14% on control plots. Native species cover was not significantly different on treated and untreated sites. Kentucky bluegrass may have utilized high nutrient levels better than the native species [134].
Fire: For information on the use of prescribed fire to control crownvetch, see Fire Management Considerations.
Prevention: It is commonly argued that the most cost-efficient and effective method of managing invasive species is to prevent their establishment and spread by maintaining "healthy" natural communities [88,116] (e.g., avoid road building in wildlands [142]) and by monitoring several times each year [62]. Discontinuing the use of crownvetch in the revegetation of roadsides and mine sites could be key to preventing crownvetch establishment in areas where it does not yet occur (review by [22]).
Managing to maintain the integrity of the native plant community and mitigate the factors enhancing ecosystem invasibility is likely to be more effective than managing solely to control the invader [51]. Weed prevention and control can be incorporated into many types of management plans, including those for logging and site preparation, grazing allotments, recreation management, research projects, road building and maintenance, and fire management [146]. See the Guide to noxious weed prevention practices [146] for specific guidelines in preventing the spread of weed seeds and propagules under different management conditions.
Cultural Seeding or planting native plants in crownvetch stands may provide some control. Grover (2001 personal communication cited in [139]) suggests planting cool-season native grasses, such as switchgrass, into sites invaded by crownvetch or on sites where crownvetch was treated. In tall fescue and crownvetch stands along roadsides in West Virginia, establishment and growth of seeded native species were best when stands were herbicide treated or tilled before seeding [118].
Physical or mechanical Small crownvetch populations may be controlled by hand-pulling, digging, or heavy shading by cloth or mulch [44]. For larger crownvetch populations, mowing may be a more useful mechanical control method.
Hand-pulling or digging of crownvetch is time consuming, labor intensive, and requires the removal of all stems, roots, and rhizomes. Sites should be monitored for seedlings and sprouts in successive years (review by [139]). Mowing to control crownvetch spread may be most effective in the late spring or at flower bud stage (reviews by [119,121,123,139,151]), although Henson [47] reported that crownvetch recovery was slow when plants were cut for hay at the full bloom stage. Repeated mowing is necessary (reviews by [119,121,123,139,151]). A review recommends mowing twice a year, first in June and again in late August [22]. Another review suggests that mowing treatments should avoid native or desirable vegetation [123]. While mowing may not be feasible in wildlands, controlling crownvetch in adjacent public use areas may limit its spread into wildlands.
Repeated mowing may be necessary to deplete the carbohydrates stored in crownvetch roots. Immediately following clipping, total available crownvetch root carbohydrates generally decreased but recovered quickly as aboveground stems grew. Researchers found that clipping crownvetch early in the growing season (before 1 June) or late in the growing season (August-October) allowed other "weedy" species to invade crownvetch stands. Clipping also limited seed production [77].
Biological Crownvetch has insect predators in North America and may be reduced by livestock grazing. During a 2-year survey of insects in Pennsylvania in the early 1970s, Wheeler [161] collected 125 phytophagous arthropods in crownvetch stands. Differential and redlegged grasshoppers were the most destructive crownvetch pests, but alfalfa plant bugs, potato leaf hoppers, clover stem borers, pollen beetles, and glechiid moths also injured or stressed crownvetch plants [161]. In Kansas, researchers found the striped willow flea beetle using crownvetch as a host plant, suggesting it may be useful as a biological control in areas where crownvetch is invasive. However, the striped willow flea beetle has other known hosts, primarily willows (Salix spp.) and cottonwoods (Populus spp.) [108]. Several studies indicate that crownvetch abundance is typically lower on grazed than ungrazed sites. Livestock may reduce crownvetch abundance by removing aboveground biomass, compacting soils, or both. For more information, see Grazing.
Biological control of invasive species has a long history that indicates many factors must be considered before using biological controls. Refer to these sources: [149,165] and the Weed control methods handbook [140] for background information and important considerations for developing and implementing biological control programs. Because crownvetch is still planted for erosion control and in pastures, release of a biological control is unlikely (review by [139]).
Chemical While herbicides are often effective in gaining initial control of a new invasion or a severe infestation, they rarely provide complete or long-term weed control [18]. See the Weed control methods handbook [140] for considerations on the use of herbicides in natural areas and detailed information on specific chemicals.
For information on herbicides and applications useful for crownvetch control, see [139]. Applying herbicides before crownvetch flowers and before it develops a thick vegetative mat (late April or early May) may be most effective (Walters 2001 personal communication cited in [139]). To kill crownvetch plants, complete coverage of aboveground material is necessary. Prescribed fire to remove litter may improve herbicide coverage (reviews by [119,121,139]).
Integrated management: An integrated weed management plan for heavily infested crownvetch sites may include cutting or burning a site, treating with an herbicide, then actively revegetating and managing to encourage dense growth of native vegetation (review by [139]).
Descriptions of crownvetch's weedy behavior range from "minor" [34] to "highly invasive" [144]. Crownvetch is often, but not exclusively, described as weedy and problematic in the Midwestern and eastern United States. Although considered aggressive in some North American locations [42,107,144], in many areas, crownvetch is restricted to disturbed sites and considered a minor threat to native vegetation [25,27,32,34,70,126,150]. Whether or not crownvetch is more invasive in the Midwest and eastern United States because of site conditions, more plantings, and/or greater time since first introductions is unknown. Oregon's Native Plant Society found crownvetch in the Willamette Valley in 2008 and suggested that crownvetch may be in the lag phase of colonization but may, in time, develop into a "high impact" species capable of modifying native habitats and/or altering ecological functions and processes [99].
Although more common in the eastern and Midwestern United States, descriptions of crownvetch's aggressive growth habit and potential to negatively impact native vegetation occur nearly throughout crownvetch's nonnative North American range. At the Konza Prairie Biological Station in Kansas, crownvetch is "aggressively invading" tallgrass prairie [42] and is considered "a serious management threat to prairies and savannas" (review by [121]). In a survey of plant and natural area experts in Wisconsin, crownvetch ranked 13th when 66 nonnative invasive plants were evaluated for their impacts on native plant communities [107]. The Eastern Region of the Forest Service ranks crownvetch as "highly invasive" [144]. As of 2001, crownvetch was considered a "significant" threat in Tennessee [135] and as of 2008, was considered a severe threat in Kentucky [69].
Introduction and spread: In some cases, where crownvetch has been used in revegetation it has displaced native plants. In a restoration handbook for the West, researchers cautioned that crownvetch is "very competitive" and often dominates revegetated sites to the exclusion of other vegetation [129]. Colorado systematists indicated that crownvetch produces dense cover rapidly and "thrives in shade" [159]. As of 2002 in Missouri, crownvetch had invaded and/or disrupted native plant communities in more than 10 counties [95]. In Michigan, crownvetch was heavily promoted as a rapidly growing ground cover good for erosion control. Crownvetch populations spread extensively along roadsides, waterways, and fields in that state [154]. For more about crownvetch spread, see Local distribution changes.
In several areas, crownvetch populations persist as monocultures. In Pennsylvania, crownvetch likely arrived as a contaminant in alfalfa seed. It was discovered in an alfalfa field in the early 1900s, and 60 years later a dense crownvetch stand remained in the field. Except for a few sumac shrubs (Rhus spp.), few other species had established in the crownvetch stand [92]. Crownvetch was used for revegetation throughout the Appalachian coal region [137]. In southeastern Kentucky, it was used to revegetate a coal mine site where, 24 years later, observers described crownvetch growth as "especially aggressive". Crownvetch spread from the original planting area into adjacent vegetation. While crownvetch has persisted on harsh and continually disturbed sites, it did not persist on Lee Canyon ski slopes in southern Nevada. Crownvetch was seeded on the slopes between 1970 and 1980 but was not found in a 1999 survey of ski runs and adjacent forests [138].
Displacement of native vegetation is the most commonly described impact in crownvetch-invaded habitats, but delayed succession and reduced reproduction were also reported. Crownvetch may limit the development of some native plants by increasing available nutrients ([134], review by [158]) or through allelopathy [90,131]. However, nutrient increases were not directly measured on invaded sites, and results from allelopathy studies are mixed. In some areas, crownvetch invasions may reduce insect diversity.
Impacts on vegetation:
Native grasses and forbs: Rapid, dense crownvetch growth can displace native plants. Experimental studies and observations suggest that impacts to native vegetation are greatest after crownvetch is established and producing substantial vegetative growth. Crownvetch growth has been described as a dense mat that "gradually chokes out more or less persistent weeds" [162]. Land managers have associated crownvetch spread with the displacement of native plants in prairies in Minnesota's Ottawa Bluffs Preserve (McGuigan 2002 cited in [139]) and in Iowa's Broken Kettle Grasslands and Ames High Prairie Preserve (Moats 2001 cited in [139]). In shale barren communities of the Allegany National Forest in Pennsylvania, crownvetch has excluded native plants from open woodlands (Keech 2002 cited in [139]). In Kentucky's Rolling Fork/Salt River Drainage area, crownvetch was associated with reductions in the abundance of tall warm-season grasses, native forbs, and 2 threatened species: Tennessee gladecress (Leavenworthia exigua) and Eggert's sunflower (Helianthus eggertii) (Mazyck 2002 cited in [139]). However, when seeded into plots with established vegetation in western Iowa, crownvetch seedling emergence and growth were "uniformly poor". Established vegetation included a bluestem (Andropogon spp.) monoculture, a mixture of tallgrass prairie species, and a mixture of tall and shortgrass species [85].
