dcsimg

Distribution

provided by Amphibians and Reptiles of the Philippines

This species is widely distributed, and is recognized to occur in China, New Guinea, the Solomon Islands, Australia, Indonesia, Bangladesh, Cambodia, India, Malaysia, Myanmar, the Philippines, Singapore, Sri Lanka, Thailand, Vietnam, and Papua New Guinea. In the Philippines, this species is widely distributed.

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Faunal Affinity

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This species has been documented throughout much of the Philippines.

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Habitat

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Although this species is classified as a marine snake, it is regularly encountered in streams and river systems in the Philippines, upstream from where the system meets the ocean.

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Conservation Status

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Little file snakes have a broad geographic range and are locally abundant. Populations are considered stable, and there are no known threats to the longterm persistence of this species. Their ability to survive in a variety of aquatic environments allows them to tolerate environmental changes reasonably well.

US Federal List: no special status

CITES: no special status

State of Michigan List: no special status

IUCN Red List of Threatened Species: least concern

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Cooper, L. 2011. "Acrochordus granulatus" (On-line), Animal Diversity Web. Accessed April 27, 2013 at http://animaldiversity.ummz.umich.edu/site/accounts/information/Acrochordus_granulatus.html
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Laken Cooper, Radford University
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Christine Small, Radford University
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John Berini, Special Projects
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Associations

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File snakes have a number of anti-predator adaptations, which are used not only to deter predators, but also serve as adaptations for surviving in an aquatic environment. During the day, they spend much of their time in the shelter of underwater grasses and mud. In addition to olfaction, little file snakes use their forked-tongue as a way to sense potential threats. They also have relatively loose skin, which allows them to flatten their bodies and move more efficiently in the water when escaping potential predators. Major predators of little file snakes include humans, which hut them for their meat, crocodiles, and sea eagles.

Known Predators:

  • humans, (Homo sapiens)
  • crocodiles, (Crocodylus)
  • sea eagles, (Haliaeetus)
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Cooper, L. 2011. "Acrochordus granulatus" (On-line), Animal Diversity Web. Accessed April 27, 2013 at http://animaldiversity.ummz.umich.edu/site/accounts/information/Acrochordus_granulatus.html
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Laken Cooper, Radford University
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Christine Small, Radford University
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John Berini, Special Projects
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Morphology

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Little file snakes are non-venomous. They are considerably smaller than other members of the genus Acrochordus, but have the most striking appearance of any Acrochordus species. Little file snakes are typically brownish-gray, with white or yellow stripes. Average body mass ranges from 69 to 191 g, and body length ranges from 0.6 to 2.43 m. Little file snakes have a number of features that aid in swimming and in deterring predators. All snakes in the genus Acrochordus have flat tails and loose skin that allows them to flatten out, resembling highly venomous sea snakes. Little file snakes have triangular bodies and raised scales, for which the common name "file snake" was given. They have about 100 roughly-textured scales, which allows them to grip and constrict their prey. In addition, these snakes are covered both dorsally and ventrally by tubercles, which allow them to quickly grasp onto plants and rocks, preventing them from being carried away by currents, and helping them to hold onto prey. Little file snakes also have nostrils located on the top of the head, which allows them to breathe without raising their head completely out of the water. Unlike land snakes, the lungs of file snakes extend throughout their body, so that they are able to stay submerged in water for hours without coming up for air. Like other reptiles, file snakes are ectotherms, resulting in a low metabolism. This also allows them to remain submerged for long periods of time. Like other snakes, little file snakes shed their skin during growth. Although terrestrial snakes use rocks and other hard surfaces to help pull off their old skin, little file snakes are highly active in the days prior to shedding. Increased activity rates results in decreased mass and helps loosen their old skin prior to shedding.

File snakes are sexually dimorphic. Females are typically larger than males, with a larger head and longer and heavier body. These larger body proportions help to support the reproductive processes. Average snout to vent length (SVL) in adult females is 686.2 mm, whereas average SVL in males is 648.6 mm. Body weight averages 168.5 g for females and 104 g for males. Females have an average head length of 19.57 mm and an average head width 13.2 mm. Males have an average head length of 17.8 mm and an average head width 11.11 mm.

Range mass: 69 to 191 g.

Range length: 0.6 to 2.43 m.

Average length: 1 m.

Other Physical Features: heterothermic

Sexual Dimorphism: female larger; sexes shaped differently

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Cooper, L. 2011. "Acrochordus granulatus" (On-line), Animal Diversity Web. Accessed April 27, 2013 at http://animaldiversity.ummz.umich.edu/site/accounts/information/Acrochordus_granulatus.html
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Laken Cooper, Radford University
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Christine Small, Radford University
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John Berini, Special Projects
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Life Expectancy

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In captivity, the lifespan of Acrochordus granulatus ranges from 3 to 5 years. This short lifespan is due to their specialized needs for a warm environment and a steady supply of live prey. Other species in the genus Acrochordus live slightly longer in captivity. Acrochordus arafura has been reported to live nearly 9 years, and A. javanicus approximately 6 years. There is no information available regarding the lifespan of this species in the wild.

Typical lifespan
Status: captivity:
3 to 5 years.

