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Cheilanthoideae

Cheilanthoideae ( Inglês )

fornecido por wikipedia EN

Cheilanthoideae is one of the five subfamilies of the fern family Pteridaceae.[1][2] The subfamily is thought to be monophyletic, but some of the genera into which it has been divided are not, and the taxonomic status of many of its genera and species remains uncertain, with radically different approaches in use as of December 2019.

Phylogenic relationships

The following phylogram shows a likely relationship between Cheilanthoideae and the other Pteridaceae subfamilies.[3]

Pteridaceae

Cryptogrammoideae

Parkerioideae (syn. Ceratopteridoideae)

Pteridoideae

Vittarioideae

Cheilanthoideae

Although subfamily Cheilanthoideae itself is thought to be monophyletic, many of the genera into which it has been divided (including Cheilanthes, Doryopteris, Notholaena, and Pellaea) have been shown to be polyphyletic.[4]

Genera

The division of the subfamily Cheilanthoideae into genera and species remains uncertain as of December 2019. Christenhusz et al. (2011), the Pteridophyte Phylogeny Group classification of 2016 (PPG I), and the November 2019 version of the Checklist of Ferns and Lycophytes of the World (World Ferns 8.11) agree on the following genera:[1][2][5]

Some other genera that have been included in the subfamily (or split off from genera included in the subfamily) are:[1][2][5]

  • Allosorus Bernh. – not included by Christenhusz et al. (2011); accepted in PPG I; World Ferns 8.11 includes it in Aleuritopteris, saying that it was wrongly typified
  • Baja Windham & L.O.George – accepted by World Ferns 8.11
  • Mickelopteris Fraser-Jenk. – accepted by World Ferns 8.11
  • Oeosporangium Vis. – included in Allosorus in PPG I; re-accepted by World Ferns 8.11
  • Parahemionitis Panigrahi – accepted by Christenhusz et al. (2011) and PPG I; considered a synonym of Paragymnopteris by World Ferns 8.11, which places the species Parahemionitis cordata in the monotypic genus Mickelopteris
  • Pteridella Mett. ex Kuhn – accepted by Christenhusz & Chase (2014);[6] considered a synonym of Pellaea by World Ferns 8.11

Other sources take a radically different approach. As of December 2019, Plants of the World Online places all of the possible genera of the Cheilanthoideae in the single genus Hemionitis.[7] (Parahemionitis is treated as a synonym of Acrostichum,[8] a member of a different subfamily.)

While much work remains to be done in delineating monophyletic genera in the cheilanthoids (Cheilanthes, Doryopteris and Pellaea remaining notably polyphyletic), several major clades have been consistently recovered in phylogenetic analyses and given informal names, as shown here:

cheilanthoids "ludens clade"

Calciphilopteris

bommeriids

Baja, Bommeria

"skinneri clade"

Cheilanthes skinneri and relatives

myriopterids

Mildella, Myriopteris

pellaeids

Argyrochosma, Astrolepis, Paragymnopteris, and Pellaea

notholaenids

Cheilanthes leucopoda, Cheiloplecton, Notholaena

hemionitids

Adiantopsis, Aleuritopteris, Aspidotis, Cheilanthes, Doryopteris, Gaga, Hemionitis, Lytoneuron, Mickelopteris, Oeosporangium, Ormopteris, Pentagramma, Trachypteris

References

  1. ^ a b c Christenhusz, Maarten; Zhang, Xian-Chun & Schneider, Harald (2011). "A linear sequence of extant families and genera of lycophytes and ferns". Phytotaxa. 19: 7–54. doi:10.11646/phytotaxa.19.1.2. Retrieved 2013-08-11.
  2. ^ a b c PPG I (2016). "A community-derived classification for extant lycophytes and ferns". Journal of Systematics and Evolution. 54 (6): 563–603. doi:10.1111/jse.12229. S2CID 39980610.
  3. ^ Schuettpelz, Eric; Pryer, Kathleen M.; Ranker, Tom A. & Haufler, Christopher H. (2008). "Chapter 15 Fern phylogeny" (PDF). In Ranker, Tom A. & Haufler, Christopher H. (eds.). Biology and Evolution of Ferns and Lycophytes. Cambridge University Press. pp. 417–467. ISBN 978-0-521-87411-3. Retrieved 2019-12-31.
  4. ^ Schuettpelz, Eric; Schneider, Harald; Huiet, Layne; Windham, Michael & Pryer, Kathleen M. (2007). "A molecular phylogeny of the fern family Pteridaceae: Assessing overall relationships and the affinities of previously unsampled genera" (PDF). Molecular Phylogenetics and Evolution. 44 (3): 1172–1185. doi:10.1016/j.ympev.2007.04.011. PMID 17570688. Archived from the original (PDF) on 2008-02-21.
  5. ^ a b Hassler, Michael & Schmitt, Bernd (November 2019). Checklist of Ferns and Lycophytes of the World. 8.11. Retrieved 2019-12-31.
  6. ^ Christenhusz, Maarten J.M. & Chase, Mark W. (2014). "Trends and concepts in fern classification". Annals of Botany. 113 (9): 571–594. doi:10.1093/aob/mct299. PMC 3936591. PMID 24532607.
  7. ^ "Hemionitis L." Plants of the World Online. Royal Botanic Gardens, Kew. Retrieved 2019-12-31.
  8. ^ "Parahemionitis Panigrahi". Plants of the World Online. Royal Botanic Gardens, Kew. Retrieved 2019-12-31.
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Cheilanthoideae: Brief Summary ( Inglês )