In Kentucky, crownvetch occurs in communities with Short's goldenrod (Solidago shortii), a federally endangered species. During a field experiment in Robertson County, Short's goldenrod seedlings established and flowering increased by about 3 times in plots where associated vegetation (dominated by crownvetch) was removed. The number of nonflowering Short's goldenrod ramets increased each year associated vegetation was removed, and Short's goldenrod seedlings occurred in 9 of 10 treated plots but not in control plots [156]. In a later vegetation survey of Harrison County, Kentucky, researchers found a Short's goldenrod population in a brush prairie gravel wash community where crownvetch also occurred. Impacts were not evaluated in this study [54].
Crownvetch can limit success in prairie restoration. On the Walnut Creek National Wildlife Refuge in Iowa, crownvetch occupies 1,000 of acres and is considered one of the most problematic species in restoration projects (Shutte personal communication [37]). On a research farm in Monoma County, Iowa, researchers attempted to restore a smooth brome (Bromus inermis) pasture to prairie. In 2002, the pasture was herbicide treated, plowed, and planted to prairie species. In 2003, crownvetch was present but only in trace amounts. By May 2005, crownvetch covered 46% of the restoration area, and the project was abandoned. When the project began, crownvetch was restricted to a ditchbank adjacent to the pasture (Wilsey unpublished data cited in [85]).
Woody plants: On some sites, crownvetch may limit shrub and tree seedling recruitment and delay succession. In a revegetation guide for coal mine spoils in the eastern United States, Vogel [152] recommended against planting crownvetch with tree seedlings, and a review reports that crownvetch often shades out shrub and tree seedlings [44].
Crownvetch plantings in Kentucky, Pennsylvania, and Maryland restricted the establishment of woody species. Along a road in Campbell County, Kentucky, crownvetch dominated the understory within and outside 8- to 10-year-old staghorn sumac (Rhus typhina) clumps. Researchers found few tree seedlings in the area and suspected crownvetch was limiting tree establishment, because in other succession studies, tree seedlings were often present in 8-year-old staghorn sumac stands [86]. On 51 sites in Pennsylvania and 12 sites in Maryland seeded with crownvetch at least 10 years earlier, woody plant density generally decreased as crownvetch cover increased. On sites with no crownvetch cover, there was 1 woody plant/160 feet² (15 m²). When crownvetch cover was 80%, there was 1 woody plant/1,540 feet ² (143 m²) [115].
On sites in West Virginia and Ontario, however, crownvetch growth did not limit the establishment of woody vegetation. Within 2 years of seeding crownvetch on a roadside in West Virginia, smooth sumac (Rhus glabra), sycamore (Platanus occidentalis), and sugar maple (Acer saccharum) were "naturally encroaching" in dense crownvetch cover [100]. For up to 5 years after crownvetch was seeded on a utility right-of-way near Tobemory, Ontario, tree regeneration was not restricted [17].
Allelopathy: Results from studies designed to uncover crownvetch's allelopathic potential are mixed. Leachate from crownvetch seeds did not significantly (P<0.05) reduce the germination of a variety of field, forage, turf, weed, flower, and vegetable species, but often seedlings were abnormal. Roots and shoots of seedlings emerging in crownvetch seed leachate were shorter than those of seeds germinated in water [90]. Studies conducted at the Ohio Agricultural Research and Development Center showed that growth of northern red oak (Quercus rubra) was inhibited when seedlings were watered with crownvetch foliage extracts (Larson unpublished data cited in [153]). In another study, crownvetch extracts stimulated germination of quackgrass (Elymus repens) and crownvetch but inhibited germination of smooth brome and meadow fescue (Schedonorus pratensis). Extracts were made from live crownvetch shoots collected from an abandoned field in Argonne, Illinois. An extract made from decomposing crownvetch roots or shoots rarely affected the growth of other plants. Often the extracts from decomposing crownvetch roots stimulated the growth of crownvetch, quackgrass, smooth brome, meadow fescue, and timothy (Phleum pratense) [131].
Impacts on insects: Several studies related crownvetch invasions to unbalanced insect herbivory and reduced insect abundance and diversity. In tallgrass prairie at the Konza Prairie Biological Station in northeastern Kansas, total leaf area reductions due to natural enemies and/or abiotic stresses were significantly greater for a native slimflower scurfpea (Psoralea tenuiflora) than for crownvetch [42]. When bee populations were compared in "restored" prairies and "weedy" roadsides in Kansas, average bee richness and abundance were significantly (P<0.05) greater in restored than weedy vegetation. Nonnative forbs including crownvetch, sweetclover (Melilotus spp.), and common teasel (Dipsacus fullonum) covered more than 50% of weedy roadsides. Restored prairies likely provided a greater diversity of insect foods than the roadsides. Before the study, prairie and roadside vegetation was mowed multiple times each year, and roadside vegetation was treated with herbicides. Effects of prior management on bee populations were not discussed [55]. In western Wisconsin and eastern Minnesota, researchers found that Melissa blue butterfly (Lycaeides melissa melissa) populations have expanded their range and are using crownvetch as a larval host. This range expansion puts Melissa blue butterfly populations near endangered Karner blue butterfly (L. m. samuelis) populations. Although it is unknown whether these species can hybridize and produce viable offspring, hybridization could threaten Karner blue butterfly as a distinct taxon [23].
The use of crownvetch to revegetate and control erosion on abandoned mine sites and roadside cuts has likely facilitated its wide distribution in North America. Crownvetch has recently been used as a cover crop in the United States. In Virginia, crownvetch was used in conjunction with biological control agents to control Canada thistle (Cirsium arvense) populations. Aboveground biomass of Canada thistle was reduced by 69% when tall fescue and crownvetch were seeded in an area where the tortoise beetle (Cassida rubiginosa) was established [4]. Canada thistle reductions were greater on defoliation plots that had been seeded with crownvetch and tall fescue than on plots with defoliation alone [5].
Although used as a cover crop and for erosion control, some studies suggest that crownvetch may persist indefinitely on these sites, may not provide good erosion control, and may not provide useful cover. In 1990, crownvetch was seeded on a coal mine site in Wise County, Virginia, and remained on the sites at least 11 years after seeding [31]. The Natural Resources Conservation Service described crownvetch as a "useful but overused erosion control plant". While crownvetch has partially stabilized rocky, steep slopes in the eastern United States, rills and gullies often occur beneath the crownvetch canopy. On dry sites, seeding switchgrass with crownvetch provided better slope stabilization than seeding crownvetch alone (review by [147]). In apple orchards in New York, long crownvetch stems allowed meadow voles to climb over the mesh guards installed to protect tree seedlings. Tree mortality was much higher in plots with a ground cover of crownvetch than on plots lacking a ground cover or with another species as ground cover [94].
Crownvetch flowers are possible from May to September throughout its US range [8,41,89,105,169]. In the northeastern United States, crownvetch flowering is "profuse" for a 6-week period in May and June (review by [83]). However, crownvetch flowers and produces seed throughout the growing season. Often both newly opened flowers and mature fruits occur on the same plant [84]. In the winter, at least the basal crownvetch leaves remain green (review by [129]).
Based on 2 years of study, total available root carbohydrates decreased from about 18% in late April to seasonal lows of 12% or 13% as vegetative growth was produced [77]. During field studies conducted near Blacksburg, Virginia, researchers found that the total nonstructural carbohydrates of crownvetch plants were generally highest in the fall (October and November) and lowest in summer (July and August), regardless of frequent or infrequent clipping [167].
Reports on the persistence of crownvetch seed in the soil vary, and experimental seed bank studies are generally lacking. Grover (2001) indicates that crownvetch is capable of producing a large soil seed bank (personal communication cited in [139]). A weed management guide reports that crownvetch seed may be viable for 15 years or more [22]. However, during studies in calcareous grasslands in Germany's middle Swabian Alb, crownvetch seeds were typically viable for less than 2 years in the soil [102]. Up to 25% of seed produced by crownvetch is hard and requires scarification before it will germinate. Likely hard seeds contribute most to the seed bank. Additional information about hard crownvetch seeds is presented in the Germination section below.
Storage studies indicate that crownvetch seed in an area with minimal temperature and humidity fluctuations remains viable for 11 to 15 years (review by [64]). After 8 years of storage in an open warehouse in Sanpete County, Utah, stored crownvetch seed germinated better (70%) than freshly collected seed (41%) (P<0.05). Seeds stored for more than 8 years failed to germinate. Temperature extremes in the open warehouse ranged from -29.9 °C to 38.3 °C over a 25-year period [128].
In Iowa, no crownvetch seedlings emerged from soil samples collected from established crownvetch patches [84,85]. Crownvetch also failed to emerge from soil samples collected from a constructed wetland along the Delaware River near Trenton, New Jersey, although crownvetch occurred in the aboveground vegetation [80]. However, up to 384 crownvetch seedlings emerged from soil samples taken from a roadside in Campbell County, Kentucky, where crownvetch was dominant. Crownvetch emergence was lower from soil samples taken beneath staghorn sumac (Rhus typhina) clumps than from samples collected outside staghorn sumac clumps [86].
