dcsimg

Pantherophis ( Azerbaijano )

fornecido por wikipedia AZ

Pantherophis (lat. Pantherophis) - suilanıkimilər fəsiləsinə aid heyvan cinsi.

Növləri

Mənbə

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Pantherophis: Brief Summary ( Azerbaijano )

fornecido por wikipedia AZ

Pantherophis (lat. Pantherophis) - suilanıkimilər fəsiləsinə aid heyvan cinsi.

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Amerikanische Kletternattern ( Alemão )

fornecido por wikipedia DE

Die Amerikanischen Kletternattern (Pantherophis) (von gr. panthera = Leopard und ophis = Schlange) sind eine Gattung ungiftiger, nordamerikanischer Schlangen aus der Familie der Nattern (Colubridae).

Beschreibung und Verhalten

Die Arten der Gattung Pantherophis zeichnen sich durch einen schlanken Körperbau mit nur leicht abgesetztem Kopf aus. Sie erreichen Körperlängen zwischen 100 cm und 250 cm. Viele Arten sind ausgesprochene Kulturfolger und folglich häufig in der Nähe menschlicher Siedlungen zu finden, darüber hinaus sind einige Arten sehr aktive Kletterer und können mitunter auch in Bäumen angetroffen werden, was der Gattung den deutschen Trivialnamen einbrachte. Alle Arten der Gattung vermehren sich ovipar und betreiben keine Brutpflege.

Verbreitung und Lebensraum

Das Verbreitungsgebiet der Gattung erstreckt sich über weite Teile der USA und Mexikos. Die von den unterschiedlichen Arten bewohnten Lebensräume reichen von sommerfeuchten Laub- und Nadelwäldern über die Prärie bis hin zu felsigen Höhenlagen in über 2500 m Seehöhe.

Systematik

Im Jahr 2002 wurde die bis dahin gültige Gattung der Kletternattern (Elaphe) nach eingehenden wissenschaftlichen Untersuchungen[1] in mehrere Gattungen aufgespalten. Die bereits 1843 beschriebene Gattung Pantherophis wurde zu diesem Zweck wiedereingeführt. Diese Änderung ist weitgehend akzeptiert.[2]

2008 erfolgte erneut eine Änderung der Einteilungen durch Collins & Taggart. Dabei wurden die Erdnattern in die Gattung Scotophis und die Fuchsnattern in die neue Gattung Mintonius gestellt. Die Gattung Pantherophis würde nur noch aus den drei Kornnatter-Arten Pantherophis guttatus, Pantherophis emoryi und Pantherophis slowinksii bestehen.[3][4] Diese Änderung ist allerdings umstritten, da sie die Verwandtschaft mit den Erdnattern und den Fuchsnattern verschleiern würden. Pyron et al. übernehmen diese Einteilung nicht.[5] Dementsprechend werden die Arten der Erdnattern und Fuchsnattern weiterhin der Gattung Pantherophis zugeteilt:[6][2]

Gruppe der Kornnattern

Gruppe der Erdnattern

Gruppe der Fuchsnattern

Die nächstverwandte Gattung von Pantherophis ist Pituophis.[5]

Einzelnachweise

  1. Urs Utiger, Notker Helfenberger, Beat Schätti, Catherine Schmidt, Markus Ruf, Vincent Ziswiler: Molecular systematics and phylogeny of old and new world ratsnakes, Elaphe auct., and related genera. In: Russian Journal of Herpetology. Band 9, 2002, S. 105–124.
  2. a b Pantherophis In: The Reptile Database
  3. Philipp Berg: Änderungen der Taxonomie der Kletternattern (Elaphe)
  4. Joseph T. Collins, Travis W. Taggart: An alternative classification of the New World rat snakes (genus Pantherophis [Reptilia: Squamata: Colubridae]). In: Journal of Kansas Herpetology. Band 26, 2008, S. 16–18.
  5. a b R. Alexander Pyron, Frank T. Burbrink: Neogene diversification and taxonomic stability in the snake tribe Lampropeltini (Serpentes: Colubridae). In: Molecular Phylogenetics and Evolution. Band 52, 2009, S. 524–529.
  6. Sabrina Schwarzgruber: Eine Übersicht über den aktuellen taxonomischen Stand innerhalb der Gattung Pantherophis@1@2Vorlage:Toter Link/www.schlangengrube.de (Seite nicht mehr abrufbar, Suche in Webarchiven)  src= Info: Der Link wurde automatisch als defekt markiert. Bitte prüfe den Link gemäß Anleitung und entferne dann diesen Hinweis. 2012.
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Amerikanische Kletternattern: Brief Summary ( Alemão )

