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American Cow Parsnip

Heracleum sphondylium subsp. montanum (Schleicher ex Gaudin) Briq.

Comprehensive Description

provided by North American Flora
Heracleum lanatum Michx. Fl. Bor. Am. 1: 166. 1803
Heradeum maximum Bartr. Trav. 344. 1791. (Nomen illegitimum.) Heracleum Douglasii DC. Prodr. 4: 193. 1830. Sphondvlium lanatum Greene, Man. Bay Reg. 157. 1894. Pastinaca lanala K.-Pol. Bull. Soc. Nat. Mosc. II. 29: 113. 1916.
Plants 1-3 m. high, tomentose; leaves orbicular to reniform in general outline, excluding the petioles 20-50 cm. long, 25-50 cm. broad, ternately compound, the leaflets ovate to orbicular, 15-40 cm. long, 10-35 cm. broad, cordate, coarsely serrate and variously lobed; petioles 1-4 dm. long, the upper cauline leaves with conspicuously dilated sheaths; peduncles 5-20 cm. long, tomentose but densely villous below the umbels; involucre of 5-10 deciduous, lanceolateacuminate bracts, 5-20 mm. long; involucel of bractlets similar to the bracts; rays 15-30, unequal, 5-10 cm. long; pedicels 8-20 mm. long; petals white, obovate; fruit obovate to obcordate, 8-12 mm. long, 6-9 mm. broad, somewhat pubescent.
Type locality: "Canada," Michaux.
Distribution: Newfoundland and Labrador to Georgia, west to Alaska, California, and Arizona; Siberia aud the Kurile Islands. {Elmer 5073, Payson & Payson 2299.)
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bibliographic citation
Albert Charles Smith, Mildred Esther Mathias, Lincoln Constance, Harold William Rickett. 1944-1945. UMBELLALES and CORNALES. North American flora. vol 28B. New York Botanical Garden, New York, NY
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North American Flora

Heracleum lanatum

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Heracleum maximum, commonly known as cow parsnip, is the only member of the genus Heracleum native to North America. It is also known as American cow-parsnip,[4] Satan celery, Indian celery, Indian rhubarb[5] or pushki.

Description

The leaves are up to 40 cm (16 in) across and divided into lobes.
The seeds are 8–12 mm (51612 in) long and 5–8 mm (316516 in) wide.

Cow parsnip is a tall herbaceous perennial plant,[6] reaching heights of 3 metres (10 feet).[7] The stems are hollow and densely hairy.[8] The leaves are very large, up to 40 centimetres (16 inches) across and divided into three lobes.[7] Cow parsnip has the characteristic flower umbels of the carrot family (Apiaceae), blooming from February to September.[7] The umbels can reach 30 cm (12 in) across,[7] flat-topped or rounded, and composed of small white flowers. Sometimes the outer flowers of the umbel are much larger than the inner ones. The seeds are 8–12 millimetres (3812 in) long and 5–8 mm (316516 in) wide.[9]

Similar species

It is commonly confused with Heracleum mantegazzianum (giant hogweed),[10] a much larger plant that typically has purplish spots on the stems, as well as more sharply serrated leaves.[11]

Other tall invasive Heracleum species include H. mantegazzianum, H. sosnowskyi, and H. persicum. Other similar species include wild carrot, cow parsley, poison hemlock, wild parsnip, and species of the genus Angelica.[10]

Taxonomy

The Plant List (which was last updated in 2013) classified H. maximum, H. lanatum, and H. sphondylium subsp. montanum as distinct species.[12][13][14] According to both the Integrated Taxonomic Information System (ITIS) or the National Plant Germplasm System (NPGS), H. lanatum and H. maximum are synonyms for H. sphondylium subsp. montanum,[15][16] a name proposed by Brummitt in 1971.[17][18]

According to the Plant List and Plants of the World Online, H. lanatum and H. maximum are both accepted names, the latter with no infraspecific taxa.[12][19][a] On the other hand, neither are recognized as accepted names by either the ITIS or NPGS. Besides H. lanatum and H. maximum, various scientific names have been attributed to this species.[20]

Synonyms of the grouping previously classified H. lanatum include:

  • Heracleum montanum
  • Heracleum sphondylium subsp. lanatum
  • Heracleum sphondylium var. lanatum
  • Heracleum sphondylium subsp. montanum

