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Anolis proboscis Peters & Orces 1956

Brief Summary ( 英語 )

由EOL authors提供

The Ecuadorian Horned Anole (Anolis proboscis) is one of three species of anole lizards known as "proboscis anoles". This group, which is also known as the A. laevis group (Williams 1979), consists of A. laevis, A. proboscis, and A. phyllorhinus. Members of all three species have a striking scaly anterior (forward) extension of the snout (Poe et al. 2012).

All three species of proboscis anoles were originally described based on extremely few specimens, all of which were males, and for many decades there were no reported observations of living specimens. The first females of any of these three species were reported by Rodrigues et al. (2002, cited in Losos et al. 2012), who described females of A. phyllorhinus from Amazonian Brazil. Prior to 2005, when a group of birders encountered and photographed a male A. proboscis crossing a road in Mindo, Ecuador (Almendáriz and Vogt 2007), A. proboscis had not been reported since the last specimen was collected in 1966 (Losos et al. 2012). Subsequently, there have been a small number of additional records. Yánez-Muñoz et al. (2010, cited in Losos et al. 2012) described variation in five specimens of A. proboscis (including the first females of this species) that were located in 2007 and 2008 in two areas 11 to 13 km north of previous records. In August and December of 2009, Poe et al. (2012) found and collected 11 additional specimens along a paved road within 5 km of Cunuco, the type locality for this species. Losos et al. (2012) captured 20 individuals as part of their field study in and near Mindo.

Poe et al. (2012) note that the scarcity of specimens of A. laevis and A. phyllorhinus is perhaps unsurprising given their ranges: The single specimen of A. laevis (collected in the mid-19th century) was from a mountain trail connecting two lower elevation towns in Peru and A. phyllorhinus is an Amazonian forest form with an apparently narrow distribution. The scarcity of A. proboscis in collections may be a bit more surprising since the type locality is in a populated area within a 3 hour drive of Quito along a well-traveled, paved road and is frequented by both ecotourists and scientists. However, Losos et al. (2012) attribute the fact that these anoles are rarely observed to their morphology and behavior. They are cryptic in pattern and coloration, move very slowly, and appear to spend most of their time in dense vegetation high off the ground where they are almost impossible to observe.

Based on field observations and measurements, Losos et al. (2012) concluded that, except for its striking proboscis, A. proboscis closely matches the "twig ecomorph" that has evolved multiple times on islands in the Greater Antilles: among other similarities, these anoles are cryptically colored, small, and short-limbed with a narrow head and a tail that is at least somewhat prehensile. The proboscis that gives this group of anoles its name is absent in females of both A. phyllorhinus and A. proboscis (A. laevis females are still unknown). Losos et al. note that although the function of the male's proboscis has not been studied, it is extremely flexible and would clearly not be an effective weapon, although it seems likely that it serves to intimidate other males and/or help attract potential mates.

Poe et al. (2012) reported morphological data from both newly collected and previously preserved specimens of the species, including some characters not previously scored for this species; discussed the results of a phylogenetic analysis aimed at determining the placement of this species within Anolis; and presented new information on the color, behavior, and ecology of A. proboscis in life. Losos et al. (2012) described the results of their behavioral and ecological field study of this species near Mindo, Ecuador.

Although much about A. proboscis remains unknown, much has been learned about this species since its rediscovery, making it by far the best studied of the proboscis anoles. Anolis phyllorhinus remains poorly known and A. laevis is still known from just the preserved specimen from Peru described in 1876. The phylogenetic relationships of all proboscis anoles remain poorly understood (Poe et al. 2012; Losos et al. 2012).

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Distribution ( 英語 )

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Continent: South-America
Distribution: Ecuador (Pichincha), Peru ?
Type locality: œNeighbourhood of Cunuco, a small town at 1200 meters elevation, five kilometers northwest of Mindo, on the south bank of the Rio Mindo, a northern tributary of the upper Rio Blanco, in Pichincha Province, Ecuador
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Pinocchioechse ( 德語 )

由wikipedia DE提供

Die Pinocchioechse (Anolis proboscis, Syn.: Dactyloa proboscis), auch als Hornanolis bezeichnet, ist eine kleine Echsenart aus der Gattung Anolis in der Familie Dactyloidae. Ein einzelnes männliches Exemplar wurde 1953 in Ecuador entdeckt und 1956 von James Arthur Peters und Gustavo Orcés formell beschrieben. Anschließend blieb sie bis zu ihrer Wiederentdeckung im Jahr 2004 verschollen. Ihr derzeit bekannter Lebensraum ist ein kleiner Vegetationsabschnitt entlang einer ecuadorianischen Autobahn.