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Cooper, L. 2011. "Acrochordus granulatus" (On-line), Animal Diversity Web. Accessed April 27, 2013 at http://animaldiversity.ummz.umich.edu/site/accounts/information/Acrochordus_granulatus.html
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Laken Cooper, Radford University
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Christine Small, Radford University
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John Berini, Special Projects
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Habitat

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Like other species in the genus Acrochordus, little file snakes are fully aquatic. Unlike Javan wart snakes and Arafura snakes, which are strictly freshwater species, little file snakes can tolerate a broad range salinities, from salt to freshwater. Little file snakes have been found at depths ranging from 4 to 20 m, but generally prefer shallow water habitats because they must surface often for air. File snakes can stay completely submerged for hours at a time while hunting for prey and can withstand both slow and fast-moving currents.

Range depth: 4 to 20 m.

Habitat Regions: temperate ; saltwater or marine ; freshwater

Aquatic Biomes: lakes and ponds; rivers and streams; coastal ; brackish water

Wetlands: swamp

Other Habitat Features: estuarine

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Cooper, L. 2011. "Acrochordus granulatus" (On-line), Animal Diversity Web. Accessed April 27, 2013 at http://animaldiversity.ummz.umich.edu/site/accounts/information/Acrochordus_granulatus.html
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Laken Cooper, Radford University
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Christine Small, Radford University
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John Berini, Special Projects
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Distribution

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Little file snakes, also known as wart snakes, occur in aquatic habitats along coastal regions of southeast Asia, Indonesia, northern Australia, and surrounding islands. They are native to the Paleartic, Oriental and Australian regions. There is limited information as to whether little file snakes have been introduced into other regions, but their importance in the pet trade suggest that they are present in captivity in many regions of the world.

Biogeographic Regions: palearctic (Native ); oriental (Native ); australian (Native )

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Cooper, L. 2011. "Acrochordus granulatus" (On-line), Animal Diversity Web. Accessed April 27, 2013 at http://animaldiversity.ummz.umich.edu/site/accounts/information/Acrochordus_granulatus.html
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Laken Cooper, Radford University
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Christine Small, Radford University
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John Berini, Special Projects
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Trophic Strategy

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File snakes are piscivores and their diet primarily consists of small fish, snails, and small crustaceans. More specifically, little file snakes prey heavily on sleeper fish and gobies.

File snakes exhibit sexually dimorphic feeding behavior. Males locate food by sliding along the bottom of the ocean floor and searching in cracks for fish. Females are ambush predators and wait for prey to pass by before attacking. Both sexes have rough skin that helps them keep hold of prey. File snakes do not store food in their body, but digest it as soon as it is caught.

Animal Foods: fish; mollusks; aquatic crustaceans

Primary Diet: carnivore (Piscivore , Eats non-insect arthropods)

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Cooper, L. 2011. "Acrochordus granulatus" (On-line), Animal Diversity Web. Accessed April 27, 2013 at http://animaldiversity.ummz.umich.edu/site/accounts/information/Acrochordus_granulatus.html
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Laken Cooper, Radford University
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Christine Small, Radford University
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John Berini, Special Projects
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Associations

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As a piscivore, Acrochordus granulatus likely has a significant impact on local fish diversity and abundance. They are also prey for a number of different vertebrate predators throughout their geographic range. Parasites of this species have not been documented.

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Cooper, L. 2011. "Acrochordus granulatus" (On-line), Animal Diversity Web. Accessed April 27, 2013 at http://animaldiversity.ummz.umich.edu/site/accounts/information/Acrochordus_granulatus.html
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Laken Cooper, Radford University
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Christine Small, Radford University
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John Berini, Special Projects
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Benefits

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Humans hunt little file snakes for their meat and for trade on the aquaria market. Occasionally, they are also captured for their skin, which can be used as a substitute for leather to make handbags, shoes, wallets and a number of other items.

Positive Impacts: pet trade ; food ; body parts are source of valuable material

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Cooper, L. 2011. "Acrochordus granulatus" (On-line), Animal Diversity Web. Accessed April 27, 2013 at http://animaldiversity.ummz.umich.edu/site/accounts/information/Acrochordus_granulatus.html
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Laken Cooper, Radford University
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Christine Small, Radford University
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John Berini, Special Projects
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Benefits

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There are no known adverse effects of Acrochordus granulatus on humans.

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Cooper, L. 2011. "Acrochordus granulatus" (On-line), Animal Diversity Web. Accessed April 27, 2013 at http://animaldiversity.ummz.umich.edu/site/accounts/information/Acrochordus_granulatus.html
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Laken Cooper, Radford University
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Christine Small, Radford University
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John Berini, Special Projects
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Life Cycle

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Unlike most snakes, little file snakes are ovoviviparous. Embryos do not contain a skull and have very few other developed organs. Little file snakes have indeterminate growth. The metotic fissure, a bony canal connecting the inner ear to the brain case that helps transmit stimuli to the nervous system, is present in all snakes except members of the genus Acrochordus. This simple autapomorphy suggests that acrochords are only distantly related to other groups of snakes.

Development - Life Cycle: indeterminate growth

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Cooper, L. 2011. "Acrochordus granulatus" (On-line), Animal Diversity Web. Accessed April 27, 2013 at http://animaldiversity.ummz.umich.edu/site/accounts/information/Acrochordus_granulatus.html
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Laken Cooper, Radford University
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Christine Small, Radford University
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John Berini, Special Projects
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Behavior

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Outside of breeding season, little file snakes do not communicate with conspecifics. During breeding season females produce pheromones in order to attract potential mates, which detect this pheromone via Jacobson't organ. Like terrestrial snakes, little file snakes use their tongues to detect danger and to locate prey via olfaction. Most snakes have advanced haptic and auditory senses and limited vision, however, these have not been formally tested in little file snakes.

