Archaeidae

Figure 7.Lateral ‘head’ profiles of females of species of Austrarchaea from mid-eastern Australia, showing variation in carapace shape as quantified by the post-ocular ratio (P.O. Ratio) and ratio of highest point of carapace relative to post-ocular length (HPC Ratio): A, allotype A. alani sp. n.; B, allotype A. aleenae sp. n.; C, allotype A. judyae sp. n.; D, allotype A. raveni sp. n.; E, allotype A. harmsi sp. n.; F, allotype A. monteithi sp. n.; G, allotype A. cunninghami sp. n.; H, allotype A. dianneae sp. n.; I, A. nodosa (Forster, 1956); J, allotype A. platnickorum sp. n.; K, allotype A. binfordae sp. n.; L, A. milledgei sp. n. (WAM T112568); M, allotype A. mascordi sp. n.; N, allotype A. smithae sp. n.; O, allotype A. mcguiganae sp. n. Asterisks (*) denote concave depressions.

Figure 9.Lateral ‘head’ profiles of females of species of Zephyrarchaea, showing variation in carapace shape as quantified by the post-ocular ratio (P.O. Ratio) and ratio of highest point of carapace relative to post-ocular length (HPC Ratio) (see Fig. 8): A, allotype Zephyrarchaea vichickmani sp. n.; B, allotype Zephyrarchaea marae sp. n.; C, holotype Zephyrarchaea grayi sp. n.; D, holotype Zephyrarchaea austini sp. n.; E, Zephyrarchaea mainae (Platnick, 1991b); F, allotype Zephyrarchaea janineae sp. n.; G, holotype Zephyrarchaea robinsi (Harvey, 2002a); H, allotype Zephyrarchaea melindae sp. n.; I, allotype Zephyrarchaea barrettae sp. n.

Figure 12.Austrarchaea westi sp. n. A–E, Holotype male (QMB S59537) from Mount Williams, Dinden National Park, NE. Queensland: A, cephalothorax and abdomen, lateral view; B, chelicerae, lateral view, showing lack of defined accessory setae; C–D, pedipalpal bulb, ventral view; E, detail of distal tegular sclerites, retrolateral view. C = conductor; E = embolus; T = tegulum; (TS)2-3 = tegular sclerites 2-3. Scale bars: A = 1.0 mm; D = 0.2 mm.

Figure 8.Lateral ‘head’ profiles of males of species of Austrarchaea from south-eastern Queensland and extreme north-eastern New South Wales (including the Border Ranges), showing variation in carapace shape as quantified by the post-ocular ratio (P.O. Ratio) and ratio of highest point of carapace relative to post-ocular length (HPC Ratio): A, holotype A. alani sp. n.; B, holotype A. aleenae sp. n.; C, holotype A. judyae sp. n.; D, holotype A. raveni sp. n.; E, holotype A. harmsi sp. n.; F, holotype A. clyneae sp. n.; G, holotype A. cunninghami sp. n.; H, holotype A. dianneae sp. n.; I, A. nodosa (Forster, 1956) (QMB S75416). Asterisks (*) denote concave depressions.

Figure 19.Zephyrarchaea grayi sp. n. and Zephyrarchaea austini sp. n. A–B, holotype female Zephyrarchaea grayi sp. n. (AMS KS109448) from Delley’s Dell, Grampians National Park, Victoria: A, cephalothorax and abdomen, lateral view; B, internal genitalia, antero-dorsal view. C–D, holotype female Zephyrarchaea austini (SAM NN28000) from Western River Wilderness Protection Area, Kangaroo Island, South Australia: C, cephalothorax and abdomen, lateral view; D, internal genitalia, antero-dorsal view. GP = genital plate. Scale bars: A, C = 1.0 mm.