Synonyms listed at Plants of the World Online include:[19]

  • Heracleum douglasii DC.
  • Heracleum inperpastum Koidz.
  • Heracleum lanatum Michx.
  • Heracleum sphondylium var. lanatum (Michx.) Dorn
  • Heracleum sphondylium subsp. lanatum (Michx.) Á.Löve & D.Löve
  • Heracleum sphondylium var. tsurugisanense (Honda) H.Ohba
  • Heracleum turugisanense Honda
  • Pastinaca lanata Koso-Pol.
  • Sphondylium lanatum (Michx.) Greene

As H. lanatum, it is listed as having the variety asiaticum.[5][20][19]

The genus name Heracleum (from "Heracles") refers to the very large size of all parts of these plants.[21]

Heracleum maximum Bart. Batiscan River banks, Quebec

Distribution and habitat

Specimen in Homer, Alaska
The same spot in late winter, showing the dead white stalks

The species is native to North America, being distributed throughout most of the continental United States (except the Gulf Coast and a few neighboring states), ranging from the Aleutian Islands and Alaska in the far northwest to Newfoundland on the east coast, extending south as far as California, New Mexico, Kansas, Ohio, and Georgia. It occurs from sea level to elevations of about 2,700 m (9,000 ft).[9] It is especially prevalent in Alaska, where it is often found growing amongst plants like devil's club, which is nearly identical in size and somewhat similar in appearance, and monkshood, a very toxic flower. In Canada, it is found in every province and territory except Nunavut. It is listed as "Endangered" in Kentucky and "Special Concern" in Tennessee.[22] The plant is also found in Siberia, East Asia, and the Kuril Islands.[20][5][23][24]

Ecology

The species is widely recognized as a valuable pasture plant for cows, sheep, and goats. It is also known to be important in the diets of numerous wild animals, especially bears, both grizzly bears and black bears.[20] It is a host plant for the black swallowtail butterfly (Papilio polyxenes), short-tailed swallowtail butterfly (Papilio brevicauda), old world swallowtail butterfly (Papilio machaon), anise swallowtail butterfly (Papilio zelicaon), and the moths Eupithecia tripunctaria, Papaipema harrisii, Papaipema impecuniosa, Agonopterix clemensella, Agonopterix flavicomella, and Depressaria pastinacella.[25]

Toxicity

The plant contains furanocoumarins such as xanthotoxin,[26] angelicin, pimpinellin and isopimpinellin, isoimperatorin, bergapten and isobergapten, 6‐isopentenyloxyisobergapten, and sphondin.[27][28][26] In one study, the young leaves did not contain xanthotoxin, but older, senescing leaves contained "substantial amounts".[26] Some of these furanocoumarins found in cow parsnip are known to have antimicrobial[27][26] properties and are responsible for a rash producing erythematous vesicles (burn-like blisters) and hyperpigmentation that occurs after getting the clear sap onto one's skin.[29][26][30] They are photosensitive, with the rash occurring only after exposure to ultraviolet light.[26][30] Because of this, phytophotodermatitis causing skin blistering may occur after coming into contact with the sap on a sunny day.[30] The scars and pigmentation from these blisters caused by some Heracleum species can last for months or years.[27]

Uses

The thick flower stems, coming into season in early summer, can be peeled and eaten cooked when young, as was done by Native Americans.[31][7] Caution should be taken as the flowers resemble those of the extremely poisonous Cicuta maculata.[32]

Indigenous North Americans have had a variety of uses for cow parsnip, often traveling long distances in the spring—80 kilometres (50 miles) or more—to find the succulent plant shoots.[5] The young stems and leafstalks were peeled and usually eaten raw, while early American settlers cooked the plant.[33] In terms of taste, texture, and nutrients, the peeled stalks resembled celery, which gave rise to the common name "Indian celery". The natives were aware of the toxic effects of the plant, knowing that if the outer skin were not removed, one would get an "itchy mouth" or blistering skin.[5][29] Pregnant women were warned away from the flower bud stalks to prevent newborns from asphyxiating when crying.[5]

At least seven native groups in North America used the plant as a dermatological aid.[5] It could be an ingredient in poultices applied to bruises or sores.[5][34] A poultice prepared from the roots of cow parsnip was applied to swellings, especially of the feet.[20] The dried stems were used as drinking straws for the old or infirm, or made into flutes for children. An infusion of the flowers can be rubbed on the body to repel flies and mosquitoes. A yellow dye can be made from the roots.[34]