Communication Channels: chemical

Other Communication Modes: pheromones

Perception Channels: visual ; infrared/heat ; tactile ; vibrations ; chemical

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Cooper, L. 2011. "Acrochordus granulatus" (On-line), Animal Diversity Web. Accessed April 27, 2013 at http://animaldiversity.ummz.umich.edu/site/accounts/information/Acrochordus_granulatus.html
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Laken Cooper, Radford University
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Christine Small, Radford University
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John Berini, Special Projects
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Reproduction

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Only limited information exists regarding the mating system of little file snakes. Females release pheromones in order to attract males, and in many cases multiple males arrive. It is not known how females choose a mate, since there is no competition among male file snakes. In sea snakes, males wrap around the larger females to begin the mating process. Mating may last for hours. Some males drown during mating, as females determine when the pair surfaces for air. Male snakes have two penises, but can only mate with one at a time.

Acrochordus granulatus mates once every 2 years, typically during June or July. Gestation ranges from 5 to 7 months. Males produce sperm throughout the year, which moves through the ductus to the cloaca during mating periods. Sperm levels vary depending on time of year and typically increases as mating approaches. Female file snakes accumulate yolk in high amounts during mating season. Females are capable of delaying conception and holding sperm until conditions are suitable for giving birth.

No information is available on sexual maturation in Acrochordus granulatus. In closely related Acrochordus arafura, males become sexually mature as early as 4 years old, with an average age of maturation closer to 5 years. Most A. arafura females reach sexual maturity by 7 years of age. Acrochordus granulatus has clutch sizes ranging from 1 to 12 eggs, with an average of 6 eggs, and birth mass ranges from 6 to 9 g.

Breeding interval: Little file snakes breed once every two years.

Breeding season: Little file snakes breed during June and July and give birth during December.

Range number of offspring: 1 to 12.

Average number of offspring: 6.

Range gestation period: 5 to 7 months.

Key Reproductive Features: iteroparous ; seasonal breeding ; sexual ; ovoviviparous ; sperm-storing

Neither male or female little file snakes provide parental care for young. Offspring are completely independent upon birth.

Parental Investment: no parental involvement

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Cooper, L. 2011. "Acrochordus granulatus" (On-line), Animal Diversity Web. Accessed April 27, 2013 at http://animaldiversity.ummz.umich.edu/site/accounts/information/Acrochordus_granulatus.html
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Laken Cooper, Radford University
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John Berini, Special Projects
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Brief Summary

provided by EOL authors

The three extant Acrochordus species (the only three species included in the family Acrochordidae) exhibit differences in their ecology, scalation, and anatomy. Acrochordus granulatus, which was previously classified in a separate monotypic genus (Chersydrus), are the smallest Acrochordus (~1 m in total length). Acrochordus granulatus is also the most widely distributed species, reported from coastal northwest India to the Solomon Islands in diverse habitats, including freshwater lakes and rivers, mangroves, mudflats, reefs, and open sea (up to 10 km offshore and at depths of 20 m).

(Sanders et al. 2010 and references therein)

The blood oxygen carrying capacity of Acrochordus granulatus exceeds that of other reptiles and approaches levels characteristic of endothermic mammals and birds.Acrochordus, especiallyA. granulatus, maintain an exceptionally large oxygen store (due to both high blood volume and high blood oxygen carrying capacities). The unusually high hematocrit in this species results in increased blood viscosity, which is is at least partially compensated for by unusually low metabolic rates and sluggish habits (Lillywhite et al. 1988). It is not clear whether the evolution of its remarkable blood oxygen carrying capacity is related to the marine habits of A. granulatus or other other ecological factors (e.g., use of calm, anoxic water bodies) (Brischoux et al. 2011)

Lillywhite et al. (2013) studying dehydration and drinking behavior in Acrochordus granulatus. Acrochordus (file snakes) dehydrate in seawater and do not drink seawater when dehydrated in air and offered seawater to drink. However, they will drink freshwater and the threshold of dehydration for first drinking response is a loss of around 7% of original body mass. Detailed studies of A. granulatus support a growing conclusion that diverse taxa of marine snakes require environmental sources of freshwater to maintain water balance, contrary to earlier belief.

References

  • Brischoux, F., G.E.A. Gartner, et al. 2011. Is Aquatic Life Correlated with an Increased Hematocrit in Snakes? PLoS ONE 6(2): e17077. doi:10.1371/journal.pone.0017077
  • Lillywhite H.B., A.W. Smits, M.E. Feder. 1988. Body fluid volumes in the aquatic snake, Acrochordus granulatus. J Herpetol 22: 434-438.
  • Sanders, K.L, A.H. Mumpuni, J.J. Head, and D.J. Gower. 2010. Phylogeny and divergence times of filesnakes (Acrochordus): Inferences from morphology, fossils and three molecular loci. Molecular Phylogenetics and Evolution 56: 857–867.
  • Lillywhite, H.B., H. Heatwole, and C.M. Sheehy III. 2013. Dehydration and Drinking Behavior of the Marine File Snake Acrochordus granulatus. Physiological and Biochemical Zoology 87(1):46–55.