Crownvetch typically produces seed in its 2nd year of growth (Bender personal observation cited in [11]), and seeds typically mature 6 to 10 weeks after flower pollination. Generally, seed production is low, and high precipitation, low light, and/or low temperatures may reduce production. Seed production of 75 to 175 lb/acre is common, although 400 lb/acre has been reported [92]. Seed production may be limited by pollinators. When bee hives were located near a crownvetch population in Pennsylvania, production averaged 16.4 metric tons of air dry seed/ha [1].
Seed maturation and production can vary with growing conditions. In a year with above-average growing season precipitation, production averaged 257 kg/ha, and in a year with slightly below-average growing season precipitation, production averaged 530 kg/ha [1]. In years with abundant moisture, crownvetch often produces a "flush" of vegetative growth, which shades developing seed pods and delays or limits seed development and maturation [92,111]. On low fertility, droughty soils, vegetative growth rarely interferes with seed development and maturity [111]. In a field study, crownvetch plants that were covered by screens with bees inside produced lighter and fewer seeds than plants in the open. The researcher considered reduced light beneath the screens the likely reason for reduced production [3]. Low night temperatures in late summer or early fall can "retard seed maturation" [92].
Seedling establishment and plant growth: In the first 1 to 2 years after seeding, crownvetch establishment and growth are limited, but 3 to 4 years after establishment, crownvetch can form dense stands (reviews by [83,127]). Site conditions including winter temperatures, soils, moisture availability, and associated vegetation affect crownvetch establishment and productivity.
Two reviews report that crownvetch establishes and spreads well from seed [22,121], but a review from the Great Plains reports that crownvetch is often "found close to where it was originally planted" and does "not seed well" [133]. An Intermountain revegetation guide also rated crownvetch establishment and spread from seed as "poor" [101].
Slow emergence, growth, and establishment are often reported for 1st-year crownvetch seedlings (reviews by [47,83,111,129]). Six- to seven-week-old crownvetch seedlings had an average relative growth rate of 0.117 g/g/day in sand with nutrients added [29]. While crownvetch may completely cover a site within 2 or 3 years of seeding (review by [83]), in the western United States it typically takes 3 to 4 years to flower (review by [127]). In a garden, crownvetch completely covered an area by its 2nd growing season, but aboveground growth was slow in the 1st several weeks after seeding when roots were developing [39].
Temperature and soils: Crownvetch seedlings are sensitive to cold temperatures, soil acidity, and soil compaction. Seedlings emerging after July will not likely survive the winter [92]. Crownvetch seedlings less than 4 inches (10 cm) tall at the time of the first killing frost rarely survive (Woodruff and Blaser 1970 as cited in [100]). Acidic and/or compacted soils can limit crownvetch seedling establishment and growth [40,111]. Soon after seeding in West Virginia, crownvetch cover averaged 70% on mechanically scarified slopes and 32% on compacted slopes. By 18 months after seeding, however, crownvetch cover was 95% to 100% on both slopes [100].
Low moisture levels at the seedling stage will likely reduce crownvetch establishment. Crownvetch establishment was poor in a cleared forest along a utility right-of-way near Tobemory, Ontario. Crownvetch established on only half of the seeded plots, and cover averaged only 0.15% at the end of the 1st growing season. Summer drought was blamed for poor establishment [17]. In the revegetation of Lesser Slave Lake Basin in central Alberta, crownvetch establishment was best in low areas between the sand dunes and poorest on sand dune crests [81]. For more information on crownvetch seedling and plant growth as related to soil characteristics, see Soils.
Associated vegetation: Crownvetch seedling establishment is best on open sites lacking other vegetation [92]. When crownvetch was seeded in a 10-year-old switchgrass (Panicum virgatum) stand near Ames, Iowa, establishment was low and averaged 0.9% in July after the 1st seeding and 7% in August after a 2nd seeding. Crownvetch was seeded at a rate of 171 seeds/m² [12]. The mechanisms by which crownvetch establishment is limited in intact, undisturbed vegetation were not discussed in the available literature (2010), but researchers showed that initial establishment is not limited by shading. Crownvetch seedlings survived 1 month of receiving just 1% to 2% incident radiation beneath an oat (Avena spp.) canopy (Buxton and Wedin 1970 cited in [100]).
Crownvetch is an early- to mid-seral species that is moderately shade tolerant. In many areas, crownvetch is restricted to disturbed sites, although grazing and mowing may decrease its abundance.
Studies in its native range indicate that crownvetch occurs in early- and mid-seral stages of secondary succession. Crownvetch may appear earlier when succession begins on bare ground than when succession begins with field abandonment. During the succession of bare sites created during road construction in eastern Bohemia in the Czech Republic, crownvetch occurred with low cover in the first 2 years. Crownvetch dominated the site in years 3 to 5, decreased in year 6, and was rare in year 7 [71]. In another successional study of 16 seres in the Czech Republic, crownvetch was most common after 10 years of succession. Seres began as bare ground resulting from human-caused disturbances and covered up to 76 years of vegetation development [104]. In old field succession in the Bohemian Karst Protected Region of Czech Republic, crownvetch was more common on fields abandoned 30 to 60 years earlier than on those abandoned 6 to 20 years earlier [9]. In Germany, crownvetch was common in old meadows (>30 years abandoned) but not in pioneer communities or early-seral fields (1-4 years abandoned) [124].
Shade tolerance: Crownvetch is described as preferring full sun but tolerating partial shade (reviews and management guides by [22,123,129]). However, Lorenz and others [83] suggest that crownvetch has "poor" shade tolerance, and Weber and Wittmann [159] indicate that crownvetch thrives in shaded conditions.
Field experiments and observations indicate that crownvetch does tolerate some shading, but long-term persistence in partial or full shade was rarely evaluated. In West Virginia, crownvetch was seeded with black locust (Robinia pseudoacacia). Two years after seeding, crownvetch cover was nearly 100%. Four years after seeding, black locust produced a nearly closed canopy, but cover of crownvetch was not "diminished" [111]. In a field study, normal light level reductions of 60% or more were required to reduce crownvetch growth (review by [100]). In established crownvetch stands in Pennsylvania, total herbage yield of crownvetch was significantly lower (P<0.05) in shaded than unshaded plots. In areas shaded 58% to 78% for 2 years from late July to late September, crownvetch yield was 6.7 to 8.8 mg/ha and in unshaded areas was 8.6 to 10.0 mg/ha [20]. During experiments conducted near Blacksburg, Virginia, crownvetch plants dropped leaflets when subjected to "prolonged" shading [167].
Disturbances: Crownvetch is often found on disturbed sites. During surveys of state forests, parks, recreation areas, and wildlife refuges from Oklahoma to Maryland, crownvetch was most common or restricted to disturbed sites that included roads, trails, ditches, utility corridors, parking areas, and abandoned fields [10,45,50,65,70,126]. In Pennsylvania, a single crownvetch plant appeared in 1905 in an alfalfa field and persisted in the field even with repeated plowing and cultivation [39].
Grazing: Several reviews indicate that crownvetch is reduced or eliminated by grazing [120,133,152]. Three seasons of continuous grazing "deteriorates" crownvetch stands [120], and stands established on coal mine spoils can be "weakened or lost if overgrazed" [152]. In north-central Romania, researchers compared continuously grazed and ungrazed feather grass (Stipa spp.) stands. Crownvetch was characteristic of sites that were ungrazed for 29 to 57 years. Grazed sites were used by both sheep and cattle from spring to fall [30]. In field experiments conducted near Blacksburg, Virginia, clipping at 7- and 14-day intervals for 2 growing seasons reduced crownvetch cover from 90% to 10% [167]. In the Bohemian Karst Protected Region of Czech Republic, crownvetch biomass increased on sites mowed once and sites mowed repeatedly [9], but Vogel [152] indicates in a revegetation guide that crownvetch stands can be "weakened or lost" if cut more than 1 time/year for hay making.
Field observations and studies in Indiana indicate that soil compaction and not biomass removal limits crownvetch growth on grazed sites. Crownvetch successfully invaded adjacent ungrazed tall fescue (Schedonorus phoenix) stands but failed to invade adjacent grazed tall fescue stands. In crownvetch-mixed grass stands that were grazed for 3 years, crownvetch plants were thin, "lacked vigor", and failed to spread. In stands that were mowed and baled for hay, crownvetch plants had "excellent vigor of growth and spreading qualities" [46].
The scientific name of crownvetch is Securigera varia (L.) Lassen (Fabaceae) [60,78,141,159].
There are at least 3 common crownvetch cultivars: 'Penngift', 'Chemung', and 'Emerald'. These cultivars are essentially ecotypes that developed with 20 or more years of natural selection at different sites [92].
Crownvetch produces extensive vegetative growth from multibranched rhizomes (weed and forage management handbooks by [121,162]) and is capable of regenerating from stem and rhizome fragments [85]. Once established, crownvetch plants can form a dense mat of vegetation that "gradually chokes out" other vegetation [162]. A weed management guide reports that crownvetch rhizomes can reach 10 feet (3 m) long, and a single plant may cover 750 to 1,100 feet² (70-100 m²) in 4 years (review by [119]). On a cut slope in West Virginia, an 8-month-old crownvetch plant had rhizomes over 7 feet (2 m) long that supported 87 stems. On this site, soils were moist, loose, and had "ample" phosphorus [100]. Field observations made in central Bohemia, Czech Republic, indicated that within a year, vegetative offspring rarely occurred more than 1.5 feet (0.5 m) from the parent crownvetch plant [103].
