Archaeidae

Figure 9.Lateral ‘head’ profiles of males of species of Austrarchaea from New South Wales (excluding the Border Ranges), showing variation in carapace shape as quantified by the post-ocular ratio (P.O. Ratio) and ratio of highest point of carapace relative to post-ocular length (HPC Ratio): A, holotype A. monteithi sp. n.; B, holotype A. christopheri sp. n.; C, holotype A. platnickorum sp. n.; D, holotype A. binfordae sp. n.; E, holotype A. milledgei sp. n.; F, holotype A. mascordi sp. n.; G, holotype A. smithae sp. n.; H, holotype A. mcguiganae sp. n.; I, holotype A. helenae sp. n. Asterisks (*) denote concave depressions.

Figure 1.Habitus images of live Archaeidae from southern Australia: A paratype female Zephyrarchaea vichickmani sp. n. from Acheron Gap, Central Highlands, Victoria; B holotype male Zephyrarchaea barrettae sp. n. from Talyuberlup Peak, Stirling Range National Park, Western Australia; C paratype female Zephyrarchaea janineae sp. n. from Karri Valley, Western Australia; D male Zephyrarchaea marki sp. n. from Thistle Cove, Cape Le Grand National Park, Western Australia; E female Zephyrarchaea mainae (Platnick) from William Bay National Park, Western Australia; F female Zephyrarchaea mainae from Bremer Bay, Western Australia. 

Figure 25. Summary distribution of the Austrarchaea daviesae species-group in tropical north-eastern Queensland, showing collections records for described species (labelled, with black circles) and unidentified juveniles or females (yellow circles) (see Table 1). Note the high proportion of unidentified specimens, especially within the Wet Tropics bioregion between Cooktown and Ingham.

Figure 21.Austrarchaea platnickorum sp. n. A–B, Cephalothorax and abdomen, lateral view: A, allotype female (AMS KS114970) from New England National Park, New South Wales; B, holotype male (AMS KS114971) from New England National Park, New South Wales. C, Holotype male chelicerae, lateral view, showing accessory setae. D–F, Holotype male pedipalp: D–E, bulb, retrolateral view; F, detail of distal tegular sclerites, prodistal view. G, Allotype female internal genitalia, dorsal view. Note the broken left tegular sclerite 1 (TS 1) in (F), highlighted (*) at the point of breakage, compared to the long, sharply-pointed right TS 1 (see inset). C = conductor; E = embolus; Es = embolic sclerite; T = tegulum; (TS)1–3 = tegular sclerites 1–3. Scale bars: A–B = 1.0 mm; E = 0.2 mm.

Figure 4.Diagnostic characters of Zephyrarchaea gen. n. and Austrarchaea Forster & Platnick. A–B, Cephalothorax, lateral view, showing differences in carapace height and the position of accessory setae on male chelicerae: A, male Austrarchaea harmsi Rix & Harvey; B, male Zephyrarchaea marki sp. n. C–D, Expanded male pedipalps, retro-ventral view, showing differences in the articulation and fusion of the conductor sclerites: C, Austrarchaea helenae Rix & Harvey; D, Zephyrarchaea marae sp. n. bH = basal haematodocha; C = conductor; C1–2 = conductor sclerites 1–2; Cy = cymbium; E = embolus; Es = embolic sclerite; H = distal haematodocha; T = tegulum; (TS)1–3 = tegular sclerites 1–3. Scale bars: C–D = 0.2 mm.

Figure 1.Habitus and habitat images of species Archaeidae from north-eastern Queensland. A–D, Habitus images of live paratype specimens of Austrarchaea griswoldi sp. n. from Eungella National Park: A newly-moulted female with recently cast cuticle; B–C, female, lateral view; D, female carrying egg-sac. E–F, Habitat images: E, tropical rainforest at Broken River, Eungella National Park – type locality of Austrarchaea griswoldi sp. n.; F, dense tropical rainforest at Malaan National Park, Atherton Tableland – locality of Austrarchaea daviesae Forster & Platnick.