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Distribution

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Continent: Oceania Asia Australia
Distribution: China (Hainan), New Guinea, Solomon Islands, Australia (along the coast of North Territory, Queensland, West Australia) Indonesia (Ambon, Flores, Irian Jaya, Java, Jobi, Schouten, Sulawesi, Sumatra, Ternate, Timor, We, Komodo); Bangladesh; Cambodia; India (including Nicobar Islands); Malaysia (Malaya and East Malaysia); Myanmar (Burma); Philippine Islands (including Palawan: Calamian Islands, Panay); Singapore; Sri Lanka, Thailand; Vietnam. Papua New Guinea (including many satellite islands)
Type locality: not given (fide MANTHEY & GROSSMANN 1997)
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Bradavičník malý ( Czech )

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Bradavičník malý (Acrochordus granulatus) je nejmenší zástupce čeledi Bradavičníkovitých. Je to vodní a nejedovatý had, který jen zřídka leze na souš. Druh se vyznačuje hnědavou či modravou (forma z Filipín, která není běžně dostupná) barvou a příčným černým pruhováním, které napodobuje některé prudce jedovaté mořské hady. Oproti všem jiným rodům hadů chybí bradavičníkům některé kosti v lebce a jsou tedy považováni jen za vzdálené příbuzné běžných hadů.

Popis

Bradavičník malý dorůstá až 100 centimetrů délky. Má drobné šupiny, které se nepřekrývají a jsou poměrně drsné. Jejich kůže je velmi volná, což v kombinaci s drsnými šupinami pomáhá při lovu ryb, kterými se především živí. Jejich zbarvení je různorodé, většinou však různé odstíny modravé (hadi z Filipín), hnědé až béžové s klikatou čárou na hřbetu. Modrá forma z Filipín může na první pohled připomínat jedovatého mořského hada Laticauda laticaudata. Má poměrně krátkou hlavu a nevýrazný krk. Oči i nozdry jsou postavené spíše nahoře, aby mohl být ponořený a vidět nad hladinu. Jejich čelisti nejsou srostlé, mezičelist však není ozubená. Dolní čelist je pevná.

Rozšíření

Sladkovodní had. Tito hadi se vyskytují v jižní Indii, Číně, Indonésii (ostrovy Sumatra, Jáva a Borneo), Vietnamu, Kambodži, Myanmaru, Thajsku, Malajsii, Singapuru až po Filipíny. Obývá rybníky, močály a jezera se stojatou vodou. Z tohoto důvodu

Chov

Chov je doporučován ve stojaté vodě v akváriu s nízkým sloupcem vody ca do 12 cm podle velikosti hada. Nejvhodnější potravou jsou ryby, u malých hadů např paví očka. Filtr by v akváriu neměl vytvářet silný proud vody.

Způsob života

Bradavičník malý je skutečně vodní had, který na břeh vystupuje jen zřídka. Samice rodí živá mláďata.

Reference

  1. Červený seznam IUCN 2018.1. 5. července 2018. Dostupné online. [cit. 2018-08-09]

Externí odkazy

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Bradavičník malý: Brief Summary ( Czech )

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Bradavičník malý (Acrochordus granulatus) je nejmenší zástupce čeledi Bradavičníkovitých. Je to vodní a nejedovatý had, který jen zřídka leze na souš. Druh se vyznačuje hnědavou či modravou (forma z Filipín, která není běžně dostupná) barvou a příčným černým pruhováním, které napodobuje některé prudce jedovaté mořské hady. Oproti všem jiným rodům hadů chybí bradavičníkům některé kosti v lebce a jsou tedy považováni jen za vzdálené příbuzné běžných hadů.

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કાનસીયો (સર્પ) ( Gujarati )

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કાનસીયો (અંગ્રેજી:File Snake, Western Wart Snake, Little File Snake; દ્વિપદ-નામ: Acrochordus granulatus) એ ગુજરાતમાં દેખાતી સર્પોના કુલ બાર(૧૨) કુટુંબોની ત્રેસઠ[૧] (૬૩) જાતિઓમાંની એક બિનઝેરી સર્પની જાતી છે.

ઓળખ

આ સર્પને શરીર પર કાનસ (કાનસ, આરી કે રતેડી તરીકે ઓળખાતું ઓઝારો ઘસવા માટેનું સાધન) ના જેવા ભીંગડા હોવાથી આવું નામકરણ થયું છે. આ જાતીના સર્પમાં માદા નર કરતા લગભગ બમણી લાંબી જોવા મળે છે. વસવાટ પાણીમાં કે પાણીકાંઠે કરે છે. સમગ્ર ગુજરાતના દરીયાકાઠાના વિસ્તારોમાં જોવા મળે છે. સહેલાઈથી હાથમાં આવી જતો હોવાથી અને સ્વભાવે નરમ હોવાથી કેટલાક લોકો આ સાપનો ખાવામાં ઉપયોગ કરે છે.

આહાર

આ સર્પને ભોજન તરીકે માછલી, દેડકા અને અન્ય નાના જળચર ગમે છે[૨].

પ્રજનન

એક પ્રજનન ગાળામાં ૬ થી ૧૦ બચ્ચાને જન્મ આપે છે[૨].

સંદર્ભ

  1. દેસાઈ, અજય મ. (એપ્રિલ ૨૦૧૭). સર્પ સંદર્ભ (ગુજરાતનાં સાપ વિષે માહિતિ). પ્રકૃતિ મિત્ર મંડળ. p. ૧૪.
  2. ૨.૦ ૨.૧ દેસાઈ, અજય મ. (એપ્રિલ ૨૦૧૭). સર્પ સંદર્ભ (ગુજરાતનાં સાપ વિષે માહિતિ). પ્રકૃતિ મિત્ર મંડળ. p. ૧૪૯.
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કાનસીયો (સર્પ): Brief Summary ( Gujarati )

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કાનસીયો (અંગ્રેજી:File Snake, Western Wart Snake, Little File Snake; દ્વિપદ-નામ: Acrochordus granulatus) એ ગુજરાતમાં દેખાતી સર્પોના કુલ બાર(૧૨) કુટુંબોની ત્રેસઠ (૬૩) જાતિઓમાંની એક બિનઝેરી સર્પની જાતી છે.

