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Figure 1.Adults of subgenera Antiperyphanes and Chilioperyphus. A Bembidion (Antiperyphanes) rufoplagiatum. Argentina: Neuquén: Arroyo Queñi at Lago Queñi, DRM voucher V100796 B Bembidion (Antiperyphanes) hirtipes, Argentina: Mendoza: Pampa Palauco, DRM voucher V100792 C Bembidion (Antiperyphanes) spinolai, Argentina: Chubut: Rio Azul at Lago Puelo, DRM voucher V100788 D Bembidion (Antiperyphanes) zanettii, Ecuador: Napo: Rio Quijos W of Baeza, DRM voucher V100791 E Bembidion (Antiperyphanes) mandibulare. Chile: Reg. X, Chiloé: Cucao, DRM voucher V100789 F Bembidion (Chilioperyphus) orregoi, Argentina: Chubut: Rio Azul at Lago Puelo, DRM voucher V100674. Scale bar 1 mm.
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Figure 3.Adults of subgenus Ecuadion. A Bembidion chimborazonum, ECUADOR: Pichincha: Paso de la Virgen, DRM voucher V100793 B Bembidion sanctaemarthae, ECUADOR: Napo: Rio Chalpi Grande, DRM voucher V100798 C Bembidion andersoni, ECUADOR: Pichincha: Reserva Yanacocha, start Andean Snipe Trail, DRM voucher V100655 D Bembidion walterrossi, ECUADOR: Napo: Vinillos, 4.1 km S Cosanga, DRM voucher V100794 E Bembidion cotopaxi, ECUADOR: Pichincha: km 17 on route 28 W of Papallacta DRM voucher V100797 F Bembidion ricei, ECUADOR: Napo: Rio Chalpi Grande, DRM voucher V100622. Scale bar 1 mm.
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Figure 5.Adults of subgenus Nothonepha. A Bembidion tetrapholeon (black form), Argentina: Neuquén: Arroyo Queñi at Lago Queñi, DRM voucher V100781 B Bembidion tetrapholeon (orange form), Argentina: Chubut: Rio Azul at Lago Puelo, DRM voucher V100780 C Bembidion germainianum, Argentina: Neuquén: Rio Salado at route 40, DRM voucher V100782 D Bembidion lonae, Argentina: Mendoza: Salinas del Diamante, DRM voucher V100786 E Bembidion eburneonigrum, Argentina: Neuquén: Rio Neuquén at Chos Malal, DRM voucher V100785 F Bembidion tucumanum, Argentina: Mendoza: Salinas del Diamante, DRM voucher V100783 G Bembidion engelhardti engelhardti, Argentina: Neuquén: Rio Salado at route 40, DRM voucher V100784. Scale bar 1 mm.
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Figure 8.Maximum likelihood bootstrap tree showing only those clades appearing in 90% of the bootstrap replicates; taxa outside of the Antiperyphanes Complex not shown. Numbers below branches indicate maximum likelihood bootstrap percentage / parsimony bootstrap percentage. Circled letters on branches correspond to groups documented in Table 3.
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Figure 8.Maximum likelihood bootstrap tree showing only those clades appearing in 90% of the bootstrap replicates; taxa outside of the Antiperyphanes Complex not shown. Numbers below branches indicate maximum likelihood bootstrap percentage / parsimony bootstrap percentage. Circled letters on branches correspond to groups documented in Table 3.
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Figure 1.Adults of subgenera Antiperyphanes and Chilioperyphus. A Bembidion (Antiperyphanes) rufoplagiatum. Argentina: Neuquén: Arroyo Queñi at Lago Queñi, DRM voucher V100796 B Bembidion (Antiperyphanes) hirtipes, Argentina: Mendoza: Pampa Palauco, DRM voucher V100792 C Bembidion (Antiperyphanes) spinolai, Argentina: Chubut: Rio Azul at Lago Puelo, DRM voucher V100788 D Bembidion (Antiperyphanes) zanettii, Ecuador: Napo: Rio Quijos W of Baeza, DRM voucher V100791 E Bembidion (Antiperyphanes) mandibulare. Chile: Reg. X, Chiloé: Cucao, DRM voucher V100789 F Bembidion (Chilioperyphus) orregoi, Argentina: Chubut: Rio Azul at Lago Puelo, DRM voucher V100674. Scale bar 1 mm.
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Figure 1.Adults of subgenera Antiperyphanes and Chilioperyphus. A Bembidion (Antiperyphanes) rufoplagiatum. Argentina: Neuquén: Arroyo Queñi at Lago Queñi, DRM voucher V100796 B Bembidion (Antiperyphanes) hirtipes, Argentina: Mendoza: Pampa Palauco, DRM voucher V100792 C Bembidion (Antiperyphanes) spinolai, Argentina: Chubut: Rio Azul at Lago Puelo, DRM voucher V100788 D Bembidion (Antiperyphanes) zanettii, Ecuador: Napo: Rio Quijos W of Baeza, DRM voucher V100791 E Bembidion (Antiperyphanes) mandibulare. Chile: Reg. X, Chiloé: Cucao, DRM voucher V100789 F Bembidion (Chilioperyphus) orregoi, Argentina: Chubut: Rio Azul at Lago Puelo, DRM voucher V100674. Scale bar 1 mm.
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Figure 8.Maximum likelihood bootstrap tree showing only those clades appearing in 90% of the bootstrap replicates; taxa outside of the Antiperyphanes Complex not shown. Numbers below branches indicate maximum likelihood bootstrap percentage / parsimony bootstrap percentage. Circled letters on branches correspond to groups documented in Table 3.