fornecido por wikipedia DE

Die Amerikanischen Kletternattern (Pantherophis) (von gr. panthera = Leopard und ophis = Schlange) sind eine Gattung ungiftiger, nordamerikanischer Schlangen aus der Familie der Nattern (Colubridae).

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Pantherophis ( Inglês )

fornecido por wikipedia EN

Pantherophis is a genus of nonvenomous colubrid snakes endemic to central and eastern regions of North America. It consists of the North American ratsnakes, the foxsnakes, and the cornsnakes. The genus, which contains 10 recognized species, first appeared in the fossil record in the Middle Miocene around 16.3 million years ago. They are a large terrestrial snake genus that lack subocular scales. Originally classified in the genus Elaphe, phylogenetic studies have found this taxon to be closely related to Pituophis. As with all snakes Pantherophis is an obligate faunivore with a diet that consists of small mammals, birds, reptiles and amphibians, and even insects. While many species conservation status is categorized as "least concern", many local populations in some species have declined where some places have them listed as federally protected. The corn snake (P. guttatus) is a popular pet reptile, due to the availability of captive-bred animals, their low maintenance and calm disposition, and the variety of color morphs. There are other species of Pantherophis that are in the pet trade, though are not as popular as the corn snake.

Field Characteristics

Member species in the genus Pantherophis are large terrestrial snakes, built for constriction. Pantherophis is characterized by having a divided cloacal plate. The dorsal scales are arranged in more than 30 rows around the body, and at least some of these are keeled, albeit faintly. All species lack subocular scales.[4] Each of the ten or so species can be further distinguished based on subtle characteristics, such as color markings, scale nuances, and geographic range.[4]

Systematics and Paleontology

Taxonomy

The taxonomy of Pantherophis has been a complicated area of research. The genus was named by the Austrian zoologist Leopold Fitzinger in 1843, with the type species being Pantherophis guttatus. Shortly after, the genus was then considered to be a junior synonym of the otherwise Old World genus Elaphe. However the morphology of the hemipenes, the dentition, and the squamatization found the New World species to be closer to the genus Pituophis (gopher snakes, pine snakes, and bullsnakes). Subsequent molecular studies have found support of the sister grouping between Pituophis and the North American Elaphe species, which resulted in the resurrection of the genus Pantherophis.[5] These two genera, along with several other North American endemic taxa like Lampropeltis belong to the tribe Lampropeltini.[6]

There are currently four subgenera of Pantherophis, of which three are extant and one is extinct: Scotophis Baird & Girard, 1853, for the ratsnakes; †Palaeoelaphe Gilmore, 1938, for a Miocene fossil species;[2] Mintonius Collins & Taggart, 2008, for the foxsnakes; and Pantherophis for the cornsnake complex. There are a few studies that suggested the possibility that Pantherophis is paraphyletic in respect to Pituophis.[7] To maintain taxonomic stability, some of the aforementioned subgenera are reevaluated into proper genera. This, however, has not been supported by larger scale molecular trees concerning the relationships of various snake taxa.[6][8][9]

Phylogeny

The interspecies relationships of Pantherophis usually has the subgenera Pantherophis and Mintonius being sister taxa, with Scotophis at the root of the genus. Below is the widely supported phylogenetic tree of the species in the genus.[6][8][9]

Pantherophis Scotophis

Pantherophis bairdi

Pantherophis obsoletus

Pantherophis alleghaniensis

Pantherophis spiloides

Mintonius

Pantherophis vulpinus

Pantherophis ramspotti

Pantherophis

Pantherophis guttatus

Pantherophis slowinskii

Pantherophis emoryi

Extant Species

Below is the list of valid species with their geographic distributions.[3][10]

Nota bene: In this list, a binomial authority in parentheses indicates that the species was originally described in a genus other than Pantherophis.