References

Footnotes

  1. ^ Heracleum maximum is also accepted by the Database of Vascular Plants of Canada (VASCAN), referencing the in prep family treatment in the Flora of North America project.[3]

Citations

  1. ^ Maiz-Tome, L. (2016). "Heracleum maximum". IUCN Red List of Threatened Species. 2016: e.T64314237A67729681. Retrieved April 17, 2022.
  2. ^ "Heracleum maximum". ipni.org. International Plant Names Index. Retrieved 2018-09-20.
  3. ^ a b "Heracleum maximum W. Bartram". data.canadensys.net. Database of Vascular Plants of Canada (VASCAN). Retrieved 16 December 2018.
  4. ^ "Heracleum maximum". Go Botany. New England Wildflower Society. Retrieved 4 January 2019.
  5. ^ a b c d e f g h Kuhnlein, Harriet V.; Turner, Nancy J. (1986). "Cow-Parsnip (Heracleum lanatum Michx.): An Indigenous Vegetable of Native People of Northwestern North America" (PDF). J. Ethnobiol. 6 (2): 309–324.
  6. ^ Umberto Quattrocchi (19 April 2016). CRC World Dictionary of Medicinal and Poisonous Plants: Common Names, Scientific Names, Eponyms, Synonyms, and Etymology (5 Volume Set). CRC Press. pp. 1959–. ISBN 978-1-4822-5064-0.
  7. ^ a b c d e Spellenberg, Richard (2001) [1979]. National Audubon Society Field Guide to North American Wildflowers: Western Region (rev ed.). Knopf. pp. 340–341. ISBN 978-0-375-40233-3.
  8. ^ Steven Foster; Christopher Hobbs (2002). A Field Guide to Western Medicinal Plants and Herbs. Houghton Mifflin Harcourt. pp. 67–. ISBN 0-395-83806-1.
  9. ^ a b Norman F. Weeden (1996), A Sierra Nevada Flora, Wilderness Press, ISBN 0-89997-204-7
  10. ^ a b "Heracleum maximum: Similar Species". iNaturalist.org. iNaturalist. Retrieved 2018-08-26.
  11. ^ "Giant Hogweed, Heracleum mantegazzianum". maine.gov. State of Maine: Department of Agriculture, Conservation, and Forestry. Retrieved 2018-08-26.
  12. ^ a b "Heracleum maximum W. Bartram". The Plant List, Version 1.1. 2013. Retrieved September 21, 2018.
  13. ^ "Heracleum lanatum Michx". The Plant List, Version 1.1. 2013. Retrieved September 21, 2018.
  14. ^ "Heracleum sphondylium subsp. montanum (Schleich. ex Gaudin) Briq". The Plant List, Version 1.1. 2013. Retrieved September 21, 2018.
  15. ^ "Heracleum sphondylium ssp. montanum (Schleich. ex Gaudin) Briq". Integrated Taxonomic Information System. Retrieved September 21, 2018.
  16. ^ "Taxon: Heracleum sphondylium L. subsp. montanum (Schleich. ex Gaudin) Briq". Germplasm Resources Information Network. United States Department of Agriculture, Agricultural Research Service. Retrieved September 21, 2018.
  17. ^ Brummitt, R. K. (1971). "Relationship of Heracleum lanatum Michx. of North America to H. sphondylium of Europe". Rhodora. 73 (796): 578–584. JSTOR 23311734.