Figure 39.Austrarchaea platnickorum sp. n., distribution and habitat: A, topographic map showing the known distribution of Archaeidae in south-eastern Queensland and eastern New South Wales, with collection localities for A. platnickorum highlighted in yellow; B, satellite image showing detail of inset (A); C, snow gum woodland adjacent to cool-temperate Nothofagus moorei rainforest at the type locality – Banksia Point, New England National Park, New South Wales (April 2010). Image (C) by M. Rix.

Figure 8.Lateral ‘head’ profiles of males of species of Zephyrarchaea, showing variation in carapace shape as quantified by the post-ocular ratio (P.O. Ratio) and ratio of highest point of carapace relative to post-ocular length (HPC Ratio): A, holotype Zephyrarchaea marae sp. n., showing the derivation of morphometric ratios; B, holotype Zephyrarchaea vichickmani sp. n.; C, holotype Zephyrarchaea marae sp. n.; D, holotype Zephyrarchaea porchi sp. n.; E, holotype Zephyrarchaea melindae sp. n.; F, holotype Zephyrarchaea barrettae sp. n.; G, holotype Zephyrarchaea mainae (Platnick, 1991b); H, holotype Zephyrarchaea marki sp. n.; I, holotype Zephyrarchaea janineae sp. n.

Figure 15.Austrarchaea griswoldi sp. n. A–B, Cephalothorax and abdomen, lateral view: A, allotype female (QMB S92213) from Broken River, Eungella National Park, NE. Queensland; B, holotype male (QMB S92212) from Broken River, Eungella National Park, NE. Queensland. C, Holotype male chelicerae, lateral view, showing accessory setae. D–F, Holotype male pedipalp: D–E, bulb, ventral view; F, detail of distal tegular sclerites, retrolateral view. G, Allotype female internal genitalia, postero-ventral view (genital plate removed). C = conductor; E = embolus; Es = embolic sclerite; T = tegulum; (TS)1-3 = tegular sclerites 1-3. Scale bars: A-B = 1.0 mm; E = 0.2 mm.

Figure 7.Lateral ‘head’ profiles of females of species of Austrarchaea from mid-eastern Australia, showing variation in carapace shape as quantified by the post-ocular ratio (P.O. Ratio) and ratio of highest point of carapace relative to post-ocular length (HPC Ratio): A, allotype A. alani sp. n.; B, allotype A. aleenae sp. n.; C, allotype A. judyae sp. n.; D, allotype A. raveni sp. n.; E, allotype A. harmsi sp. n.; F, allotype A. monteithi sp. n.; G, allotype A. cunninghami sp. n.; H, allotype A. dianneae sp. n.; I, A. nodosa (Forster, 1956); J, allotype A. platnickorum sp. n.; K, allotype A. binfordae sp. n.; L, A. milledgei sp. n. (WAM T112568); M, allotype A. mascordi sp. n.; N, allotype A. smithae sp. n.; O, allotype A. mcguiganae sp. n. Asterisks (*) denote concave depressions.

Figure 12.Zephyrarchaea marki sp. n. A–E, Holotype male (WAM T118985) from Thistle Cove, Cape Le Grand National Park, Western Australia: A, cephalothorax and abdomen, lateral view; B, chelicerae, lateral view, showing accessory setae; C–D, pedipalpal bulb, retrolateral view; E, detail of distal tegular sclerites, prolateral view. C1–2 = conductor sclerites 1–2; E = embolus; T = tegulum; (TS)1–3 = tegular sclerites 1–3. Scale bars: A = 1.0 mm; D = 0.2 mm.

Figure 24.Distribution of Austrarchaea griswoldi sp. n.: A, topographic map showing the known distribution of Archaeidae in the north-eastern Queensland Mackay and Whitsundays Hinterland, with collection localities for Austrarchaea griswoldi highlighted in yellow; B–C, topographic and satellite maps showing detail of inset (A). Small circles in (B–C) denote unidentified female specimens; large circles denote described species of Austrarchaea.