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Acrochordus granulatus

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Acrochordus granulatus is a snake species found from India through Southeast Asia to the Solomon Islands. It is known as the little file snake, marine file snake,[3] and little wart snake.[4] It is completely aquatic and almost helpless on land. No subspecies are currently recognized.[5]

Description

Acrochordus granulatus is the smallest of the three members of the family Acrochordidae, and is commonly called the "little file snake". Acrochordus granulatus is also the only Acrochord that permanently inhabits estuaries as well as coastal seas, dawning its other common name the "marine file snake".[6] All members of Acrochordus are completely aquatic and nearly helpless on land, with the exception of A. granulatus. The file snake has hygroscopic skin that retards desiccation, allowing it to travel out of water.[7]

As with the other members of the genus, Acrochordus granulatus has uniquely spinose scales with an almost rough texture.[8] Members of Acrochordus also possess specialized tubercles with nerve endings on the skin between their scales which provide an extra sensory organ used to feel water movements of prey.[9] Acrochordus granulatus also have laterally compressed tails, and they can flatten dorsoventrally to assist in swimming.[8] Acrochordus granulatus are the most marine of the Acrochordidae and have specialized sublingual salt glands similar to those found in the true sea snake subfamily Hydrophiinae. Despite this, they are still susceptible to dehydration at sea and rely on freshwater lenses built up on the surface of marine water for freshwater. Members of the true sea snake subfamily Hydrophiinae have been shown to exhibit this same behavior.[10][11] Acrochordus granulatus are sexually divergent, with females being slightly larger than males.[12]

Geographic range

Found from both coasts of peninsular India though Southeast Asia, the Indo-Australian Archipelago and northern Australia to the Solomon Islands. This includes Bangladesh, Myanmar, Sri Lanka, the Andaman and Nicobar Islands, Thailand, Cambodia, Vietnam, China (Hainan), the Philippines (Luzon, Cebu and Batayan), Malaysia, Indonesia (Sumatra, Java, Borneo, Flores, Timor, Sulawesi, Ternate, Ambon, and coastal Irian Jaya), Papua New Guinea, the Solomon Islands the coast along northern Australia (Northern Territory and eastern Queensland). No type locality was given with the original description, although Smith (1943) gives "India" and Saint-Girons (1972) gives "Inde."[2]

Feeding

Harold Voris reports field studies revealing a diet of Gobiodei, Eleotridae, Trypauchenidae and small crustaceans in the straits of Malacca. Due to the species' extensive range, the diet likely varies regionally.[13] In captivity they have been known fairly non-preferential to take a variety of fish.[8]

References

  1. ^ Sanders, K.; Murphy, J.; Lobo, A.; Gatus, J. (2010). "Acrochordus granulatus". IUCN Red List of Threatened Species. 2010: e.T176769A7300762. doi:10.2305/IUCN.UK.2010-4.RLTS.T176769A7300762.en. Retrieved 20 November 2021.
  2. ^ a b McDiarmid RW, Campbell JA, Touré T. 1999. Snake Species of the World: A Taxonomic and Geographic Reference, vol. 1. Herpetologists' League. 511 pp. ISBN 1-893777-00-6 (series). ISBN 1-893777-01-4 (volume).
  3. ^ Acrochordus granulatus at the Reptarium.cz Reptile Database. Accessed 16 August 2007.
  4. ^ Western Australian Reptile Species at Frank O'Connor's Birding Western Australia. Accessed 20 September 2007
  5. ^ "Acrochordus granulatus". Integrated Taxonomic Information System. Retrieved 16 August 2007.
  6. ^ Lillywhite, Harvey B.; Ellis, Tamir M. (1994). "Ecophysiological Aspects of the Coastal-Estuarine Distribution of Acrochordid Snakes". Estuaries. 17 (1): 53–61. doi:10.2307/1352334. JSTOR 1352334. S2CID 83854655.
  7. ^ Comanns, Philipp; Withers, Philip C.; Esser, Falk J.; Baumgartner, Werner (November 2016). "Cutaneous water collection by a moisture-harvesting lizard, the thorny devil (Moloch horridus)". Journal of Experimental Biology. 219 (21): 3473–3479. doi:10.1242/jeb.148791. PMID 27807218. S2CID 22725331.
  8. ^ a b c Lillywhite, Harvey B. (1996). "Husbandry of the little file snake,Acrochordus granulatus". Zoo Biology. 15 (3): 315–327. doi:10.1002/(SICI)1098-2361(1996)15:3<315::AID-ZOO10>3.0.CO;2-9.
  9. ^ Van Der Kooij, Jeroen; Povel, David (1996). "Scale Sensillae of the File Snake (Serpentes: Acrochordidae) and Some Other Aquatic and Burrowing Snakes". Netherlands Journal of Zoology. 47 (4): 443–456. doi:10.1163/156854297X00111.
  10. ^ Lillywhite, Harvey B.; Heatwole, Harold; Sheehy, Coleman M. (2014). "Dehydration and Drinking Behavior of the Marine File Snake Acrochordus granulatus". Physiological and Biochemical Zoology. 87 (1): 46–55. doi:10.1086/673375. PMID 24457920. S2CID 24775631.
  11. ^ Lillywhite, Harvey B.; Sheehy, Coleman M.; Sandfoss, Mark R.; Crowe-Riddell, Jenna; Grech, Alana (2019). "Drinking by sea snakes from oceanic freshwater lenses at first rainfall ending seasonal drought". PLOS ONE. 14 (2): e0212099. Bibcode:2019PLoSO..1412099L. doi:10.1371/journal.pone.0212099. PMC 6366689. PMID 30730972.
  12. ^ Shine, Richard (1991). "Intersexual Dietary Divergence and the Evolution of Sexual Dimorphism in Snakes" (PDF). The American Naturalist. 138: 103–122. doi:10.1086/285207. S2CID 84056277.
  13. ^ Voris, Harold K.; Glodek, Garrett S. (1980). "Habitat, Diet, and Reproduction of the File Snake, Acrochordus granulatus, in the Straits of Malacca". Journal of Herpetology. 14 (1): 108–111. doi:10.2307/1563896. JSTOR 1563896.
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Acrochordus granulatus: Brief Summary