  18. ^ Page, N. A.; Wall, R. E.; Darbyshire, S. J.; Mulligan, G. A. (2006). "The Biology of Invasive Alien Plants in Canada. 4. Heracleum mantegazzianum Sommier & Levier". Can. J. Plant Sci. 86 (2): 569–589. doi:10.4141/P05-158.
  19. ^ a b c "Heracleum maximum W.Bartram". Plants of the World Online. Royal Botanical Gardens Kew. Retrieved 4 January 2019.
  20. ^ a b c d e Campbell, Robert B. (1991). Ecology of Heracleum lanatum Michx. (cow parsnip) communities in northwestern Montana (MS thesis). University of Montana. Retrieved September 25, 2018.
  21. ^ Elizabeth L. Horn (1998), Sierra Nevada Wildflowers, Mountain Press, ISBN 0-87842-388-5
  22. ^ USDA, NRCS (n.d.). "Heracleum maximum". The PLANTS Database (plants.usda.gov). Greensboro, North Carolina: National Plant Data Team. Retrieved 2008-03-30.
  23. ^ "Heracleum lanatum Michx". Global Biodiversity Information Facility. Retrieved September 25, 2018.
  24. ^ "Heracleum lanatum: Cow Parsnip". Encyclopedia of Life. Retrieved September 25, 2018.
  25. ^ "HOSTS - The Hostplants and Caterpillars Database at the Natural History Museum". www.nhm.ac.uk.
  26. ^ a b c d e f Camm, Edith L.; Wat, Chi-Kit; Towers, G. H. N. (15 November 1976). "An assessment of the roles of furanocoumarins in Heracleum lanatum". Canadian Journal of Botany. 54 (22): 2562–2566. doi:10.1139/b76-275.
  27. ^ a b c Bahadori, Mir Babak; Dinparast, Leila; Zengin, Gokhan (November 2016). "The Genus Heracleum: A Comprehensive Review on Its Phytochemistry, Pharmacology, and Ethnobotanical Values as a Useful Herb". Comprehensive Reviews in Food Science and Food Safety. 15 (6): 1018–1039. doi:10.1111/1541-4337.12222. PMID 33401836.
  28. ^ O'Neill, Taryn; Johnson, John A.; Webster, Duncan; Gray, Christopher A. (May 2013). "The Canadian medicinal plant Heracleum maximum contains antimycobacterial diynes and furanocoumarins". Journal of Ethnopharmacology. 147 (1): 232–237. doi:10.1016/j.jep.2013.03.009. PMID 23501157.
  29. ^ a b Turner, N. J. 1973. The ethnobotany of the Bella Coola Indians of British Columbia. Syesis, 6: 193-220.
  30. ^ a b c Meades, S.J.; Schnare, D.; Lawrence, K.; Faulkner, C. "Heracleum maximum W.Bartram". Northern Ontario Plant Database. Algoma University College and Great Lakes Forestry Centre, Sault Ste. Marie, Ontario, Canada. Retrieved 8 October 2018.
  31. ^ Lyons, C. P. (1956). Trees, Shrubs and Flowers to Know in Washington (1st ed.). Canada: J. M. Dent & Sons. pp. 125, 196.
  32. ^ Niering, William A.; Olmstead, Nancy C. (1985) [1979]. The Audubon Society Field Guide to North American Wildflowers, Eastern Region. Knopf. p. 331. ISBN 0-394-50432-1.
  33. ^ Reiner, Ralph E. (1969). Introducing the Flowering Beauty of Glacier National Park and the Majestic High Rockies. Glacier Park, Inc. p. 18.
  34. ^ a b "BRIT - Native American Ethnobotany Database". herb.umd.umich.edu.