fornecido por wikipedia EN

Cheilanthoideae is one of the five subfamilies of the fern family Pteridaceae. The subfamily is thought to be monophyletic, but some of the genera into which it has been divided are not, and the taxonomic status of many of its genera and species remains uncertain, with radically different approaches in use as of December 2019.

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Wikipedia authors and editors
original
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wikipedia EN

Cheilanthoideae ( Espanhol; Castelhano )

fornecido por wikipedia ES

Cheilanthoideae es una de las cinco subfamilias de la familia de helechos Pteridaceae. Esta subfamilia contiene 20 géneros y 400 especies.[1][2][3]

Taxonomía

El siguiente diograma, muestra la relación entre Cheilanthoideae y las otras subfamilias de Pteridaceae.[2]

Pteridaceae

Cryptogrammoideae

       

Ceratopteridoideae

   

Pteridoideae

       

Vittarioideae

   

Cheilanthoideae

       

Géneros

Referencias

  1. Christenhusz, Maarten J. M.; Zhang, Xian-Chun; Schneider, Harald (18 de febrero de 2011). «A linear sequence of extant families and genera of lycophytes and ferns». Phytotaxa 19: 7-54. ISSN 1179-3163.
  2. a b Schuettpelz & Pryer, 2008 Eric Schuettpelz & Kathleen M. Pryer: "Ch. 15. Fern phylogeny" in Biology and Evolution of Ferns and Lycophytes], ed. Tom A. Ranker and Christopher H. Haufler. Cambridge University Press (2008)
  3. Schuettpelz et al. (2007) Eric Schuettpelz, Harald Schneider, Layne Huiet, Michael D. Windham, Kathleen M. Pryer: "A molecular phylogeny of the fern family Pteridaceae: Assessing overall relationships and the affinities of previously unsampled genera." Molecular Phylogenetics and Evolution 44 (2007) 1172–1185
 title=
licença
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Autores y editores de Wikipedia
original
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wikipedia ES

Cheilanthoideae: Brief Summary ( Espanhol; Castelhano )

fornecido por wikipedia ES

Cheilanthoideae es una de las cinco subfamilias de la familia de helechos Pteridaceae. Esta subfamilia contiene 20 géneros y 400 especies.​​​

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Autores y editores de Wikipedia
original
visite a fonte
site do parceiro
wikipedia ES

Cheilanthoideae ( Neerlandês; Flamengo )

fornecido por wikipedia NL

Cheilanthoideae is een onderfamilie binnen de lintvarenfamilie (Pteridaceae), een familie van varens met twaalf geslachten en ongeveer 300 soorten.

Het is een wereldwijd verspreidde groep van temporeel xerofyte (aan droogte aangepaste) varens, met slechts enkele vertegenwoordigers in het zuiden van Europa

Naamgeving en etymologie

De botanische naam Cheilanthoideae is ontleend aan het geslacht Cheilanthes.

Taxonomie

De onderfamilie zoals beschreven door Smith et al. (2006) omvat 14 geslachten met ongeveer 300 soorten. Ondertussen worden regelmatig nieuwe geslachten beschreven, meestal afgescheiden van Cheilanthes:

Bronnen, noten en/of referenties
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original
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Cheilanthoideae: Brief Summary ( Neerlandês; Flamengo )

fornecido por wikipedia NL

Cheilanthoideae is een onderfamilie binnen de lintvarenfamilie (Pteridaceae), een familie van varens met twaalf geslachten en ongeveer 300 soorten.

Het is een wereldwijd verspreidde groep van temporeel xerofyte (aan droogte aangepaste) varens, met slechts enkele vertegenwoordigers in het zuiden van Europa

licença
cc-by-sa-3.0
direitos autorais
Wikipedia-auteurs en -editors
original
visite a fonte
site do parceiro
wikipedia NL