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Acrochordus granulatus is a snake species found from India through Southeast Asia to the Solomon Islands. It is known as the little file snake, marine file snake, and little wart snake. It is completely aquatic and almost helpless on land. No subspecies are currently recognized.

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Acrochordus granulatus ( Basque )

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(RLQ=window.RLQ||[]).push(function(){mw.log.warn("Gadget "ErrefAurrebista" was not loaded. Please migrate it to use ResourceLoader. See u003Chttps://eu.wikipedia.org/wiki/Berezi:Gadgetaku003E.");});
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Acrochordus granulatus: Brief Summary ( Basque )

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Acrochordus granulatus Acrochordus generoko animalia da. Narrastien barruko Acrochordidae familian sailkatuta dago.

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Kölikäärme ( Finnish )

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Kölikäärme (Acrochordus granulatus) on käsnäkäärmeisiin kuuluva käärmelaji, jota tavataan Intian itärannikolta Ceylonille ja aina Filippiineille saakka ulottuvien alueiden jokisuissa ja rannikoilla. Nimensä kölikäärme on saanut vatsapuolella olevasta kölimäisestä suomurivistä, joka ilmeisesti on apuna uimisessa. Käsnäkäärmeitä pyydystetään vuosittain suuria määriä, sillä niiden nahkoja voidaan käyttää kenkien ja laukkujen raaka-aineena.

Lähteet

  1. Sanders, K., Murphy, J., Lobo, A. & Gatus, J.: Acrochordus granulatus IUCN Red List of Threatened Species. Version 2014.1. 2010. International Union for Conservation of Nature, IUCN, Iucnredlist.org. Viitattu 22.6.2014. (englanniksi)
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Kölikäärme: Brief Summary ( Finnish )

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Kölikäärme (Acrochordus granulatus) on käsnäkäärmeisiin kuuluva käärmelaji, jota tavataan Intian itärannikolta Ceylonille ja aina Filippiineille saakka ulottuvien alueiden jokisuissa ja rannikoilla. Nimensä kölikäärme on saanut vatsapuolella olevasta kölimäisestä suomurivistä, joka ilmeisesti on apuna uimisessa. Käsnäkäärmeitä pyydystetään vuosittain suuria määriä, sillä niiden nahkoja voidaan käyttää kenkien ja laukkujen raaka-aineena.

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Acrochordus granulatus ( French )

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Acrochordus granulatus ou serpent lime est une espèce de serpents marins de la famille des Acrochordidae[1].

Il est non venimeux.

Répartition

Cette espèce se rencontre le long des côtes[1] :

Description

 src=
Tête d'Acrochordus granulatus

C'est un serpent marin qui mesure en moyenne 1,20 m. Sa peau est brun gris avec des bandes blanc jaune pâle. Ses écailles sont rugueuses d'où son nom de serpent lime. La femelle est plus grande et plus lourde que le mâle ; elle est ovovivipare et donne naissance tous les 2 ans, après 5 à 7 mois de gestation de 1 à 12 œufs, à des serpenteaux autonomes vers le mois de décembre[2].

 src=
Serpent lime dans la nature

Publication originale

  • Schneider, 1799 : Historiae amphibiorum naturalis et literariae fasciculus primus, vol. 1 (texte intégral).

Notes et références

  1. a et b (en) Référence Reptarium Reptile Database : Acrochordus granulatus
  2. Alain Diringer (préf. Marc Taquet), Mammifères marins et reptiles marins de l'océan Indien et du Pacifique, Éditions Orphie, 2020, 272 p. (ISBN 979-10-298-0254-6), Serpent lime page 237

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Acrochordus granulatus: Brief Summary ( French )

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Acrochordus granulatus ou serpent lime est une espèce de serpents marins de la famille des Acrochordidae.

Il est non venimeux.