Fossil Remains

The fossil record of Pantherophis is the Middle Miocene to Upper Pleistocene.[2] In addition to fossil remains of some of the extant species, there are fragmentary Pantherophis remains throughout the second half of the Cenozoic in North America. Only three fossil species have been described from more complete remains. The Early Pliocene P. buisi which might be a basal species in the Scotophis subgenus,[1] and P. kansensis which is the sole species in the subgenus of Palaeoelaphe, the trunk vertebrae similar to Mintonius subgenus.[1] P. nebraskensis was another valid species, but it is considered to be a junior synonym of P. kansensis.[2]

References

  1. ^ a b c Holman, J.A. (2000). Fossil Snakes of North America: Origin, Evolution, Distribution, Paleoecology. Indiana University Press. ISBN 0253337216.
  2. ^ a b c d Wallach, V.; Williams, K.L.; Boundy, J. (2014). Snakes of the World: A Catalogue of Living and Extinct Species. Boca Raton, Florida: CRC Press. ISBN 978-1-4822-0847-4
  3. ^ a b Crother, B.I. (chair) (2012). Scientific and Standard English Names of Amphibians and Reptiles of North America North of Mexico. 7th ed. SSAR Herpetological Circular 39. 84 pp. PDF at SSAR. Accessed 4 July 2011.
  4. ^ a b Powell, R.; Collins, J.T.; Hooper, Jr., E.D. (2019). Key to the Herpetofauna of the Continental United States and Canada (3rd ed.). Lawrence, Kansas: University Press of Kansas. ISBN 978-0700628902.
  5. ^ Utiger, U.; Helfenberger, N.; Schätti, B.; Schmidt, C.; Ruf, M.; Ziswiler, V. (2002). "Molecular Systematics and Phylogeny of Old and New World Ratsnakes ..." Archived 2020-07-22 at the Wayback Machine Russian Journal of Herpetology 9 (2): 105-124.
  6. ^ a b c Pyron, R.A.; Burbrink, F.T. (2009). "Neogene diversification and taxonomic stability in the snake tribe Lampropeltini (Serpentes: Colubridae)". Molecular Phylogenetics and Evolution. 52 (2): 524–529. doi:10.1016/j.ympev.2009.02.008. PMID 19236930.
  7. ^ Figueroa, A.; McKelvy, A.D.; Grismer, L.L.; Bell, C.D.; Lailvaux, S.P. (2016). "A species-level phylogeny of extant snakes with description of a new colubrid subfamily and genus". PLOS ONE. 11 (9): e0161070. Bibcode:2016PLoSO..1161070F. doi:10.1371/journal.pone.0161070. PMC 5014348. PMID 27603205.
  8. ^ a b Pyron, R.A.; Burbrink, F.T.; Wiens, J.J. (2013). "A phylogeny and revised classification of Squamata, including 4161 species of lizards and snakes". BMC Evolutionary Biology. 13: 93. doi:10.1186/1471-2148-13-93. PMC 3682911. PMID 23627680.
  9. ^ a b Dahn, H.A.; Strickland, J.L.; Osorio, A.; Colston, T.J.; Parkinson, C.L. (2018). "Hidden diversity within the depauperate genera of the snake tribe Lampropeltini (Serpentes, Colubridae)". Molecular Phylogenetics and Evolution. 129: 214–225. doi:10.1016/j.ympev.2018.08.018. PMID 30189319. S2CID 52170106.
  10. ^ Pantherophis. The Reptile Database. www.reptile-database.org.
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Pantherophis: Brief Summary ( Inglês )

fornecido por wikipedia EN

Pantherophis is a genus of nonvenomous colubrid snakes endemic to central and eastern regions of North America. It consists of the North American ratsnakes, the foxsnakes, and the cornsnakes. The genus, which contains 10 recognized species, first appeared in the fossil record in the Middle Miocene around 16.3 million years ago. They are a large terrestrial snake genus that lack subocular scales. Originally classified in the genus Elaphe, phylogenetic studies have found this taxon to be closely related to Pituophis. As with all snakes Pantherophis is an obligate faunivore with a diet that consists of small mammals, birds, reptiles and amphibians, and even insects. While many species conservation status is categorized as "least concern", many local populations in some species have declined where some places have them listed as federally protected. The corn snake (P. guttatus) is a popular pet reptile, due to the availability of captive-bred animals, their low maintenance and calm disposition, and the variety of color morphs. There are other species of Pantherophis that are in the pet trade, though are not as popular as the corn snake.