Photo © Mandy TuIncreased vegetative growth can limit crownvetch seed production. When crownvetch produces abundant vegetative growth as seed is maturing, seed pods are shaded, seed development and maturity are delayed, and seed production is ultimately reduced. This is most common in high moisture years or areas [111].
In some areas, crownvetch persistence and spread is heavily dependent on vegetative regeneration. In Iowa, no crownvetch seedlings emerged from soil collected in crownvetch patches, but all 0.8- to 4-inch-long (2-10 cm) stem fragments with a node produced new growth. Aboveground stem segments that received water only once a week produced new growth but died within 3 weeks. Stem sections without nodes did not produce new growth but were green after a month in well-watered pots [84,85]. The researcher concluded that in this area, crownvetch persistence and spread were due to vegetative growth and regeneration [85].
La coroniella rosa (Securigera varia) ye una planta herbal de la familia de les fabacees.
Yerba viviega de 0,2-06 m, bien ramificada, difusu-ascendente, endurecida na so base y multicaule con tarmos fistulosos. Fueyes alternes estremaes n'hasta 12 pareyes de foliolos oblongos o elípticos. Flores papilionácees de color ablancazáu, rosáu o púrpura que tiende al azul, dispuestes en grupos de 10 a 12 flores.
Distribución submediterránea. Forma parte de les orles herbales de montes caducifolios submediterráneos y yerbazales mesófilos que colonicen les escamplaes d'esti tipu de montes, anque n'ocasiones apaez naturalizada en dellos puntos de la mariña, en yerbazales desenvueltos sobre suelos fondos, húmedos y subnitrificaos, en terremplénes, veres d'acequies y marxes de campos de regadío, ente 100 y 1.300 m.
Cardiotónico d'usu llindáu a prescripción facultativa. Les sos granes contienen un heterósido (ircanósido) que'l so aglicón (ircanogenina), una anhidroestrofantidina, amosó amás d'una aición cardiotónica, un marcáu efeutu antitumoral. Dosis: Usu so prescripción y control médicos.[1]
Esta páxina forma parte del wikiproyeutu Botánica, un esfuerciu collaborativu col fin d'ameyorar y organizar tolos conteníos rellacionaos con esti tema. Visita la páxina d'alderique del proyeutu pa collaborar y facer entrugues o suxerencies. La coroniella rosa (Securigera varia) ye una planta herbal de la familia de les fabacees.
Ala acıyonca (lat. Coronilla varia)[1] - acıyonca cinsinə aid bitki növü.[2]
Coronilla varia L. = Securigera varia Hündürlüyü 80 sm-ə qədər, gövdəsi zəif, əyilib qalxan və ya düz, uzun, tin-tin və içi boş, çılpaq
və ya tüklü, adətən budaqlanan çoxillik ot bitkisidir. Yalançı zoğları sərbəst olub, kiçikdir, xətvarineştərşəkilidir və uzunluğu 3 mm-dir [3].
Yarpaqları tək lələkvaridir, qısa saplaq üzərində 15-25 ədəd oval və ya uzunsov, küt yarpaqcıqlar yerləşmişdir [3].
Çiçəkləri 12-20 ədəd olmaqla, çiçək oxunun uzunluğu 3-15 sm-ə qədər olan, qısa başcığaoxşar çətirlərdə toplanmışdır; çiçək saplağı kasacıqdan iki dəfə uzun, ona bərabərvə ya ondan qısadır. Tac çəhrayı və ya bənövşəyi rəngdə olub, ağ zolaqlıdır [3].
Paxlaları qəhvəyi rəngdədir, düz və ya qövsşəkilli əyilmişdir, 1-10 ədəd yastı-dördtilli buğumlardan ibarətdir. Toxumları uzunsovovaldır, qəhvəyi və ya qonur rəngdədir [3].
May[3].
May-İyul[3].
Azərbaycanın bütün rayonlarında. Ovalıqdan, yuxarı dağ qurşağına qədər [3].
Çəmənliklərdə, meşə kənarında, kolluqlarda və bağlarda rast gəlinir[3].
Yaşıl yarpaq və gövdəsində aşı maddəsi vardır. Xalq təbabətində istifadə olunur. Cavan yarpaq və gövdəsi zəhərli olur[3].
İkiləpəlilər ilə əlaqədar bu məqalə qaralama halındadır. Məqaləni redaktə edərək Vikipediyanı zənginləşdirin. Ala acıyonca (lat. Coronilla varia) - acıyonca cinsinə aid bitki növü.
Coronilla varia L. = Securigera varia Hündürlüyü 80 sm-ə qədər, gövdəsi zəif, əyilib qalxan və ya düz, uzun, tin-tin və içi boş, çılpaq
və ya tüklü, adətən budaqlanan çoxillik ot bitkisidir. Yalançı zoğları sərbəst olub, kiçikdir, xətvarineştərşəkilidir və uzunluğu 3 mm-dir .
La coroneta rosa, coronil·la rosa, carolina o canari (Securigera varia) és una planta amb flor de la família de les Fabòidies.
Es troba a l'Europa meridional a vores de camins i herbassars xeròfils. A l'Amèrica del Nord es considera una espècie invasora.
És una planta menuda amb flors rosades. Floreix del juny al setembre. Les coronetes roses s'utilitzen en alguns llocs per controlar l'erosió del sól.[1]
Les fulles de la coroneta rosa són l'aliment principal de les erugues d'algunes papallones conegudes com a blavetes.
La coroneta rosa, coronil·la rosa, carolina o canari (Securigera varia) és una planta amb flor de la família de les Fabòidies.
Llysieuyn blodeuol (neu legume) yw Ffacbysen goronog sy'n enw benywaidd. Mae'n perthyn i'r teulu Fabaceae. Yr enw gwyddonol (Lladin) yw Securigera varia a'r enw Saesneg yw Crown vetch.[1]
Eraill yn yr un teulu yw: ffa soya (Glycine max), y ffa cyffredin (Phaseolus), pys gyffredin (Pisum sativum), chickpea (Cicer arietinum), cnau mwnci (Arachis hypogaea), pys per (Lathyrus odoratus) a licrs (Glycyrrhiza glabra).
Llysieuyn blodeuol (neu legume) yw Ffacbysen goronog sy'n enw benywaidd. Mae'n perthyn i'r teulu Fabaceae. Yr enw gwyddonol (Lladin) yw Securigera varia a'r enw Saesneg yw Crown vetch.
Eraill yn yr un teulu yw: ffa soya (Glycine max), y ffa cyffredin (Phaseolus), pys gyffredin (Pisum sativum), chickpea (Cicer arietinum), cnau mwnci (Arachis hypogaea), pys per (Lathyrus odoratus) a licrs (Glycyrrhiza glabra).
Čičorečka pestrá[1] (Securigera varia), česky též čičorka pestrá, patří mezi vytrvalé byliny z čeledi bobovitých a je okrajovou barevnou jetelovinou.
Čičorečka má vystoupavou či poléhavou lodyhu, obvykle 50 až 100 cm dlouhou (výjimečně až 200 cm). Hranatý stonek je dutý, rýhovaný a větvený. Lichozpeřeně dělené listy mají 6 až 12 párů krátce řapíkatých lístků. Květy jsou uspořádané do okolíku a vyrůstající z paždí listů v počtu 10 až 20. Kvete v květnu až září. Plodem je struk. Má hluboký kůlový kořen a odnožuje kořenovými výhony.
Čičorečka je rozšířena po celé Evropě s výjimkou Skandinávie a Velké Británie, kam byla jen druhotně zavlečena podobně jako do Severní Ameriky. Dále se vyskytuje v zemích jihovýchodní a střední Asie. V České republice je čičorečka rozšířena od nížin až po podhorské oblasti, ve vyšších polohách se vyskytuje zřídka.
Čičorečka pestrá se přednostně vyskytuje na výživných a vápencových půdách. Nejčastěji ji můžeme najít na suchých či mírně vlhkých loukách, travnatých stráních, mezích, železničních náspech a na okrajích cest.
Čičorečka pestrá má význam jako doplňková pícnina, obohacující krmivo rostlinnými bílkovinami. Přestože je mírně jedovatá, nejsou dokumentovány otravy u koní, skotu ani ovcí. U monogastrických zvířat může však být ve větším množství nebezpečná. Vysévá se koncem dubna na šířku řádků 12,5 až 25 cm, výsevek okolo 25 kg/ha. Semeno mahagonové barvy je podlouhlé a válečkovité. Výnosy suché hmoty činí v užitkovém roce 9 – 10 t/ha, se spalným teplem 18,896 MJ/kg, průměrný výnos semene je 200 – 300 kg/ha.
Čičorečka pestrá obsahuje přibližně 28 % vlákniny, 23 % dusíkatých látek, 8 – 9 % popele, 3 % tuku a okolo 37 % bezdusíkatých látek z absolutní sušiny. Porost lze využívat až čtyři užitkové roky.