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Figure 2.Adults of subgenera Antiperyphus and Notholopha. A Bembidion (Antiperyphus) philippii, Argentina: Neuquén: Rio Collón Curá ca 13 km S La Rinconada, DRM voucher V100787 B Bembidion (Notholopha) scitulum, CHILE: Reg. VII: Los Niches E of Curicó, DRM voucher V100790 C Bembidion (Notholopha) sexfoveolatum, CHILE: Reg. IX: 16.3 km E Malalcahuello, Cuesta Las Raices, DRM voucher V100598. Scale bar 1 mm.
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Figure 2.Adults of subgenera Antiperyphus and Notholopha. A Bembidion (Antiperyphus) philippii, Argentina: Neuquén: Rio Collón Curá ca 13 km S La Rinconada, DRM voucher V100787 B Bembidion (Notholopha) scitulum, CHILE: Reg. VII: Los Niches E of Curicó, DRM voucher V100790 C Bembidion (Notholopha) sexfoveolatum, CHILE: Reg. IX: 16.3 km E Malalcahuello, Cuesta Las Raices, DRM voucher V100598. Scale bar 1 mm.
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Figure 7.Tree of highest likelihood found for the combined, seven-gene matrix with partitioning scheme chosen by PartitionFinder. Only members of the Bembidion Series shown; more distant outgroups not depicted. Scale bar: 0.01, branch lengths as reconstructed by RAxML.
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Figure 8.Maximum likelihood bootstrap tree showing only those clades appearing in 90% of the bootstrap replicates; taxa outside of the Antiperyphanes Complex not shown. Numbers below branches indicate maximum likelihood bootstrap percentage / parsimony bootstrap percentage. Circled letters on branches correspond to groups documented in Table 3.
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Figure 3.Adults of subgenus Ecuadion. A Bembidion chimborazonum, ECUADOR: Pichincha: Paso de la Virgen, DRM voucher V100793 B Bembidion sanctaemarthae, ECUADOR: Napo: Rio Chalpi Grande, DRM voucher V100798 C Bembidion andersoni, ECUADOR: Pichincha: Reserva Yanacocha, start Andean Snipe Trail, DRM voucher V100655 D Bembidion walterrossi, ECUADOR: Napo: Vinillos, 4.1 km S Cosanga, DRM voucher V100794 E Bembidion cotopaxi, ECUADOR: Pichincha: km 17 on route 28 W of Papallacta DRM voucher V100797 F Bembidion ricei, ECUADOR: Napo: Rio Chalpi Grande, DRM voucher V100622. Scale bar 1 mm.
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Figure 4.Habitats of the Antiperyphanes Complex. A River shore, Argentina: Neuquén: Rio Collón Curá, about 13 km S La Rinconada, 625m. On the sandy bank in the foreground Bembidion philippii is abundant, as is Bembidion (Nothonepha) sp. nr. lonae. Also occurring on the sand banks are Bembidium orregoi and Bembidion (Nothonepha) eburneonigrum. On the upper sand banks across the river are Bembidium mandibulare, and in the gravel are Bembidium spinolai B Edges of snowfields at Chile: Reg. IX: Volcán Lonquimay, 1910m. Habitat of Bembidion (Notholopha) sexfoveolatum C Open high-elevation grassland at Ecuador: Pichincha: Paso de la Virgen, 4060m, habitat of three species of subgenus Ecuadion: Bembidion chimborazonum, Bembidion guamani, and Bembidion humboldti D Leaf litter in cloud forest, Ecuador: Pichincha: Reserva Yanacocha, 0.1152°S, 78.5837°W, 3540m, habitat of Bembidion andersoni, Bembidion georgeballi, and Bembidion onorei.
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Figure 5.Adults of subgenus Nothonepha. A Bembidion tetrapholeon (black form), Argentina: Neuquén: Arroyo Queñi at Lago Queñi, DRM voucher V100781 B Bembidion tetrapholeon (orange form), Argentina: Chubut: Rio Azul at Lago Puelo, DRM voucher V100780 C Bembidion germainianum, Argentina: Neuquén: Rio Salado at route 40, DRM voucher V100782 D Bembidion lonae, Argentina: Mendoza: Salinas del Diamante, DRM voucher V100786 E Bembidion eburneonigrum, Argentina: Neuquén: Rio Neuquén at Chos Malal, DRM voucher V100785 F Bembidion tucumanum, Argentina: Mendoza: Salinas del Diamante, DRM voucher V100783 G Bembidion engelhardti engelhardti, Argentina: Neuquén: Rio Salado at route 40, DRM voucher V100784. Scale bar 1 mm.
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CBG Photography Group. Centre for Biodiversity Genomics. CBG Photography Group. Year: 2015. Contact: collectionbio@gmail.com.
Barcode of Life Data Systems
Dorsal. Catalog no.: NFRC-2014-02742. Specimen ID: 5437077. Taxon rep.: Bembidion. Image quality: 6. Aspect ratio: 1.333.
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CBG Photography Group. Centre for Biodiversity Genomics. CBG Photography Group. Year: 2014. Contact: ccdbcol@uoguelph.ca.
Barcode of Life Data Systems
Dorsal. Catalog no.: ENT991-146567. Specimen ID: 4116756. Taxon rep.: Tachyta. Image quality: 4. Aspect ratio: 1.342.
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CBG Photography Group. Smithsonian Institution. Smithsonian Institution. Year: 2018. Contact: collectionsBIO@gmail.com.
Barcode of Life Data Systems
Lateral. Catalog no.: USNMENT01452889. Specimen ID: 9046045. Taxon rep.: Lymnastis. Image quality: 1. Aspect ratio: 1.342.