Figure 9.Lateral ‘head’ profiles of males of species of Austrarchaea from New South Wales (excluding the Border Ranges), showing variation in carapace shape as quantified by the post-ocular ratio (P.O. Ratio) and ratio of highest point of carapace relative to post-ocular length (HPC Ratio): A, holotype A. monteithi sp. n.; B, holotype A. christopheri sp. n.; C, holotype A. platnickorum sp. n.; D, holotype A. binfordae sp. n.; E, holotype A. milledgei sp. n.; F, holotype A. mascordi sp. n.; G, holotype A. smithae sp. n.; H, holotype A. mcguiganae sp. n.; I, holotype A. helenae sp. n. Asterisks (*) denote concave depressions.

Figure 22.Zephyrarchaea marki sp. n., distribution and habitat: A, topographic map showing the known distribution of Archaeidae in south-western Western Australia, with collection localities for Zephyrarchaea marki highlighted in yellow; B, satellite image showing detail of inset (A); C, temperate coastal heathland at the type locality – Thistle Cove, Cape Le Grand National Park, Western Australia (June 2010). Image (C) by M. Rix.

Figure 25. Summary distribution of the Austrarchaea daviesae species-group in tropical north-eastern Queensland, showing collections records for described species (labelled, with black circles) and unidentified juveniles or females (yellow circles) (see Table 1). Note the high proportion of unidentified specimens, especially within the Wet Tropics bioregion between Cooktown and Ingham.

Figure 22.Austrarchaea binfordae sp. n. A–B, Cephalothorax and abdomen, lateral view: A, allotype female (AMS KS13891) from Kerewong State Forest, New South Wales; B, holotype male (AMS KS114969) from Kerewong State Forest, New South Wales. C, Holotype male chelicerae, lateral view, showing accessory setae. D–F, Holotype male pedipalp: D–E, bulb, retrolateral view; F, detail of distal tegular sclerites, prodistal view. G, Allotype female internal genitalia, dorsal view. C = conductor; E = embolus; Es = embolic sclerite; T = tegulum; (TS)1–3 = tegular sclerites 1–3. Scale bars: A–B = 1.0 mm; E = 0.2 mm.

Figure 8.Lateral ‘head’ profiles of males of species of Zephyrarchaea, showing variation in carapace shape as quantified by the post-ocular ratio (P.O. Ratio) and ratio of highest point of carapace relative to post-ocular length (HPC Ratio): A, holotype Zephyrarchaea marae sp. n., showing the derivation of morphometric ratios; B, holotype Zephyrarchaea vichickmani sp. n.; C, holotype Zephyrarchaea marae sp. n.; D, holotype Zephyrarchaea porchi sp. n.; E, holotype Zephyrarchaea melindae sp. n.; F, holotype Zephyrarchaea barrettae sp. n.; G, holotype Zephyrarchaea mainae (Platnick, 1991b); H, holotype Zephyrarchaea marki sp. n.; I, holotype Zephyrarchaea janineae sp. n.

Figure 8.Austrarchaea wallacei sp. n. A–D, Holotype male (QMB S25964) from Mount Misery, Monkhouse Timber Reserve, NE. Queensland: A, cephalothorax and abdomen, lateral view; B, chelicerae, lateral view, showing accessory setae; C–D, right pedipalpal bulb (expanded; flipped horizontal for inter-specific comparison), retrolateral view. bH = basal haematodocha; C = conductor; E = embolus; ESp = embolic spur; H = haematodocha; T = tegulum; (TS)1-3 = tegular sclerites 1-3. Scale bars: A = 1.0 mm; D = 0.2 mm.

Figure 1.Habitus images of live Archaeidae from mid-eastern Australia: A–B, female Austrarchaea nodosa (Forster, 1956) from Binna Burra, Lamington National Park, Queensland; C–D, female A. mascordi sp. n. from Coolah Tops National Park, New South Wales; E–F, juvenile A. raveni sp. n. from Mount Glorious, Queensland. Images A–D by M. Rix; images E–F by Greg Anderson, used with permission.