Pěstování čičorečky pestré je v zemích Evropské unie povoleno. Podle nařízení vlády č. 86/2001 Sb. ve znění pozdějších předpisů lze dostat na pěstování této plodiny dotaci. Podmínky pro udělení této dotace jsou uvedeny v Praktické příručce MZe, č. 58/2005 „Zásady, kterými se stanovují podmínky pro poskytování dotací pro r. 2005 na základě §2 a §2d zákona č. 252/1997 Sb., o zemědělství“.
Mimo pícninářství může být využita jako energetická bylina.
V minulosti se sušená nať čičorečky Herba coronillae variae používala jako kardiotonikum vzhledem k obsahu koronillinu.
Čičorečka pestrá byla v roce 1989 přeřazena z rodu Coronilla Linné, 1753 do rodu Securigera de Candolle, 1805. Toto nové taxonomické zařazení je v současnosti běžně akceptováno. V české botanické nomenklatuře se po nějaký čas používalo původní rodové jméno čičorka i pro rod Securidaca (např. v Klíči ke květeně ČR), v roce 2012 se v Botanickém slovníku k odlišení objevuje nové české rodové jméno čičorečka.[2][1]
Obrázky, zvuky či videa k tématu čičorka pestrá ve Wikimedia Commons
Čičorečka pestrá (Securigera varia), česky též čičorka pestrá, patří mezi vytrvalé byliny z čeledi bobovitých a je okrajovou barevnou jetelovinou.
Die Bunte Kronwicke[1] (Securigera varia (L.) Lassen, Syn.: Coronilla varia L.) ist eine Pflanzenart aus der Gattung Securigera innerhalb der Unterfamilie der Schmetterlingsblütler (Faboideae). Sie wird auch Bunte Beilwicke[2] genannt.
Die Bunte Kronwicke ist eine ausdauernde krautige Pflanze.[3] Die niederliegenden bis aufsteigenden Stängel sind kantig und 30 bis 60, selten bis zu 120 Zentimeter lang.[1]
Die wechselständig angeordneten Laubblätter sind in kurzen Blattstiel und Blattspreite gegliedert. Die unpaarig gefiederte Blattspreite enthält vier bis, meist sechs bis zwölf Paare von Fiederblättchen. Die Fiederblättchen sind bei einer Länge von 0,5 bis 2 Zentimetern länglich-eiförmig und enden in Spitzchen.[1] Es sind Nebenblätter vorhanden.
Die Blütezeit reicht je nach Standort von Mai bis September, meist von Juni bis August[3]. Auf einem relativ langem Blütenstandsschaft sind 5 bis 20 Blüten sind in einem kopfigen, doldenförmigen Blütenstand angeordnet.[1] Eine Anzahl von 20 Blüten wurde schon von Linné bei der Erstveröffentlichung angegeben.[4]
Die zwittrige Blüte ist zygomorph und fünfzählig mit doppelter Blütenhülle. Der grüne Kelch ist weitglockig. Die rötlich-lilafarbene und weiße Blütenkrone[1] hat die typische Form eine Schmetterlingsblüte.[3] Die Fahne ist rosafarben, das Schiffchen weiß mit dunkelviolettem oberen Ende.[1] Die Flügel sind weiß.
Die aufrechte Hülsenfrucht ist bei einer Länge von 2,5 bis 5 (2 bis 8) Zentimetern schmal-linealisch, vierkantig mit hakig gebogenem Schnabel und durch drei bis sechs Einschnürungen schwach gegliedert.[1]
Die Chromosomengrundzahl beträgt x = 6 oder 8; es kommen Diploidie und Tetraploidie mit Chromosomenzahlen von 2n = 16 oder 24 vor.[3][1][5]
Die Bunte Kronwicke ist ein ausdauernder Hemikryptophyt[3] und eine Schaftpflanze. Sie ist erst ab dem 2. Jahr blühfähig. Die Blätter zeigen eine Nyktinastie: Sie werden zur Nacht hin in eine "Schlafstellung" nach oben geklappt. Die Bunte Kronwicke ist ein Tiefwurzler und bildet Wurzelknöllchen durch Symbiose mit Stickstoff bindenden Bakterien der Gattung Bradyrhizobium. Die Bunte Kronwicke bildet bis in 90 Zentimeter Bodentiefe ein ausgedehntes Wurzelsystem und wird zu den Rohboden-Pionierpflanzen gerechnet. Vegetative Vermehrung erfolgt durch wurzelbürtige Sprosse, also durch Wurzelbrut.
Die Blüten sind „Pollen-Schmetterlingsblumen“ mit Pumpeinrichtung. Im Gegensatz zu anderen Fabaceen mit einem freien Staubblatt wird in der Blüte kein Nektar abgesondert, wohl aber an der fleischigen Außenseite des Kelches. Bestäuber sind Honigbienen und andere Hautflügler. Die Blüten sind selbststeril.[3]
Die Früchte sind bei der Reife senkrecht stehende Bruchfrüchte. Sie sind zwischen den Samen durch sekundäre Trennwände eingeschnürt und zerfallen in vier bis zehn einsamige, nussartige, 4 bis 5 mg schwere Glieder. Es handelt sich hier also um Bruchhülsen. Die spezifisch leichten Teilfrüchte können herunterfallen und eine Schwerkraftausbreitung bewirken oder sie werden durch starke Winde fortgetragen. Auch eine Ausbreitung als Wasserhafter ist möglich.
Die Bunte Kronwicke ist in Mittel- und Südeuropa bis Westasien mit Schwerpunkt im nordöstlichen Mittelmeerraum und der Balkanhalbinsel verbreitet. In West- und Nordeuropa wird sie zum Teil als Futterpflanze kultiviert und ist eingebürgert. In Österreich ist sie in allen Bundesländern von der collinen bis montanen Höhenstufe häufig.
Diese kalkliebende Pflanzenart gedeiht in Mitteleuropa in trockenen Wiesen, Wald- und Gebüschsäumen, Rainen, Steinbrüchen und Bahndämmen. Sie gedeiht auf basenreichen, neutralen bis milden Böden und ist eine Charakterart der Ordnung Origanetalia, kommt aber auch in Pflanzengesellschaften der Verbände Mesobromion, Onopordion oder der Ordnung Agropyretalia vor.[5] In den Allgäuer Alpen steigt sie im Tiroler Teil im Gfällwald oberhalb Hägerau bis zu einer Höhenlage von 1400 Metern auf.[6]
Die ökologischen Zeigerwerte nach Landolt et al. 2010 sind in der Schweiz: Feuchtezahl F = 2+w (frisch aber wechselnd), Lichtzahl L = 3 (halbschattig), Reaktionszahl R = 4 (neutral bis basisch), Temperaturzahl T = 4 (kollin), Nährstoffzahl N = 2 (nährstoffarm), Kontinentalitätszahl K = 4 (subkontinental).[1]
Die Erstveröffentlichung erfolgte 1753 unter dem Namen (Basionym) Coronilla varia durch Carl von Linné in Species Plantarum, Tomus 2, S. 743.[4] Die Neukombination zu Securigera varia (L.) Lassen wurde 1989 durch Per Lassen in Svensk Botanisk Tidskrift, Band 83, S. 86 veröffentlicht. Synonyme Securigera varia (L.) Lassensind Coronilla haussknechtii Boiss. und Coronilla hirta Boiss.[7]
Diese Art wurde früher der Gattung Coronilla zugeordnet. Um die verwandtschaftliche Beziehung besser darzustellen, wurden einige Arten im letzten Jahrhundert der monophyletischen Gattung Securigera zugeordnet.
Alle Pflanzenteile, davon besonders die Samen, sind giftig. Hauptwirkstoffe sind Coronilla-Glykoside mit digitalisartiger Wirkung und Psoralen.
Für die Bunte Kronwicke bestehen bzw. bestanden auch die weiteren deutschsprachigen Trivialnamen Beilkraut (Schlesien), Giftwicki (Schweiz), Klaft (Österreich), Kronwicke (Schlesien, Schwaben), Peltschen (Schwaben), Schaflinse (Schlesien, Bern), Falsche Sparsette (Schweiz) und Bunte Vogelswicken.[8]
Die Bunte Kronwicke (Securigera varia (L.) Lassen, Syn.: Coronilla varia L.) ist eine Pflanzenart aus der Gattung Securigera innerhalb der Unterfamilie der Schmetterlingsblütler (Faboideae). Sie wird auch Bunte Beilwicke genannt.
Securigera varia (synonym Coronilla varia), commonly known as crownvetch[2] or purple crown vetch, is a low-growing legume vine. It is native to Africa, Asia and Europe and is commonly used throughout the United States and Canada for erosion control, roadside planting and soil rehabilitation. It has become an invasive species in many states of the US.