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Ular kadut ( Indonesian )

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Ular kadut (Acrochordus granulatus) atau ular karung belang, adalah spesies ular air dari famili Acrochordidae. Ular ini tersebar luas di Asia Tenggara hingga Australia bagian utara.[2]

Pengenalan

 src=
Ilustrasi 1
 src=
Ilustrasi 2

Panjang tubuh ular kadut antara 0.6 sampai 1.2 meter (60-120 cm). Badannya gemuk dan padat, serta dilapisi dengan sisik-sisik kecil yang kasar. Ekornya pendek dan berujung tumpul. Kepala bagian atas berwarna hitam keabu-abuan atau hitam kecokelatan menyerupai lumpur. Tubuh bagian atas berwarna cokelat kelabu atau cokelat lumpur, dengan pola zigzag tebal berwarna hitam keabu-abuan atau hitam kecokelatan. Bagian bawah tubuhnya berwarna lebih muda.[3][4]

Peyebaran dan habitat

Tersebar luas mulai dari India bagian timur laut, Bangladesh, Tiongkok (Hainan) Myanmar, Laos, Vietnam, Kamboja, Thailand, Malaysia, Singapura, kepulauan Nusantara, Filipina, Papua Nugini, hingga pesisir utara Australia.[3][5]

Ular kadut adalah salah satu jenis ular yang toleran terhadap air asin. Habitat utamanya adalah sungai berlumpur, kolam, rawa-rawa, dan muara dekat pantai. Ular ini juga sering dijumpai di daerah tambak ikan atau kolam pancing.[3][4]

Perilaku, makanan, dan reproduksi

Ular kadut aktif pada malam hari dan menghabiskan sebagian besar waktunya di dasar perairan. Akan tetapi, ular ini lebih sering berkelana di tempat yang lebih dangkal karena harus sering menuju permukaan untuk mmenghirup udara. Walaupun begitu, bila ditaruh di darat, ular ini menjadi lemas dan lamban. Makanan utama ular ini adalah ikan. Ular ini juga memangsa belut serta bebrapa jenis kepiting dan udang. Di daerah tambak, ular ini dianggap sebagai hama ikan yang sangat rakus, dan sering merugikan para pemilik dan pengelola tambak ikan karena memakan ikan-ikan mereka.[3][4]

Ular kadut berkembangbiak dengan melahirkan (ovovivipar). Jumlah anak yang dihasilkan sebanyak 5 sampai 10 ekor, masing-masing ular muda berukuran panjang sekitar 22 cm.[3][4]

Daftar pustaka


  • Greer, A.E. 2006. Encyclopedia of Australian Reptiles. Australian Museum Online. Accessed 16 August 2007.
  • Shine, R. 1991. Australian Snakes, a Natural History. Ithaca, New York: Cornell University Press.
  • Smith, M.A. 1943 The Fauna of British India, Ceylon and Burma, Including the Whole of the Indo-Chinese Sub-Region. Reptilia and Amphibia. 3 (Serpentes). Taylor and Francis, London. 583 pp.
  • Wall, Frank 1921 Ophidia Taprobanica or the Snakes of Ceylon. Colombo Mus. (H. R. Cottle, govt. printer), Colombo. xxii, 581 pages

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Ular kadut: Brief Summary ( Indonesian )

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Ular kadut (Acrochordus granulatus) atau ular karung belang, adalah spesies ular air dari famili Acrochordidae. Ular ini tersebar luas di Asia Tenggara hingga Australia bagian utara.

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Acrochordus granulatus ( Italian )

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L'acrocordo granulato (Acrochordus granulatus (Schneider, 1799)), detto anche serpente lima minore, è una specie di serpente che vive nei mari dell'India e dell'Asia meridionale, fino alle Isole Salomone e all'Australia settentrionale.[2]

Descrizione

 src=
Esemplare di acrocordo granulato

È un animale completamente acquatico e quasi inerme sulla terraferma. La sua pelle si ferisce facilmente, per lo meno in cattività, nonostante abbia una superficie decisamente ruvida (da qui il nome comune).

Gli acrocordi granulati sono sessualmente dimorfici: i maschi sono molto più piccoli delle femmine, e possiedono un tipo di colorazione decisamente più vivace. Questi acrocordi sono meno massicci degli altri appartenenti alla famiglia, e sono caratterizzati da una strana piega cutanea, simile a una sutura, posta longitudinalmente in posizione ventrale.

Biologia

L'acrocordo granulato si rinviene principalmente in piccole pozze, estuari e acquitrini, anche se vive abitualmente in mare aperto. Secondo alcuni studi, questi acrocordi si nutrono principalmente di pesci gatto, di pesci arcobaleno e dei cosiddetti "grunters". Occasionalmente si cibano di anguille, ma sembra che non caccino anfibi. Gli acrocordi granulati usano la loro pelle ruvida per stritolare le prede sott'acqua. Le femmine di questa specie tendono agguati, mentre i maschi sono predatori attivi.

Note

  1. ^ (EN) Sanders, K., Murphy, J., Lobo, A. & Gatus, J. 2010, Acrochordus granulatus, su IUCN Red List of Threatened Species, Versione 2020.2, IUCN, 2020.
  2. ^ Acrochordus granulatus, in The Reptile Database. URL consultato il 2 luglio 2014.

Bibliografia

  • Greer, A.E. 2006. Encyclopedia of Australian Reptiles. Australian Museum Online
  • Shine, R. 1991. Australian Snakes, a Natural History. Ithaca, New York: Cornell University Press.
  • Smith, M.A. 1943 The Fauna of British India, Ceylon and Burma, Including the Whole of the Indo-Chinese Sub-Region. Reptilia and Amphibia. 3 (Serpentes). Taylor and Francis, London. 583 pp.
  • Wall, Frank 1921 Ophidia Taprobanica or the Snakes of Ceylon. Colombo Mus. (H. R. Cottle, govt. printer), Colombo. xxii, 581 pages

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Acrochordus granulatus: Brief Summary ( Italian )

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L'acrocordo granulato (Acrochordus granulatus (Schneider, 1799)), detto anche serpente lima minore, è una specie di serpente che vive nei mari dell'India e dell'Asia meridionale, fino alle Isole Salomone e all'Australia settentrionale.