Figure 9.Lateral ‘head’ profiles of females of species of Zephyrarchaea, showing variation in carapace shape as quantified by the post-ocular ratio (P.O. Ratio) and ratio of highest point of carapace relative to post-ocular length (HPC Ratio) (see Fig. 8): A, allotype Zephyrarchaea vichickmani sp. n.; B, allotype Zephyrarchaea marae sp. n.; C, holotype Zephyrarchaea grayi sp. n.; D, holotype Zephyrarchaea austini sp. n.; E, Zephyrarchaea mainae (Platnick, 1991b); F, allotype Zephyrarchaea janineae sp. n.; G, holotype Zephyrarchaea robinsi (Harvey, 2002a); H, allotype Zephyrarchaea melindae sp. n.; I, allotype Zephyrarchaea barrettae sp. n.

Figure 17.Distribution of Austrarchaea wallacei sp. n.: A, topographic map showing the known distribution of Archaeidae in the north-eastern Queensland Wet Tropics bioregion, with collection localities for Austrarchaea wallacei highlighted in yellow; B–C, topographic and satellite maps showing detail of inset (A). Labelled boundaries in (B) denote upland subregional zones of faunal endemism identified by Winter et al. (1984), Williams et al. (1996) and other authors for the northern Wet Tropics (modified from Edward 2011). Small squares in (B–C) denote unidentified juvenile specimens; small circles denote unidentified female specimens; large circles denote described species of Austrarchaea. BM = Black Mountain Corridor; CU = Carbine Uplands; FU = Mt Finnigan Uplands; TU = Thornton Uplands; WU = Windsor Uplands.

Figure 7.Lateral ‘head’ profiles of females of species of Austrarchaea from mid-eastern Australia, showing variation in carapace shape as quantified by the post-ocular ratio (P.O. Ratio) and ratio of highest point of carapace relative to post-ocular length (HPC Ratio): A, allotype A. alani sp. n.; B, allotype A. aleenae sp. n.; C, allotype A. judyae sp. n.; D, allotype A. raveni sp. n.; E, allotype A. harmsi sp. n.; F, allotype A. monteithi sp. n.; G, allotype A. cunninghami sp. n.; H, allotype A. dianneae sp. n.; I, A. nodosa (Forster, 1956); J, allotype A. platnickorum sp. n.; K, allotype A. binfordae sp. n.; L, A. milledgei sp. n. (WAM T112568); M, allotype A. mascordi sp. n.; N, allotype A. smithae sp. n.; O, allotype A. mcguiganae sp. n. Asterisks (*) denote concave depressions.

Figure 14.Zephyrarchaea melindae sp. n. A–B, Cephalothorax and abdomen, lateral view: A, allotype female (WAM T97468) from Toolbrunup Peak, Stirling Range National Park, Western Australia; B, holotype male (WAM T118986) from Mount Hassell, Stirling Range National Park, Western Australia. C, Holotype male chelicerae, lateral view, showing accessory setae. D–F, Holotype male pedipalp: D–E, bulb, retrolateral view, with inset showing twisted apex of tegular sclerite 1 in retroventral view; F, detail of distal tegular sclerites, prolateral view. G, Allotype female internal genitalia, dorsal view. C1–2 = conductor sclerites 1–2; E = embolus; GP = genital plate; T = tegulum; (TS)1 = tegular sclerite 1. Scale bars: A–B = 1.0 mm; E = 0.2 mm.

Figure 25. Summary distribution of the Austrarchaea daviesae species-group in tropical north-eastern Queensland, showing collections records for described species (labelled, with black circles) and unidentified juveniles or females (yellow circles) (see Table 1). Note the high proportion of unidentified specimens, especially within the Wet Tropics bioregion between Cooktown and Ingham.