Crownvetch grows 1 to 2 feet tall and bears small clusters of 1/2-inch pink and white flowers from early summer to late fall. The variety Penngift produces mounds of pink flowers. Crownvetch is a tough, aggressive spreading plant that will crowd out its neighbors in a show garden but is well suited to a sunny bank, where it will grow for decades with little to no fertilizing, mowing, or weeding since the thick foliage prevents weeds from growing amid it.[3] Its deep, tenacious, complex root system[4] and thick, fern-like leaves provide excellent erosion control where it is used as a ground cover. However, this legume has a very long germination period of about 6 months and does not create full coverage until two or three years later.[4]
Crownvetch is toxic to horses and other non-ruminants because of the presence of nitroglycosides. If consumed in large amounts, it can cause slow growth, paralysis, or even death. However, this is not true for ruminant animals such as cattle, goats, and sheep.[5] These aliphatic nitro compounds are degraded in ruminant digestion and do not affect the animals.[5]
Crownvetch was originally labeled as a poor forage for farm animals but more recently it has been deemed as a good forage when fed as hay to or grazed by ruminants.[6] It consists of 21.7% crude protein and 22.2% crude fibre.[7] In an experiment involving the digestibility of crownvetch in sheep, it was found that the crude protein is 65.6% and the crude fibre is 46.2% digestible which is similar, but slightly less than the digestibility of alfalfa forage.[7] As previously mentioned, this only applies to ruminant animals as it is toxic to non-ruminants.[5]
Crownvetch has been identified as an invasive in several midwestern states. It is very hard to eradicate once established.[8]
It is also a common host-plant for the moth Zygaena ephialtes.[9]
Securigera varia (synonym Coronilla varia), commonly known as crownvetch or purple crown vetch, is a low-growing legume vine. It is native to Africa, Asia and Europe and is commonly used throughout the United States and Canada for erosion control, roadside planting and soil rehabilitation. It has become an invasive species in many states of the US.
Crownvetch planted to prevent erosion 
A field with crownvetch Crownvetch grows 1 to 2 feet tall and bears small clusters of 1/2-inch pink and white flowers from early summer to late fall. The variety Penngift produces mounds of pink flowers. Crownvetch is a tough, aggressive spreading plant that will crowd out its neighbors in a show garden but is well suited to a sunny bank, where it will grow for decades with little to no fertilizing, mowing, or weeding since the thick foliage prevents weeds from growing amid it. Its deep, tenacious, complex root system and thick, fern-like leaves provide excellent erosion control where it is used as a ground cover. However, this legume has a very long germination period of about 6 months and does not create full coverage until two or three years later.
La coronilla rosa (Securigera varia) es una planta herbácea de la familia de las fabáceas.
Hierba vivaz de 0,2-06 m, muy ramificada, difuso-ascendente, endurecida en su base y multicaule con tallos fistulosos. Hojas alternas divididas en hasta 12 parejas de folíolos oblongos o elípticos. Flores papilionáceas de color blanquecino, rosado o púrpura que tiende al azul, dispuestas en grupos de 10 a 12 flores.
Distribución submediterránea. Forma parte de las orlas herbáceas de bosques caducifolios submediterráneos y herbazales mesófilos que colonizan los claros de este tipo de bosques, aunque en ocasiones aparece naturalizada en algunos puntos del litoral, en herbazales desarrollados sobre suelos profundos, húmedos y subnitrificados, en terraplenes, orillas de acequias y márgenes de campos de regadío, entre 100 y 1.300 m.
Cardiotónico de uso limitado a prescripción facultativa. Sus semillas contienen un heterósido (ircanósido) cuyo aglicón (ircanogenina), una anhidroestrofantidina, ha mostrado además de una acción cardiotónica, un marcado efecto antitumoral. Dosis: Utilización bajo prescripción y control médicos.[1]
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Datos: Q158908
Multimedia:
Especies: Securigera varia
Kirju sarikhernes (Coronilla varia) on liblikõieliste sugukonda kuuluv taimeliik.
Taim on kantud Eesti ohustatud liikide punasesse nimestikku.[1]
Kirju sarikhernes (Coronilla varia) on liblikõieliste sugukonda kuuluv taimeliik.
Taim on kantud Eesti ohustatud liikide punasesse nimestikku.
Kirjokärsäpalko eli kirjonivelvirna (Securigera varia) on monivuotinen hernekasvilaji, jota esiintyy Euroopassa. Sen rento varsi kasvaa 40:stä sataan senttimetriä pitkäksi. Lehdissä on viidestä kymmeneen paria soikeita lehdyköitä. Kukinto on 5–20 kukasta koostuva pallomainen sarja. Teriöt ovat kirjavia, terälehdistä purje on tavallisesti vaaleanpunainen, venho ja siivet valkoisia ja venhon kärki tummanvioletti. Palko on kolmesta viiteen senttimetriä, nelisärmäinen ja katkeilee nivelkohdista.
Kirjokärsäpalko eli kirjonivelvirna (Securigera varia) on monivuotinen hernekasvilaji, jota esiintyy Euroopassa. Sen rento varsi kasvaa 40:stä sataan senttimetriä pitkäksi. Lehdissä on viidestä kymmeneen paria soikeita lehdyköitä. Kukinto on 5–20 kukasta koostuva pallomainen sarja. Teriöt ovat kirjavia, terälehdistä purje on tavallisesti vaaleanpunainen, venho ja siivet valkoisia ja venhon kärki tummanvioletti. Palko on kolmesta viiteen senttimetriä, nelisärmäinen ja katkeilee nivelkohdista.
Securigera varia
Securigera varia, la coronille bigarrée ou coronille changeante, est une espèce de plantes de la famille des Fabaceae, sous-famille des Faboideae, originaire d'Europe et d'Asie tempérée. Ce sont des plantes herbacées vivaces qui peuvent atteindre 60 cm de haut environ. La coronille bigarrée était utilisée en médecine traditionnelle pour ses propriétés cardiotoniques et diurétiques et pour soigner les maladies de la prostate. Toutefois, il fallait l'utiliser avec de grandes précautions à cause de sa toxicité[1].
La plante a été introduite dans de nombreux pays, en particulier aux États-Unis, où elle est utilisée à diverses fins : plante ornementale, plante de couverture, engrais vert, amélioration des sols et lutte contre l'érosion, plante fourragère (malgré sa toxicité spécialement pour les non-ruminants). Elle s'est révélée être une plante envahissante dans certaines régions[2].
Securigera varia est une plante herbacée vivace, essentiellement glabre, qui pousse normalement à une hauteur de 60 cm environ ou plus en floraison, voire seulement de 30 cm à l'état végétatif. La plante a un système racinaire de type pivotant, profond et très ramifié Elle produit aussi de forts rhizomes charnus qui peuvent atteindre 3 mètres de long voire plus. Les tiges, grossières, relativement faibles et fortement ramifiées proviennent d'un caudex court et ramifié. Dépourvues de vrilles, rampantes ou redressées, les tiges peuvent atteindre 2 mètres de long. La plante apparaît souvent sous la forme d'un épais tapis de tiges[5].
Les feuilles, de couleur vert foncé, de 4 à 16 cm de long, sont composées imparipennées, comptant généralement de 7 à 25 folioles. Celles-ci sont généralement oblongues, de forme elliptique à obovale, et mesurent de 1 à 3 cm de long. En hiver, les feuilles basales et parfois supérieures restent vertes[6],[7],[5].
Les fleurs, hermaphrodites, sont groupées en ombelles simples composées de 6 à 25 fleurs et portées par un long pédoncule. Elles ont une corolle de type papilionacé de 10 à 12 mm de long, de couleur variant du blanc-rosâtre au rose foncé et comptent 10 étamines.
Les fruits sont des gousses minces, linéaires, segmentées (lomentacées), pointues, groupées en ombelles. Les gousses peuvent atteindre 6 cm de long et compter jusqu'à 12 segments contenant chacun une seule graine. Elles sont déhiscentes lorsqu'elles sont sèches. Les graines sont cylindriques et mesurent 3 à 4 mm sur 1 à 1,2 mm[6],[7],[5].
Port de la plante.
Feuille.
Inflorescence.
Fleur.
Fruits.
Securigera varia est une espèce tétraploïde comptant 24 chromosomes (2n = 4x = 24)[8].
Coronilla varia contient diverses substances phytochimiques.
Des hydroxycoumarines (scopolétine, daphnorétine, ombelliférone) et des glycosides cardiotoniques (coronilloside[9], déglucohyrcanoside, hyrcanoside) ont été isolés des graines[10].
La plante contient une neurotoxine, l'acide β-nitropropionique, principalement dans les feuilles et dans les tiges en croissance[11].
Des flavonoïdes ont été trouvés dans les feuilles. Les inflorescences contiennent également de la cytisine[9], du kaempférol et de l'astragaline. Deux composés isoflavones ont été isolés de Coronilla varia, 1,10 μg/ml de génistine et 0,34 μg/ml de génistéine. La plante a une faible teneur en saponines, teneur plus élevée dans les feuilles que dans les tiges. La concentration de tanins condensés (proanthocyanidines) s'élève à 16,0 g/kg de poids sec[10].
Dans la classification des types biologiques de Raunkier, Securigera varia est une plante chamaephyte et hémicryptophyte[5].
La plante se reproduit par les graines et par les rhizomes, ces derniers jouant un rôle important dans la croissance et la persistance de l'espèce. La période de floraison s'étend sur le printemps et l'été (de mai à septembre dans l'hémisphère nord)[5].
La pollinisation est entomogame et dépend principalement des abeilles. La dissémination des graines est épizoochore.
