Pyrgulopsis micrococcus (Pilsbry 1893)

Pyrgulopsis micrococcus (Pilsbry, 1893)

Amnicola micrococcus Pilsbry in Stearns, 1893:277, fig. 1.—Pilsbry, 1899:121 [in part].—Stearns, 1901:286 [in part; fig. 4].—Hannibal, 1912a:38; 1912b:185.—Walker, 1918:134.—Baker, 1964:174.—Richardson et al., 1991:64.

Fontelicella (Microamnicola) micrococcus.—Gregg and Taylor, 1965:109.—Burch, 1982:26, figs. 231, 244.

Fontelicella micrococcus.—Taylor, 1975:123.—Turgeon et al., 1988:61.

Pyrgulopsis micrococcus.—Hershler and Thompson, 1987:29, figs. 7, 33.—Hershler and Sada, 1987:788, figs. 8a, 9–16).—Hershler, 1989:182, figs. 17c,d, 20–25.—Hershler and Pratt, 1990:285, fig. 5.—USDI, 1991b:58818.

Paludestrina stearnsiana.—Berry, 1909:78.

Amnicola stearnsiana.—Berry, 1948:59.

Paludestrina longinqua.—Hannibal, 1912a:34 [in part].

Hydrobia sp.—Taylor, 1954:69.

DIAGNOSIS.—Shell globose to ovate-conic, small to medium-sized, umbilicate. Penial filament medium length, lobe short. Penial ornament a variably shaped terminal gland.

DESCRIPTION.—Shell (Figure 20a) globose to ovate-conic; height, 1.1–3.1 mm, whorls, 3.25–3.5. Protoconch weakly punctate adapically, becoming smoother toward beginning of teleoconch; later portion with a few weak spiral lines adapically. Teleoconch whorls convex, slightly shouldered; sculpture of moderately strong growth lines. Aperture usually slightly separated from body whorl. Inner lip complete, slightly thickened; columellar lip slightly reflected. Outer lip near orthocline. Umbilicus rimate-perforate. Periostracum light brown.

Operculum (Figure 20b,c) ovate, light amber; nucleus slightly eccentric; dorsal surface weakly frilled. Attachment scar margin slightly thickened between nucleus and mid-point of inner edge; callus small.

Central radular tooth (Figure 37d) with moderately indented dorsal edge; lateral cusps, 4–7; central cusp pointed, slightly broader and longer than laterals; basal cusps, 1, medium-sized, with weak dorsal support. Basal process narrow; basal sockets deep. Lateral margins thickened; neck pronounced.

Cephalic tentacles pale or with small patch of gray-black pigment just distal to eyespots. Snout pale to dark gray-black. Foot pale to black; pigment often especially strong along anterior edge. Opercular lobe pale or black along anterior edge. Neck pale to dark gray-black. Pallial roof, visceral coil moderate to dark gray-black.

Ctenidial filaments, 17, medium height, narrow. Osphradium centered slightly posterior to middle of ctenidial axis. Kidney opening thickened, sometimes white. Stomach caecum small, broad.

Testis, 1.0–1.5 whorls, overlapping anterior stomach chamber almost to posterior edge of style sac. Prostate gland a fat bean-shape, with medium-large (20%–33%) pallial section; pallial vas deferens with proximal kink. Penis (Figure 48d) medium-sized; filament medium length, narrow, tapered; lobe usually short, squat, rounded distally. Terminal gland medium-sized (sometimes reduced-absent), circular-horizontal, borne along ventral surface of distal edge of lobe. Filament dark.

Female genitalia shown in Figure 5e. Ovary, 0.5–0.75 whorl, slightly overlapping posterior stomach chamber. Albumen gland without a pallial section. Capsule gland shorter than albumen gland. Genital aperture a subterminal slit with short vestibule. Coiled oviduct a slight horizontal twist followed by broad horizontal loop (often kinked in middle), positioned well behind pallial wall. Oviduct and bursal duct join just behind pallial wall. Bursa copulatrix ovoid, medium length and width, with up to half of length posterior to gland. Bursal duct moderately broad, medium length. Seminal receptacle pouchlike, short, partly overlapping (or slightly lateral to) anterior bursa copulatrix.

TYPE LOCALITY.—Small spring in Oasis Valley [Nye County], Nevada. Lectotype (Baker, 1964:174), ANSP 67279; paralectotypes, ANSP 368399, USNM 123622.

DISTRIBUTION.—Widespread in southeastern California and southwestern Nevada: Death Valley system, Mohave River drainage, small endorheic basins.

MATERIAL EXAMINED.—USNM 847246, unnamed springs comprising headwaters of Amargosa River, Springdale, Nye County, Nevada (T 10S, R 47E, NE1/4 sec. 31).