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wikipedia EN

Heracleum lanatum: Brief Summary

provided by wikipedia EN

Heracleum maximum, commonly known as cow parsnip, is the only member of the genus Heracleum native to North America. It is also known as American cow-parsnip, Satan celery, Indian celery, Indian rhubarb or pushki.

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Heracleum maximum ( Spanish; Castilian )

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Heracleum maximum, comúnmente conocida como la pastinaca de vaca, es el único miembro del género Heracleum originario de América del Norte. H. maximum también se conoce como apio indio, ruibarbo indio o pushki.

Taxonomía

Según The Plant List, Heracleum maximum es un nombre de especie aceptado[2]​ pero la especie no tiene sinónimos ni taxones infraespecíficos. Por otro lado, Heracleum maximum no se reconoce como un nombre aceptado ni por el Sistema Integrado de Información Taxonómica (SIIT) ni por el Sistema Nacional de Germoplasma Vegetal (NPGS). Según ITIS y NPGS, tanto Heracleum maximum como Heracleum lanatum son sinónimos de Heracleum sphondylium subsp. montanum.[3][4]​ Estos tres nombres a menudo se usan indistintamente en la literatura, lo que ha causado cierta dificultad y confusión.

La clasificación que figura a continuación sigue a The Plant List, es decir, cada una de Heracleum maximum, Heracleum lanatum y Heracleum sphondylium subsp. montanum es una especie distinta.[2][5][6]

Distribución

La pastinaca de vaca se distribuye en la mayor parte del territorio continental de los Estados Unidos, excepto en la Costa del Golfo y en algunos estados vecinos. Crece desde el nivel del mar hasta elevaciones de aproximadamente 2,700 metros.[7]​ Es especialmente frecuente en Alaska, donde a menudo se encuentra creciendo entre plantas como Oplopanax horridus, que es casi idéntico en tamaño y muy similar en apariencia, y Aconitum, una flor muy tóxica. En Canadá, se encuentra en todas las provincias y territorios, excepto Nunavut. Está listado como "En peligro" en Kentucky y "Preocupación especial" en Tennessee.

Características

 src=
Las semillas tienen una longitud de 8–12 mm y una anchura de 5–8 mm.

La pastinaca de vaca es una hierba alta, que llega a alturas de más de 2 metros. El nombre del género Heracleum (de "Hércules") se refiere al tamaño muy grande de todas las partes de estas plantas.[8]​ La pastinaca de vaca tiene las características umbelas de flores de la familia de la zanahoria (Apiaceae). Las umbelas miden aproximadamente 20 centímetros (8 pulgadas) de ancho, tienen una superficie plana o redondeada y están compuestas de pequeñas flores blancas. A veces las flores exteriores de la umbela son mucho más grandes que las internas. Las hojas son muy grandes, miden hasta 40 cm de ancho y se dividen en lóbulos. Los tallos son robustos y suculentos. Las semillas tienen una longitud de 8–12 mm y una anchura de 5–8 mm.[7]

Los tallos y las hojas contienen furocumarinas, sustancias químicas responsables de la erupción característica de las vesículas eritematosas (ampollas similares a quemaduras) y la posterior hiperpigmentación que se produce después de que la savia transparente llegue a la piel. El químico es fotosensible,[9]​ y la erupción ocurre solo después de la exposición a la luz ultravioleta. Debido a esto, la fitofotodermatitis puede ocurrir después de cortar las plantas en un día soleado.

Comúnmente se confunde con Heracleum mantegazzianum (hierba de cerdo gigante),[10]​ que es una planta mucho más grande que típicamente tiene manchas purpurinas en los tallos, así como hojas más sagradas.[11]

Usos

Los indígenas norteamericanos han tenido una variedad de usos para la pastinaca de vaca. Podría ser un ingrediente en cataplasmas aplicadas a las contusiones o llagas. Los tallos jóvenes y los tallos de las hojas se usaron como alimento una vez que se despegaba la piel exterior. Los tallos secos se usaban como pajitas para los ancianos o enfermos, o se convertían en flautas para niños.

Una infusión de las flores se puede frotar sobre el cuerpo para repeler moscas y mosquitos. Un tinte amarillo se puede hacer de las raíces.[12]

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Referencias

  1. «Heracleum maximum». ipni.org. International Plant Names Index. Consultado el 20 de septiembre de 2018.
  2. a b «Heracleum maximum W. Bartram». The Plant List, Version 1.1. 2013. Consultado el 21 de septiembre de 2018.
  3. «Heracleum sphondylium ssp. montanum (Schleich. ex Gaudin) Briq.». Integrated Taxonomic Information System. Consultado el 21 de septiembre de 2018.
  4. «Taxon: Heracleum sphondylium L. subsp. montanum (Schleich. ex Gaudin) Briq.». Germplasm Resources Information Network. United States Department of Agriculture, Agricultural Research Service. Consultado el 21 de septiembre de 2018.
  5. «Heracleum lanatum Michx.». The Plant List, Version 1.1. 2013. Consultado el 21 de septiembre de 2018.
  6. «Heracleum sphondylium subsp. montanum (Schleich. ex Gaudin) Briq.». The Plant List, Version 1.1. 2013. Consultado el 21 de septiembre de 2018.
  7. a b Norman F. Weeden (1996), A Sierra Nevada Flora, Wilderness Press, ISBN 0-89997-204-7.
  8. Elizabeth L. Horn (1998), Sierra Nevada Wildflowers, Mountain Press, ISBN 0-87842-388-5.
  9. Meades, S.J.; Schnare, D.; Lawrence, K.; Faulkner, C. «Heracleum maximum W.Bartram». Northern Ontario Plant Database. Algoma University College and Great Lakes Forestry Centre, Sault Ste. Marie, Ontario, Canada. Consultado el 8 de octubre de 2018.
  10. «Heracleum maximum: Similar Species». iNaturalist.org. iNaturalist. Consultado el 26 de agosto de 2018.
  11. «Giant Hogweed, Heracleum mantegazzianum». maine.gov. State of Maine: Department of Agriculture, Conservation, and Forestry. Consultado el 26 de agosto de 2018.
  12. «BRIT - Native American Ethnobotany Database». herb.umd.umich.edu.