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Acrochordus granulatus ( Vietnamese )

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Rắn rầm ri cá (tên khoa học: Acrochordus granulatus) là một loài rắn trong họ Acrochordidae. Loài này được Schneider mô tả khoa học đầu tiên năm 1799.[3] Nó hoàn toàn sống thủy sinh và hầu như không làm gì được trên cạn. Không có phân loài nào được công nhận.[4]

Phân bố

Loài này phân bố tại Ấn Độ (bao gồm cả quần đảo Nicobar, quần đảo Andaman), Sri Lanka, Bangladesh, Myanmar (Burma), Trung Quốc (Hải Nam), Việt Nam, Campuchia, Thái Lan, Malaysia (bán đảo Mã Lai và Đông Malaysia), Singapore, các đảo thuộc Philippines (bao gồm Palawan: quần đảo Calamian, Panay), Indonesia (Ambon, Flores, Irian Jaya, Java, Jobi, Schouten, Sulawesi, Sumatra, Ternate, Timor, We, Komodo), Papua New Guinea (kể cả các đảo vệ tinh), New Guinea, quần đảo Solomon, Vanuatu, Australia (dọc theo vùng duyên hải Lãnh thổ Bắc Úc, Queensland, Tây Úc).

Hình ảnh

Tập tin:ChersydrusGranulatusRooij.jpg

Tham khảo

  1. ^ Sanders, K., Murphy, J., Lobo, A. & Gatus, J. (2010). Acrochordus granulatus. Sách Đỏ IUCN các loài bị đe dọa. Phiên bản 2013.2. Liên minh Bảo tồn Thiên nhiên Quốc tế. Truy cập ngày 3 tháng 5 năm 2014.
  2. ^ McDiarmid RW, Campbell JA, Touré T. 1999. Snake Species of the World: A Taxonomic and Geographic Reference, vol. 1. Herpetologists' League. 511 pp. ISBN 1-893777-00-6 (series). ISBN 1-893777-01-4 (volume).
  3. ^ Acrochordus granulatus. The Reptile Database. Truy cập ngày 29 tháng 5 năm 2013.
  4. ^ Acrochordus granulatus (TSN 634832) tại Hệ thống Thông tin Phân loại Tích hợp (ITIS).


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Acrochordus granulatus: Brief Summary ( Vietnamese )

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Rắn rầm ri cá (tên khoa học: Acrochordus granulatus) là một loài rắn trong họ Acrochordidae. Loài này được Schneider mô tả khoa học đầu tiên năm 1799. Nó hoàn toàn sống thủy sinh và hầu như không làm gì được trên cạn. Không có phân loài nào được công nhận.

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瘰鱗蛇 ( Chinese )

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學名 Acrochordus granulatus
Schneider, 1799


瘰鱗蛇學名Acrochordus granulatus)又名疣鱗蛇,主要分布於東南亞印度以及所羅門群島一帶。瘰鱗蛇是一個完全生活於中的蛇種,暫時未有任何亞種被確認。

解剖學描述

瘰鱗蛇是水棲的蛇種,在陸地上幾近完全喪失行動能力。牠們的鱗皮表面質地雖然相當粗糙,但其實皮層偏薄,容易撕裂。瘰鱗蛇是同種異形的,雌雄各有不同型態,雄性瘰鱗蛇體型較小,身體顏色的深淺對比亦較為明顯。瘰鱗蛇經常出沒於河塘、沼澤等地,亦分布於海洋。

生態學描述

瘰鱗蛇主要分布於印度半島東南亞地帶,亦出沒於印尼澳洲海域的群島,與及澳洲北部的所羅門群島等。這些地區包括緬甸安達曼群島尼科巴群島泰國柬埔寨越南中國(主要指海南島)、菲律賓呂宋宿霧等)、馬來西亞印尼蘇門答臘爪哇婆羅洲蘇拉威西島弗洛勒斯島帝汶等)、巴布亞新畿內亞、澳洲北部所羅門群島與及昆士蘭東部。

行為學描述

瘰鱗蛇主要進食魚類。根據希恩(Shine, 1991)的瘰鱗蛇研究,指半數的瘰鱗蛇胃部裡曾發現過彩虹魚、鱸魚等魚類,偶爾也會進食鰻魚,而暫時仍未有發現瘰鱗蛇進食兩棲動物的證據。牠們粗糙的皮膚主要用途是為了抵禦水壓,便於在海底獵食。雌性瘰鱗蛇主要以伏擊的形式捕食獵物,相對地雄性瘰鱗蛇則會比較主動地搜掠獵物。

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瘰鱗蛇: Brief Summary ( Chinese )

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瘰鱗蛇(學名:Acrochordus granulatus)又名疣鱗蛇,主要分布於東南亞印度以及所羅門群島一帶。瘰鱗蛇是一個完全生活於中的蛇種,暫時未有任何亞種被確認。

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Distribution

provided by World Register of Marine Species
India to Australia and Solomons

Reference

van der Land, J. (ed). (2008). UNESCO-IOC Register of Marine Organisms (URMO).

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cc-by-4.0
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WoRMS Editorial Board
contributor
Jacob van der Land [email]

Habitat

provided by World Register of Marine Species
estuarine

Reference

van der Land, J. (ed). (2008). UNESCO-IOC Register of Marine Organisms (URMO).

license
cc-by-4.0
copyright
WoRMS Editorial Board
contributor
Jacob van der Land [email]