Figure 21.Distribution of Austrarchaea westi sp. n.: A, topographic map showing the known distribution of Archaeidae in the north-eastern Queensland Wet Tropics bioregion, with collection localities for Austrarchaea westi highlighted in yellow; B-C, topographic and satellite maps showing detail of inset (A). Labelled boundaries in (B) denote upland subregional zones of faunal endemism identified by Winter et al. (1984), Williams et al. (1996) and other authors for the central Wet Tropics (modified from Edward 2011). Small squares in (B–C) denote unidentified juvenile specimens; small circles denote unidentified female specimens; large circles denote described species of Austrarchaea. AU = Atherton Uplands; BK = Bellenden Ker/Bartle Frere; HI = Hinchinbrook Island; KU = Kirrama Uplands; LE = Lee Uplands; LU = Lamb Uplands; MT = Malbon-Thompson Uplands.

Figure 11.Austrarchaea dianneae sp. n. A–B, Cephalothorax and abdomen, lateral view: A, allotype female (QMB S90186) from Tamborine National Park, Queensland; B, holotype male (QMB S90185) from Tamborine National Park, Queensland. C, Holotype male chelicerae, lateral view, showing accessory setae. D–F, Holotype male pedipalp: D–E, bulb, retrolateral view; F, detail of distal tegular sclerites, prodistal view. G, Allotype female internal genitalia, dorsal view. C = conductor; E = embolus; Es = embolic sclerite; T = tegulum; (TS)1–3 = tegular sclerites 1–3. Scale bars: A–B = 1.0 mm; E = 0.2 mm.

Figure 30.Zephyrarchaea austini sp. n., distribution and habitat: A, topographic map showing the known distribution of Archaeidae in Victoria and South Australia, with collection localities for Zephyrarchaea austini highlighted in yellow; B, satellite image showing detail of inset (A); C, open eucalypt woodland and heathland at the type locality – near Billy Goat Falls, Western River Wilderness Protection Area, Kangaroo Island, South Australia (May 2010). Image (C) by M. Rix.

Figure 25. Summary distribution of the Austrarchaea daviesae species-group in tropical north-eastern Queensland, showing collections records for described species (labelled, with black circles) and unidentified juveniles or females (yellow circles) (see Table 1). Note the high proportion of unidentified specimens, especially within the Wet Tropics bioregion between Cooktown and Ingham.

Figure 29.Austrarchaea dianneae sp. n., distribution and habitat: A, topographic map showing the known distribution of Archaeidae in south-eastern Queensland and eastern New South Wales, with collection localities for A. dianneae highlighted in yellow; B, satellite image showing detail of inset (A); C, subtropical rainforest at the type locality – Joalah Section, Tamborine National Park, Queensland (June 2009). Note the sympatric occurrence of this species with A. nodosa on the ‘scenic rim’ at Binna Burra, Lamington National Park. Image (C) by M. Rix.

Figure 1.Habitus images of live Archaeidae from southern Australia: A paratype female Zephyrarchaea vichickmani sp. n. from Acheron Gap, Central Highlands, Victoria; B holotype male Zephyrarchaea barrettae sp. n. from Talyuberlup Peak, Stirling Range National Park, Western Australia; C paratype female Zephyrarchaea janineae sp. n. from Karri Valley, Western Australia; D male Zephyrarchaea marki sp. n. from Thistle Cove, Cape Le Grand National Park, Western Australia; E female Zephyrarchaea mainae (Platnick) from William Bay National Park, Western Australia; F female Zephyrarchaea mainae from Bremer Bay, Western Australia. 

Figure 13.Austrarchaea woodae sp. n. A–E, Holotype male (QMB S72988) from Boulder Caves, Mount Bartle Frere, Wooroonooran National Park, NE. Queensland: A, cephalothorax and abdomen, lateral view; B, chelicerae, lateral view, showing accessory setae; C–D, pedipalpal bulb, ventral view; E, detail of distal tegular sclerites, retrolateral view. C = conductor; E = embolus; T = tegulum; (TS)2-3 = tegular sclerites 2-3. Scale bars: A = 1.0 mm; D = 0.2 mm.