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Heracleum maximum: Brief Summary ( Spanish; Castilian )

provided by wikipedia ES

Heracleum maximum, comúnmente conocida como la pastinaca de vaca, es el único miembro del género Heracleum originario de América del Norte. H. maximum también se conoce como apio indio, ruibarbo indio o pushki.

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Berce laineuse ( French )

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Heracleum maximum

La berce laineuse ou berce très grande (Heracleum maximum, tsot-ka-re-nion pour certains amérindiens) est la seule espèce du genre Heracleum native d'Amérique du Nord, poussant du Nouveau-Mexique (32° N) jusqu'à l'ile du Prince-Patrick (76° N)[1].

Synonyme : Heracleum lanatum.

Description

C'est une plante herbacée vivace à partie aérienne annuelle, qui peut atteindre 2,5 m de hauteur[2].

Tige

Sa tige est dressée, robuste de (1-2 cm) à la base, ramifiée seulement vers le haut; côtelée et creuse.

Fleurs

Ses fleurs sont habituellement blanches, et roses chez de rares spécimens[2].

Habitats

La Berce laineuse croît naturellement dans les zones humides pas trop acides.

Usages

Usages alimentaire

Comme d'autres berces, la plante est comestible (racines y compris), mais la prudence est recommandée lors de sa manipulation ou consommation par des personnes photosensibles.

Usages médicinaux

Elle a été signalée par plusieurs auteurs (par le frère Marie-Victorin le premier) comme probablement utilisée par les Autochtones contre la fièvre, les frissons et le syndrome grippal, en particulier par les Hurons-Wendat (Wendake) lors de la pandémie de grippe espagnole[3].

Précautions

La sève de la berce laineuse, comme pour la plupart des plantes de cette famille (systématiquement pour la berce du Caucase) peut provoquer une dermatite de contact déclenchée par l'exposition de la peau au soleil.

Risques de confusion

La Berce laineuse ressemblen beaucoup à diverses autres espèces de Berce, et ne doit pas être confondue avec la berce du Caucase, invasive et phototoxique, potentiellement source de graves brûlures en cas de contacts avec sa sève.

Galerie

Notes et références

  1. http://plants.usda.gov/java/profile?symbol=HEMA80
  2. a et b « Berce laineuse : Définition simple et facile du dictionnaire », sur www.linternaute.fr (consulté le 17 novembre 2021)
  3. Jacques Rousseau et Marcel Raymond, Études ethnobotaniques québécoises, Montréal, Institut botanique, coll. « Contributions de l'Institut botanique de l'université de Montréal » (no 55), 1945, 154 p. (lire en ligne), « Le folklore botanique de Gaughnawaga », p. 15.

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Berce laineuse: Brief Summary ( French )

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Heracleum maximum

La berce laineuse ou berce très grande (Heracleum maximum, tsot-ka-re-nion pour certains amérindiens) est la seule espèce du genre Heracleum native d'Amérique du Nord, poussant du Nouveau-Mexique (32° N) jusqu'à l'ile du Prince-Patrick (76° N).

Synonyme : Heracleum lanatum.

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Heracleum maximum ( Croatian )

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Heracleum maximum, jedna od vrsti roda Heracleum, (porodica Apiaceae). Rasprostranjena je na području Sjedinjenih država (osim u državama Zaljeva) gdje ju nazivaju cow parsnip, indian celery ili pushki, i u Kanadi.

To je višegodišnja biljka[1] koja cvjeta od svibnja do lipnja bijelim cvijećem. Može narasti preko dva metra. Indijanci su je koristili i kao lijek i za hranu, a njezine osušene stabljike služile su kao slamke za piće ili dječje flaute. Kutenai su je nazivali wumaǂ, a Konkowi chou’-mē-ō.

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Izvori

  1. Wildflower pristupljeno 3. siječnja 2020
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