L'aire de répartition originelle de Securigera varia s'étend dans les régions tempérées de l'Eurasie, incluant l'Europe méditerranéenne, l'Europe centrale et l'Europe de l'Est, ainsi que l'Asie mineure, la région du Caucase et le Moyen-Orient. Elle inclut en particulier les pays suivants : Albanie, Allemagne, Biélorussie, Bulgarie, Espagne, France, Grèce, Hongrie, Iran, Irak, Italie, Liban, Pologne, Roumanie, Russie d'Europe et Crimée, Slovaquie, Suisse, Syrie, Tchéquie, Turquie, Turkménistan, Ukraine, ex-Yougoslavie[12].
L'espèce a été introduite dans diverses régions : Europe du Nord et de l'Ouest (Belgique, Danemark, pays baltes, Pays-Bas, Norvège, Royaume-Uni, Russie du Nord, Sibérie occidentale, Suède), Amérique (Argentine, Canada, États-Unis, République dominicaine), Asie (Corée), Océanie (Australie, Nouvelle-Zélande)[12]. Il s'agit d'introductions le plus souvent délibérées, mais parfois accidentelles (introduction en Belgique dans des semences de céréales)[13],[12].
Dans son aire de répartition naturelle, l'espèce est présente dans les prairies, les pelouses sèches et les zones perturbées. Dans les régions où elle a été introduite, Securigera varia se trouve dans les prairies mésiques, les pâturages, les fossés de drainage, les bords de routes et remblais de chemin de fer, les jardins et les champs ouverts, les marges forestières, les dunes secondaires, les berges de rivières et bancs de gravier le long des cours d'eau[2].
Selon The Plant List (23 septembre 2020)[14] :
Securigera varia est utilisée comme cardiotonique, pour provoquer des vomissements et pour traiter les rhumatismes articulaires et les crampes. La prudence est recommandée en raison de la toxicité de la plante et des graines. Selon certains auteurs, la plante aurait des activités antitumorales et antibactériennes, ainsi que des propriétés antioxydantes dues à des composés flavonoïdes[2].
Considérée autrefois comme un fourrage médiocre pour les animaux d'élevage, la coronille bigarrée est un bon fourrage lorsqu'il est donné aux ruminants, qu'il soit distribué sous forme de foin ou brouté au pâturage[15]. Ce fourrage se compose de 21,7 % de protéines brutes et 22,2 % de fibres brutes[16]. Dans une expérience impliquant la digestibilité de la coronille bigarrée chez le mouton, on a constaté que la digestibilité des protéines brutes s'élevaient à 65,6 % et celle des fibres brutes à 46,2 %, valeurs similaires, quoique légèrement inférieures, à celle du fourrage de luzerne[16].
Cela ne s'applique cependant qu'aux ruminants, car cette plante est toxique pour les non-ruminants, notamment les chevaux, en raison de la présence de nitroglycosides. Consommée en grande quantité, elle peut entraîner un retard de croissance, une paralysie, voire la mort. Chez les ruminants, tels que les bovins, ovins et caprins, ces composés nitrés aliphatiques sont dégradés dans le tube digestif par les microorganismes du rumen et n'affectent pas la santé des animaux[17].
Aux États-Unis, Securigera varia a été largement plantée le long des routes et autoroutes, ainsi que dans les zones perturbées pour lutter contre l'érosion ou pour stabiliser les accotements routiers. On l'a aussi utilisée pour assurer la couverture du sol, la remise en valeur des terrains miniers abandonnés et comme culture intercalaire, car par son association avec des cyanobactéries, cette plante enrichit le sol en azote[4].
Securigera varia
Securigera varia, la coronille bigarrée ou coronille changeante, est une espèce de plantes de la famille des Fabaceae, sous-famille des Faboideae, originaire d'Europe et d'Asie tempérée. Ce sont des plantes herbacées vivaces qui peuvent atteindre 60 cm de haut environ. La coronille bigarrée était utilisée en médecine traditionnelle pour ses propriétés cardiotoniques et diurétiques et pour soigner les maladies de la prostate. Toutefois, il fallait l'utiliser avec de grandes précautions à cause de sa toxicité.
La plante a été introduite dans de nombreux pays, en particulier aux États-Unis, où elle est utilisée à diverses fins : plante ornementale, plante de couverture, engrais vert, amélioration des sols et lutte contre l'érosion, plante fourragère (malgré sa toxicité spécialement pour les non-ruminants). Elle s'est révélée être une plante envahissante dans certaines régions.
Pisana čećorica (Securigera varia) je rostlina ze swójby łušćinowcow. Wona bu prjedy rodej Coronilla přirjadowana.
Pisana čećorica je wjacelětna, zelišćowa rostlina. Wona docpěwa wysokosć wot 80 cm.
Stołpiki su hranite.
Lisćowe łopjena su pjerite a z krótkim stołpikom.
Rostlina kćěje wot meje hač do oktobra. Wokołk wobsahuje maksimalnje 20 kćenjow.
Łušćinowy płód docpěwa dołhosć wot 25 hač do 50 mm. Wón je štyrihranity z hokatim pyskom.
Wona rosće na suchich łukach, pućnych kromach, pod kerčinami a na ladźe; na wapnu. Rostlina preferuje wapnojtu pódu.
Pisana čećorica je w nimale cyłej Europje, w Kawkazu a Małoaziji rozšěrjena.
Pisana čećorica je picowanska rostlina.
Pisana čećorica (Securigera varia) je rostlina ze swójby łušćinowcow. Wona bu prjedy rodej Coronilla přirjadowana.
Securigera varia (L.) Lassen, 1989 è una pianta erbacea appartenente alla famiglia delle Leguminose o Fabaceae.[1] Da non confondere assolutamente con un'altra pianta della stessa famiglia, Bituminaria bituminosa. La differenza è da ricercare anche nella forma e nel colore delle foglie.
Pianta perenne, alta 30–70 cm; fusti prostrati - diffusi, con rami ascendenti, glabri angolosi e fistolosi lunghi da 30 cm. fino a 1,5 m. ; foglie con 17-23 segmenti, ellittici; corone 10-20 fiore calice corto, conico; corolla rosea, più colorita sul grande petalo centrale superiore (vessillo). Fiorisce tra giugno ed agosto.
Pianta comune nelle radure erbose e nei boschi radi fino a 1500 m.
È considerata una specie invasiva in molti stati degli Stati Uniti d'America.
È annoverata tra le piante velenose per la presenza in tutte le sue parti di un glucoside tossico, la catartina.
Securigera varia (L.) Lassen, 1989 è una pianta erbacea appartenente alla famiglia delle Leguminose o Fabaceae. Da non confondere assolutamente con un'altra pianta della stessa famiglia, Bituminaria bituminosa. La differenza è da ricercare anche nella forma e nel colore delle foglie.
Dvispalvis raženis (lot. Securigera varia, sin. Coronilla varia) – pupinių (Fabaceae) šeimos raženių (Securigera) genties augalų rūšis.
Daugiametis, apie 80 cm aukščio augalas. Šaknis gana ilga, sumedėjusi. Lapai neporiškai plunksniški. Stiebai tuščiaviduriai, kylantys. Žiedai sutelkti kekėse. Vainikėlis rožinis, rečiau baltas. Žydi birželio – rugpjūčio mėn. Apdulkina bitės. Ankštys stačios, apie 5,5 cm ilgio, rudos spalvos su 3-6 viensėkliais narveliais. Sėklos ovalinės, rudos.
Vienintelė Lietuvoje auganti rūšis. Auga sausuose, karbonatinguose dirvožemiuose.
Tai nuodingas, taip pat ir vaistinis, bei medingas augalas.
Sėklose yra nuodingo gliukozido, todėl suėdę žolę galvijai ir arkliai suserga.
Bont kroonkruid of kroonwikke (Securigera varia, synoniem: Coronilla varia) is een vaste plant die behoort tot de vlinderbloemenfamilie (Leguminosae). De plant komt van nature voor in Midden- en Zuid-Europa en is van daaruit verbreid naar West- en Noord-Europa, omdat de plant daar als voedergewas wordt geteeld. Ook wordt de plant geteeld in de Verenigde Staten en Canada voor bodemverbetering, het tegengaan van erosie en als bermbeplanting. De plant is giftig voor paarden door de aanwezigheid van nitroglycosiden.
De plant wordt 30-120 cm hoog en heeft een liggende tot opstijgende, kantige stengel. De bladeren zijn oneven geveerd met zeven tot twaalf paar deelblaadjes en een topblaadje. De ovale blaadjes zijn 6-16 mm lang.
Bont kroonkruid bloeit van juni tot september. De bloemsteel is anderhalf tot drie keer zo lang als de kelkbuis. De kroonbladen zijn 10-15 mm lang. De vlag van de bloem is roze en de zwaarden zijn vaak wit. De kiel is roze of wit en heeft een paarsrode snavel. Soms komen ook geheel witte bloemen voor. De bloeiwijze is een scherm met vijf tot twintig bloemen.
De vrucht is een 25-50 mm lange, vierkantige peul met drie tot zes insnoeringen en een hakig gebogen snavel. De rijpe peul breekt op de insnoeringen in stukjes.
De plant komt voor op matig vochtige, kalkrijke grond op dijkhellingen, spoordijken, in bermen en duinen.