Figure 9.Lateral ‘head’ profiles of males of species of Austrarchaea from New South Wales (excluding the Border Ranges), showing variation in carapace shape as quantified by the post-ocular ratio (P.O. Ratio) and ratio of highest point of carapace relative to post-ocular length (HPC Ratio): A, holotype A. monteithi sp. n.; B, holotype A. christopheri sp. n.; C, holotype A. platnickorum sp. n.; D, holotype A. binfordae sp. n.; E, holotype A. milledgei sp. n.; F, holotype A. mascordi sp. n.; G, holotype A. smithae sp. n.; H, holotype A. mcguiganae sp. n.; I, holotype A. helenae sp. n. Asterisks (*) denote concave depressions.

Figure 8.Lateral ‘head’ profiles of males of species of Zephyrarchaea, showing variation in carapace shape as quantified by the post-ocular ratio (P.O. Ratio) and ratio of highest point of carapace relative to post-ocular length (HPC Ratio): A, holotype Zephyrarchaea marae sp. n., showing the derivation of morphometric ratios; B, holotype Zephyrarchaea vichickmani sp. n.; C, holotype Zephyrarchaea marae sp. n.; D, holotype Zephyrarchaea porchi sp. n.; E, holotype Zephyrarchaea melindae sp. n.; F, holotype Zephyrarchaea barrettae sp. n.; G, holotype Zephyrarchaea mainae (Platnick, 1991b); H, holotype Zephyrarchaea marki sp. n.; I, holotype Zephyrarchaea janineae sp. n.

Figure 22.Distribution of Austrarchaea woodae sp. n.: A, topographic map showing the known distribution of Archaeidae in the north-eastern Queensland Wet Tropics bioregion, with collection localities for Austrarchaea woodae highlighted in yellow; B–C, topographic and satellite maps showing detail of inset (A). Labelled boundaries in (B) denote upland subregional zones of faunal endemism identified by Winter et al. (1984), Williams et al. (1996) and other authors for the central Wet Tropics (modified from Edward 2011). Small squares in (B–C) denote unidentified juvenile specimens; small circles denote unidentified female specimens; large circles denote described species of Austrarchaea. AU = Atherton Uplands; BK = Bellenden Ker/Bartle Frere; HI = Hinchinbrook Island; KU = Kirrama Uplands; LE = Lee Uplands; LU = Lamb Uplands; MT = Malbon-Thompson Uplands.

Figure 20.Austrarchaea christopheri sp. n. A–E, Holotype male (AMS KS114968) from Dorrigo National Park, New South Wales: A, cephalothorax and abdomen, lateral view; B, chelicerae, lateral view, showing accessory setae; C–D, pedipalpal bulb, retrolateral view; E, detail of distal tegular sclerites, prodistal view. C = conductor; E = embolus; Es = embolic sclerite; T = tegulum; (TS)1–3 = tegular sclerites 1–3. Scale bars: A = 1.0 mm; D = 0.2 mm.

Figure 9.Lateral ‘head’ profiles of females of species of Zephyrarchaea, showing variation in carapace shape as quantified by the post-ocular ratio (P.O. Ratio) and ratio of highest point of carapace relative to post-ocular length (HPC Ratio) (see Fig. 8): A, allotype Zephyrarchaea vichickmani sp. n.; B, allotype Zephyrarchaea marae sp. n.; C, holotype Zephyrarchaea grayi sp. n.; D, holotype Zephyrarchaea austini sp. n.; E, Zephyrarchaea mainae (Platnick, 1991b); F, allotype Zephyrarchaea janineae sp. n.; G, holotype Zephyrarchaea robinsi (Harvey, 2002a); H, allotype Zephyrarchaea melindae sp. n.; I, allotype Zephyrarchaea barrettae sp. n.

Figure 25. Summary distribution of the Austrarchaea daviesae species-group in tropical north-eastern Queensland, showing collections records for described species (labelled, with black circles) and unidentified juveniles or females (yellow circles) (see Table 1). Note the high proportion of unidentified specimens, especially within the Wet Tropics bioregion between Cooktown and Ingham.