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Pantherophis ( Espanhol; Castelhano )

fornecido por wikipedia ES

Pantherophis es un género de serpientes de la familia Colubridae. Sus especies se distribuyen por Norteamérica, excepto su extremo norte.

Especies

Se reconocen las siguientes:[1]

Referencias

  1. Uetz, P. & Jirí Hošek (ed.). «Pantherophis». Reptile Database. Reptarium. Consultado el 24 de mayo de 2014.
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Pantherophis: Brief Summary ( Espanhol; Castelhano )

fornecido por wikipedia ES

Pantherophis es un género de serpientes de la familia Colubridae. Sus especies se distribuyen por Norteamérica, excepto su extremo norte.

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Pantherophis ( Estônio )

fornecido por wikipedia ET

Pantherophis on maoperekond.

Klassifikatsioon

Perekonda Pantherophis klassifitseeritakse roomajate andmebaasis järgmised maoliigid[1]:

Levila

Neid madusid võib kohata Mehhikos, Ameerika Ühendriikides ja Kanada lõunaosas.

Viited

  1. Peter Uetz & Jakob Hallermann, Pantherophis Roomajate andmebaasi veebiversioon (vaadatud 07.09.2014) (inglise keeles)

Välislingid

Selles artiklis on kasutatud prantsuskeelset artiklit fr:Pantherophis seisuga 07.09.2014.

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Pantherophis: Brief Summary ( Estônio )

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Pantherophis on maoperekond.

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Pantherophis ( Basco )

fornecido por wikipedia EU

Pantherophis Colubridae familiako narrasti genero bat da. Mexikotik Kanadaraino hedatzen dira eta Pantherophis guttatus espeziea Antilletan bizi da.

Espezieak

(RLQ=window.RLQ||[]).push(function(){mw.log.warn("Gadget "ErrefAurrebista" was not loaded. Please migrate it to use ResourceLoader. See u003Chttps://eu.wikipedia.org/wiki/Berezi:Gadgetaku003E.");});
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Pantherophis: Brief Summary ( Basco )

fornecido por wikipedia EU

Pantherophis Colubridae familiako narrasti genero bat da. Mexikotik Kanadaraino hedatzen dira eta Pantherophis guttatus espeziea Antilletan bizi da.

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Pantherophis ( Francês )

fornecido por wikipedia FR

Pantherophis est un genre de serpents de la famille des Colubridae[1].

Répartition

Les espèces de ce genre se rencontrent au Mexique, aux États-Unis et dans le sud du Canada[1]. Pantherophis guttatus a été introduit aux Antilles.

Liste des espèces

Selon Reptarium Reptile Database (13 mars 2012)[2] :

Taxinomie

Les espèces de ce genre étaient pour la plupart classées auparavant dans le genre Elaphe, Pantherophis a été relevé de sa synonymie à la suite des analyses génétiques de Utiger & al., 2002[3].

Publication originale

  • Fitzinger, 1843 : Systema Reptilium, fasciculus primus, Amblyglossae. Braumüller et Seidel, Wien, p. 1-106 (texte intégral).

Notes et références

  1. a et b (en) Référence Reptarium Reptile Database : Pantherophis
  2. Reptarium Reptile Database, consulté le 13 mars 2012
  3. Utiger, Helfenberger, Schätti, Schmidt, Ruf & Ziswiler, 2002 : Molecular systematics and phylogeny of Old and New World ratsnakes, Elaphe auct., and related genera (Reptilia, Squamata, Colubridae). Russian Journal of Herpetology, vol. 9, n. 2, p. 105–124 (texte intégral).
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Pantherophis: Brief Summary ( Francês )

fornecido por wikipedia FR

Pantherophis est un genre de serpents de la famille des Colubridae.

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Pantherophis ( Italiano )

fornecido por wikipedia IT

Pantherophis Fitzinger, 1843 è un genere di serpenti della famiglia dei Colubridi.[1]

Tassonomia

Il genere comprende le seguenti specie:[1]

Note

  1. ^ a b Genus: Pantherophis, in The Reptile Database. URL consultato il 22 luglio 2014.