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Bont kroonkruid of kroonwikke (Securigera varia, synoniem: Coronilla varia) is een vaste plant die behoort tot de vlinderbloemenfamilie (Leguminosae). De plant komt van nature voor in Midden- en Zuid-Europa en is van daaruit verbreid naar West- en Noord-Europa, omdat de plant daar als voedergewas wordt geteeld. Ook wordt de plant geteeld in de Verenigde Staten en Canada voor bodemverbetering, het tegengaan van erosie en als bermbeplanting. De plant is giftig voor paarden door de aanwezigheid van nitroglycosiden.
De plant wordt 30-120 cm hoog en heeft een liggende tot opstijgende, kantige stengel. De bladeren zijn oneven geveerd met zeven tot twaalf paar deelblaadjes en een topblaadje. De ovale blaadjes zijn 6-16 mm lang.
Bont kroonkruid bloeit van juni tot september. De bloemsteel is anderhalf tot drie keer zo lang als de kelkbuis. De kroonbladen zijn 10-15 mm lang. De vlag van de bloem is roze en de zwaarden zijn vaak wit. De kiel is roze of wit en heeft een paarsrode snavel. Soms komen ook geheel witte bloemen voor. De bloeiwijze is een scherm met vijf tot twintig bloemen.
De vrucht is een 25-50 mm lange, vierkantige peul met drie tot zes insnoeringen en een hakig gebogen snavel. De rijpe peul breekt op de insnoeringen in stukjes.
De plant komt voor op matig vochtige, kalkrijke grond op dijkhellingen, spoordijken, in bermen en duinen.
Cieciorka pstra (Securigera varia) – gatunek rośliny z rodziny bobowatych (Fabaceae). Gatunek tradycyjnie zaliczany do rodzaju cieciorka (Coronilla), po rewizji ujęć taksonomicznej od 1989 roku gatunek ten włączany jest do rodzaju Securigera[3][2][4]. Pochodzi z Azji i Europy, ale rozprzestrzenił się także w Nowej Zelandii i Północnej Ameryce[4]. Jest również uprawiany w wielu krajach świata. W Polsce na stanowiskach naturalnych jest dość pospolity[5].
Bylina, hemikryptofit. Kwitnie od maja do lipca. Najłatwiej spotkać ją można na przydrożach, miedzach, słonecznych stokach wzgórz i suchych łąkach. W górach występuje po regiel górny. W klasyfikacji zbiorowisk roślinnych gatunek charakterystyczny dla Cl. Trifolio-Geranietea[6]. Roślina trująca.
Jest rośliną żywicielską larw motyli powszelatka brunatka, wietka gorczycznika, wietka Reala, szlaczkoni: erate, siarecznika, sylwetnika, południowca, oraz modraszków: srebroplamka, korydona, adonisa, aleksisa, dafnida, ikara, argusa[7].
Cieciorka pstra (Securigera varia) – gatunek rośliny z rodziny bobowatych (Fabaceae). Gatunek tradycyjnie zaliczany do rodzaju cieciorka (Coronilla), po rewizji ujęć taksonomicznej od 1989 roku gatunek ten włączany jest do rodzaju Securigera. Pochodzi z Azji i Europy, ale rozprzestrzenił się także w Nowej Zelandii i Północnej Ameryce. Jest również uprawiany w wielu krajach świata. W Polsce na stanowiskach naturalnych jest dość pospolity.
Ranostaj pestrý (Securigera varia) je rastlina z čeľade bôbovité (Fabaceae).
Commons ponúka multimediálne súbory na tému Ranostaj pestrý
Wikidruhy ponúkajú informácie na tému Ranostaj pestrý
Ranostaj pestrý (Securigera varia) je rastlina z čeľade bôbovité (Fabaceae).
Rosenkronill (Securigera varia) är en art i familjen ärtväxter. Den kommer ursprungligen från centrala och södra Europa och har inplanterats i USA och Kanada för att hejda jorderosion.
Rosenkronill är en flerårig ört. Blommorna är ljusrosa och sitter i flockar. De blommar mellan maj och september och trivs bäst i väldränerad jord i soliga lägen.
Wikimedia Commons har media som rör Rosenkronill.
Rosenkronill (Securigera varia) är en art i familjen ärtväxter. Den kommer ursprungligen från centrala och södra Europa och har inplanterats i USA och Kanada för att hejda jorderosion.
Rosenkronill är en flerårig ört. Blommorna är ljusrosa och sitter i flockar. De blommar mellan maj och september och trivs bäst i väldränerad jord i soliga lägen.
Рослина заввишки 30—10 см, з повзучим гіллястим кореневищем. Стебла дещо розпростерті або майже прямостоячі, гіллясті, голі, борозенчасті. Листки непарноперисті, черешок з 5—12 парами еліптичних або довгасто-лінійних листочків, на вершк загострених. Верхні листки сидячі, прилистки вільні, ланцетні. Квітки зібрані в зонтикоподібні китиці з квітконосами завдовжки 3—15 см, що іноді перевищують листки. Чашечка дзвоникоподібна, 5-зубчаста. Віночок з трох зігнутим дзьобоподібно розширеним човником. Тичинок 10, з яких 9 зрослися нитками в трубочку. Плоди — лінійні боби, 4—5 см завдовжки. Цвіте в травні — серпні; насіння достигає з липня.
Росте на луках, узліссях, по чагарниках, на схилах у великій кількості.
В медицині використовується насіння в'язеля, з якого виділено маловивчені серцеві глюкозиди: коронизид, корониллін. Крім того, насіння містить вуглевод стахіозу, жирне олію, насичені жирні кислоти, сечову кислоту, каротин, вітамін С (0,27 %).
Коронизид має вибіркову дію на серце, за характером і силою подібну до строфантину.
В народній медицині застосовується від водянки. Настій з насіння в'язеля діє подібно до настою наперстянки.
Securigera varia là một loài thực vật có hoa trong họ Đậu. Loài này được (L.) Lassen miêu tả khoa học đầu tiên.[1]
Bài viết liên quan đến tông đậu Loteae này vẫn còn sơ khai. Bạn có thể giúp Wikipedia bằng cách mở rộng nội dung để bài được hoàn chỉnh hơn.
Securigera varia là một loài thực vật có hoa trong họ Đậu. Loài này được (L.) Lassen miêu tả khoa học đầu tiên.
Родина растения — Южная Европа, Юго-Западная Азия и Северная Африка, в настоящее время распространилось по всей Европе. Инвазивный вид, очень быстро распространяющийся корневыми отпрысками и семенами. Завезено в Северную Америку в 1869 году, с 1950-х годов широко культивируется в США, на северо-востоке страны используется для стабилизации грунтов. В Европе и США также выращивается в качестве кормового растения. Натурализовался в Новой Зеландии.
Также выращивается в качестве декоративного садового растения, используется в медицине. Растение морозостойко до −33 °C, засухоустойчиво, переносит и застой воды, однако является безусловным гелиофитом.
Впервые действительно описано растение было Карлом Линнеем во 2-м томе Species plantarum. Датой публикации работы считается 1 мая 1753 года.
Родина растения — Южная Европа, Юго-Западная Азия и Северная Африка, в настоящее время распространилось по всей Европе. Инвазивный вид, очень быстро распространяющийся корневыми отпрысками и семенами. Завезено в Северную Америку в 1869 году, с 1950-х годов широко культивируется в США, на северо-востоке страны используется для стабилизации грунтов. В Европе и США также выращивается в качестве кормового растения. Натурализовался в Новой Зеландии.
Также выращивается в качестве декоративного садового растения, используется в медицине. Растение морозостойко до −33 °C, засухоустойчиво, переносит и застой воды, однако является безусловным гелиофитом.
多变小冠花（学名：Coronilla varia），又名绣球小冠花，是一种豆科植物。这种植物原产于地中海一带，在欧洲中部和南部，亚洲西南部和北非均有分布。[1]
这是一篇與植物相關的小作品。你可以通过编辑或修订扩充其内容。 多变小冠花（学名：Coronilla varia），又名绣球小冠花，是一种豆科植物。这种植物原产于地中海一带，在欧洲中部和南部，亚洲西南部和北非均有分布。
分類 界 : 植物界 Plantae 門 : 被子植物門 Magnoliophyta 綱 : 双子葉植物綱 Magnoliopsida 目 : マメ目 Fabales 科 : マメ科 Fabaceae 属 : Coronilla 種 : タマザキクサフジ C. varia 学名 Coronilla varia L. シノニム Coronilla varia
和名 タマザキクサフジ 英名 Trailing Crown Vetchタマザキクサフジ（玉咲草藤、学名：Securigera varia）は、マメ科に分類される多年草の一種。クラウンベッチの名称で栽培される。
北アメリカ、中近東、日本に外来種として定着している[1]。日本では1981年に札幌市で初めて帰化が確認された[1]。
茎は斜めもしくは匍匐して伸び、1mほどまで生長する。夏から秋に淡紅紫色の蝶形花をいくつも固まって咲かせる。葉は奇数羽状複葉となる。
牧草や道路法面に利用される。
タマザキクサフジ（玉咲草藤、学名：Securigera varia）は、マメ科に分類される多年草の一種。クラウンベッチの名称で栽培される。
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