Figure 1.Habitus images of live Archaeidae from mid-eastern Australia: A–B, female Austrarchaea nodosa (Forster, 1956) from Binna Burra, Lamington National Park, Queensland; C–D, female A. mascordi sp. n. from Coolah Tops National Park, New South Wales; E–F, juvenile A. raveni sp. n. from Mount Glorious, Queensland. Images A–D by M. Rix; images E–F by Greg Anderson, used with permission.

Figure 11.Zephyrarchaea janineae sp. n. A–B, Cephalothorax and abdomen, lateral view: A, allotype female (WAM T118981) from Karri Valley, Western Australia; B, holotype male (WAM T89559) from Karri Valley, Western Australia. C, Holotype male chelicerae, lateral view, showing accessory setae. D–F, Holotype male pedipalp: D–E, bulb, retrolateral view; F, detail of distal tegular sclerites, prolateral view. G, Allotype female internal genitalia, dorsal view. C1–2 = conductor sclerites 1–2; E = embolus; GP = genital plate; T = tegulum; (TS)1–3 = tegular sclerites 1–3. Scale bars: A–B = 1.0 mm; E = 0.2 mm.

Figure 14.Austrarchaea hoskini sp. n. A–B, Cephalothorax and abdomen, lateral view: A, allotype female (QMB S17937) from Mount Elliot, Bowling Green Bay National Park, NE. Queensland; B, holotype male (QMB S30811) from Mount Elliot, Bowling Green Bay National Park, NE. Queensland. C, Holotype male chelicerae, lateral view, showing accessory setae. D–F, Holotype male pedipalp: D–E, bulb, ventral view; F, detail of distal tegular sclerites, retrolateral view. G, Allotype female internal genitalia, postero-ventral view (as seen through posterior rim of genital plate). C = conductor; E = embolus; ESp = embolic spur; GP = genital plate; T = tegulum; (TS)2-3 = tegular sclerites 2-3. Scale bars: A-B = 1.0 mm; E = 0.2 mm.

Figure 7.Lateral ‘head’ profiles of females of species of Austrarchaea from mid-eastern Australia, showing variation in carapace shape as quantified by the post-ocular ratio (P.O. Ratio) and ratio of highest point of carapace relative to post-ocular length (HPC Ratio): A, allotype A. alani sp. n.; B, allotype A. aleenae sp. n.; C, allotype A. judyae sp. n.; D, allotype A. raveni sp. n.; E, allotype A. harmsi sp. n.; F, allotype A. monteithi sp. n.; G, allotype A. cunninghami sp. n.; H, allotype A. dianneae sp. n.; I, A. nodosa (Forster, 1956); J, allotype A. platnickorum sp. n.; K, allotype A. binfordae sp. n.; L, A. milledgei sp. n. (WAM T112568); M, allotype A. mascordi sp. n.; N, allotype A. smithae sp. n.; O, allotype A. mcguiganae sp. n. Asterisks (*) denote concave depressions.

Figure 21.Zephyrarchaea janineae sp. n., distribution and habitat: A, topographic map showing the known distribution of Archaeidae in south-western Western Australia, with collection localities for Zephyrarchaea janineae highlighted in yellow; B, satellite image showing detail of inset (A); C, wet eucalypt forest at the type locality – Karri Valley, Western Australia (August 2006). Image (C) by M. Rix.

Figure 23.Distribution of Austrarchaea hoskini sp. n.: A, topographic map showing the known distribution of Archaeidae in the north-eastern Queensland Wet Tropics bioregion, with collection localities for Austrarchaea hoskini highlighted in yellow; B–C, topographic and satellite maps showing detail of inset (A). Labelled boundaries in (B) denote upland subregional zones of faunal endemism identified by Winter et al. (1984), Williams et al. (1996) and other authors for the southern Wet Tropics (modified from Edward 2011). EU = Elliot Uplands; HU = Halifax Uplands.