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Pantherophis: Brief Summary ( Italiano )

fornecido por wikipedia IT

Pantherophis Fitzinger, 1843 è un genere di serpenti della famiglia dei Colubridi.

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Pantherophis ( Polonês )

fornecido por wikipedia POL
Commons Multimedia w Wikimedia Commons

Pantherophisrodzaj węża z podrodziny Colubrinae w rodzinie z rodziny połozowatych (Colubridae).

Zasięg występowania

Rodzaj obejmuje gatunki występujące w Ameryce Północnej[5].

Systematyka

Etymologia

  • Pantherophis: gr. πανθηρ panthēr „lampart, pantera”[6]; οφις ophis, οφεως opheōs „wąż”[7].
  • Scotophis: gr. σκοτος skotos „ciemny”[8]; οφις ophis, οφεως opheōs „wąż”[7]. Gatunek typowy: Coluber alleghaniensis Holbrook, 1836.
  • Coryphodon: gr. κορυφη koruphē „czubek głowy”, od κορυς korus, κορυθος koruthos „chełm”; οδους odous, οδοντος odontos „ząb”[3]. Gatunek typowy: Coluber pantherinus Daudin, 1803.
  • Mintonius: Sherman Anthony Minton Jr. (1919–1999) oraz Madge Rutherford Minton (1920–2004), amerykańscy herpetolodzy[4]. Gatunek typowy: Scotophis vulpinus Baird & Girard, 1853.

Podział systematyczny

Do rodzaju należą następujące gatunki[5]:

Przypisy

  1. L.J.F.J. Fitzinger: Systema reptilium. Fasciculus primus, Amblyglossae. Vindobonae: Braumüller et Seidel, 1843, s. 25. (łac.)
  2. S.F. Baird & Ch.F. Girard: Catalogue of North American reptiles in the Museum of the Smithsonian institution. Cz. 1: Serpents. Washington: Smithsonian Institution, 1853, s. 73. (ang.)
  3. a b A.M.C. Duméril & G. Bibron: Erpétologie générale, ou, Histoire naturelle complète des reptiles. T. 7. Cz. 1. Paris: Roret, 1854, s. 178, 180. (fr.)
  4. a b J.T. Collins & T.W. Taggart. An alternative classification of the New World Rat Snakes (genus Pantherophis [Reptilia: Squamata: Colubridae]). „Journal of Kansas Herpetology”. 26, s. 16–17, 2008 (ang.).
  5. a b P. Uetz & J. Hallermann: Genus: Pantherophis (ang.). The Reptile Database. [dostęp 2019-02-04].
  6. Jaeger 1944 ↓, s. 161.
  7. a b Jaeger 1944 ↓, s. 154.
  8. Jaeger 1944 ↓, s. 207.
  9. Praca zbiorowa: Zwierzęta: encyklopedia ilustrowana. Warszawa: Wydawnictwo Naukowe PWN, 2005, s. 402. ISBN 83-01-14344-4.

Bibliografia

  1. E.C. Jaeger: Source-book of biological names and terms. Springfield: Charles C. Thomas, 1944, s. 1–256. (ang.)
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Pantherophis: Brief Summary ( Polonês )

fornecido por wikipedia POL

Pantherophis – rodzaj węża z podrodziny Colubrinae w rodzinie z rodziny połozowatych (Colubridae).

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Pantherophis ( Português )

fornecido por wikipedia PT

Pantherophis é uma cobra pertencente ao gênero de cobras de origem norte-americana, não venenosas, pertencentes à numerosa família Colubridae. São reproduzidas em cativeiro há vários anos, para fins comerciais, pois são excelentes animais de estimação, estando já bem adaptadas ao cativeiro. São geralmente chamadas de cobras-do-milharal.

Taxonomia

Existem várias espécies dentro do género Pantherophis (anteriormente incluída no género Elaphe), sendo as mais conhecidas a espécies Pantherophis guttatus, Pantherophis obsoletus e a recentemente descrita como sendo uma espécie (e não uma subespécie de P. guttatus) Pantherophis slowinskii, embora esta classificação não seja universalmente aceite.

A espécie Pantherophis guttatus tem várias subespécies, como P. g. guttatus, P.g. emoryi e P.g. rosacea.

Descrição

Largamente reproduzida como animal de estimação nos Estados Unidos, Brasil, Europa e outros paises asiaticos como o Japão. Existem hoje dezenas de variações de cores, porém as pigmentações são 5, o que muda dentre as variações de cores são as combinações de genes ativando ou desativando esse ou aquele pigmento ou simplesmente mudando sua intensidade como é o caso do hipomelanismo.

Os albinismo nas especies ocorrem quando uma cor é bloqueada, no caso de um amelanico a melanina esta ausente, para que isso ocorra o animal deve ser homozigoto amel, caso fosse hetero apresentaria a coloração selvagem.

Existem vários tipos de albinismo dentre as variações, não somente um amelanico é albino, existem também os Black albinos em outras palavras os Aneristicos.

Existem também as duplo homozigotos ( duplo albinos) como Snow ( amelanisco + Aneristico )por exemplo e outras.

Há muitas variações e combinações de pigmentos inclusive Co-dominancia O padrão selvagem é normalmente composto por manchas vermelhas e arredondadas na zona dorsal, que contrastam normalmente com um fundo acinzentado ou alaranjado, tendo o ventre um padrão xadrez, preto e branco.

No geral chegam a 70cm podendo ultrapassar essa marca dependendo da genetica do animal

A sua longevidade média varia entre os 12 e os 18 anos, havendo registo de um espécime que atingiu os 21 anos e 9 meses.

Biologia

São, como a maior parte das cobras, animais activos nas horas crepusculares e nocturnas, embora na Primavera possam estar activas durante o dia. As fêmeas produzem cerca de 20 ovos de cada vez, os quais, ao eclodirem, darão origem a cobras com cerca de 20 centímetros. Não são venenosas, embora na natureza, quando ameaçadas, possam morder, apesar da sua natureza pacífica.

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Pantherophis: Brief Summary ( Português )

fornecido por wikipedia PT

Pantherophis é uma cobra pertencente ao gênero de cobras de origem norte-americana, não venenosas, pertencentes à numerosa família Colubridae. São reproduzidas em cativeiro há vários anos, para fins comerciais, pois são excelentes animais de estimação, estando já bem adaptadas ao cativeiro. São geralmente chamadas de cobras-do-milharal.

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Pantherophis ( Eslovaco )

fornecido por wikipedia SK

Pantherophis, po slovensky užovka alebo štíhlovka, je rod z čeľade užovkovité.

Druhy

Rod bol vytvorený pomerne nedávno vyčlenením niektorých druhov z rodu Elaphe (pozri tam).

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Pantherophis: Brief Summary ( Eslovaco )

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Pantherophis, po slovensky užovka alebo štíhlovka, je rod z čeľade užovkovité.

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옥수수뱀속 ( Coreano )

fornecido por wikipedia 한국어 위키백과

옥수수뱀속(Pantherophis 판테로피스[*])은 북미중미에 서식하는 무독성 뱀들의 속이다. 뱀과 뱀아과에 속한다. 독은 없는 반면 모두 크게 자라 설치류를 졸라 죽이기에 사람에게 이롭다. 모식종옥수수뱀이다.

옥수수뱀속은 1843년 모식종 옥수수뱀과 함께 기재된 오래된 속이다. 하지만 옥수수뱀 외에 현재 옥수수뱀속으로 분류되는 대형 뱀들은 원래 구대륙구렁이들(한국구렁이, 일본구렁이 등)과 함께 뱀속에 속해 있었다. 2002년에야 이 신대륙구렁이들이 구대륙구렁이들과 분류학적으로 상이하다는 학설이 제기되었고,[1] 2009년 계통학 연구로 확증되면서 현재와 같이 재분류되었다.[2]

각주

  1. Utiger, Urs; Notker Helfenberger; Beat Schätti; Catherine Schmidt; Markus Ruf; and Vincent Ziswile (2002). "Molecular Systematics and Phylogeny of Old and New World Ratsnakes ..." Archived 2020년 7월 22일 - 웨이백 머신 Russian Journal of Herpetology 9 (2): 105-124.
  2. Pyron RA, Burbrink FT (2009). "Neogene diversification and taxonomic stability in the snake tribe Lampropeltini (Serpentes: Colubridae)". Mol. Phylogenet. Evol. 52: 524-529.
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