Merkmale

Die Pinocchioechse erreicht eine Kopf-Rumpf-Länge von 5 bis 7,5 cm. Die Männchen besitzen eine auffälligen 1 bis 2 cm langen Nasenfortsatz oder Rüssel, eine längliche Struktur, die von der Mitte der Schnauze ausgeht und offenbar bei der Balz Verwendung findet, ebenso wie die Wamme, die bei beiden Geschlechtern vorhanden ist. Es gibt mehrere Farbmorphe, wobei die Färbung von gelblich-grün bis braun mit orangefarbenen oder schwarzen Markierungen reicht und der Bauch im Allgemeinen weiß ist.[1]

Verbreitung und Lebensraum

Die Pinocchioechse bewohnt montane Waldlebensräume im Chocó, wo sie sich in hohen Bäumen aufhält.[2] Die Terra typica befindet sich in der Nachbarschaft von Cunuco, einer Kleinstadt auf 1200 Metern Höhe, fünf Kilometer nordwestlich von Mindo, am Südufer des Río Mindo, eines nördlichen Nebenflusses des oberen Río Blanco, in der Provinz Pichincha, Ecuador. Exemplare wurden vorwiegend in der Primär- und Sekundärvegetation entlang einer gut befahrenen Schotterstraße gefunden. Derzeit sind insgesamt nur fünf Fundorte bekannt, wobei die beiden am weitesten entfernten maximal 13 km voneinander entfernt liegen. Während das bekannte Verbreitungsgebiet Weideland und Sekundärwald umfasst, ist es wahrscheinlich, dass die Art auch in anderen Gebieten mittlerer Höhe (von 1200 bis 1650 m) in den ecuadorianischen Anden vorkommt.

Wiederentdeckung und Status

Für die Pinocchioechse wurde bereits das Aussterben befürchtet, da sie nach der Sammlung des Holotypus im Jahr 1953 nicht mehr nachgewiesen wurde. Sie wurde 2004 wiederentdeckt, als ein Individuum in einem Nebelwald bei Mindo, Ecuador, von einem Ornithologen, der zu Besuch war, beobachtet und fotografiert wurde.[3] Im August 2009 entdeckte eine herpetologische Expedition der University of New Mexico die Art in einer abgelegenen Region Ecuadors. Insgesamt fanden sie fünf Individuen, darunter drei Männchen und die ersten beiden Weibchen, die jemals gesehen und gesammelt wurden.[1] Seit 2009 konnten mehrere andere Expeditionen die Art in abgelegene Regionen Ecuadors umsiedeln.[4]

Die Art wird von der IUCN aufgrund ihrer möglicherweise eingeschränkten Verbreitung in Kombination mit anhaltenden Bedrohungen in Form von Lebensraumverlust durch Abholzung, menschliche Besiedlung, Landwirtschaft und Beweidung als „stark gefährdet“ (endangered) eingeschätzt.

Einzelnachweise

  1. a b Steven Poe, Fernando Ayala, Ian M. Latella, Thomas L. Kennedy, James A. Christensen, Levi N. Gray, Natalie J. Blea, Beeling M. Armijo, Eric W. Schaad: Morphology, phylogeny, and behavior of Anolis proboscis. In: Breviora. 530, 2012, S. 1–11. doi:10.3099/530.1.
  2. Mario H. Yánez-Muñoz, Miguel A. Urgilés, Marco Altamirano-Benavides, Stalin R. Cáceres: Redescripción de Anolis proboscis Peters & Orcés (Reptilia: Polychrotidae), con el descubrimiento de las hembras de la especie y comentarios sobre su distribución y taxonomía. In: Avances en Ciencias e Ingeniería. 2, Nr. 3, 2010, S. 1–14. doi:10.18272/aci.v2i3.39.
  3. A. C. Almendariz, C. Vogt: Anolis proboscis (Squamata: Polychrotidae), Una lagartija rara pero no extinta. In: Politécnica. 27, Nr. 4, 2007, S. Biología 7 133–135.
  4. D. Main: Once 'Extinct' Pinocchio Lizard Pokes His Nose Out. In: LiveScience.com. 4. Oktober 2013. Abgerufen am 11. September 2020.
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Pinocchioechse: Brief Summary ( 德語 )

由wikipedia DE提供

Die Pinocchioechse (Anolis proboscis, Syn.: Dactyloa proboscis), auch als Hornanolis bezeichnet, ist eine kleine Echsenart aus der Gattung Anolis in der Familie Dactyloidae. Ein einzelnes männliches Exemplar wurde 1953 in Ecuador entdeckt und 1956 von James Arthur Peters und Gustavo Orcés formell beschrieben. Anschließend blieb sie bis zu ihrer Wiederentdeckung im Jahr 2004 verschollen. Ihr derzeit bekannter Lebensraum ist ein kleiner Vegetationsabschnitt entlang einer ecuadorianischen Autobahn.

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Anolis proboscis ( 英語 )

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A close-up of a male specimen.

Anolis proboscis, commonly known as the horned anole, Ecuadorian horned anole[2] or Pinocchio lizard,[3] is a small anole lizard in the family Dactyloidae. A single male specimen was discovered in 1953 in Ecuador and formally described by Peters and Orces in 1956, but the species then went unreported until its rediscovery in 2004. Its currently known habitat is a small stretch of vegetation along an Ecuadorian highway. It has been classified as Endangered by the IUCN due to its restricted distribution and ongoing habitat loss.

Description

Anolis proboscis has very short limbs and a small body. Four main movement behaviors have been observed: crawling (most common), walking, running, and jumping.[4]

Anolis proboscis has a total body length (excluding tail) of roughly 5–7.5 cm (2.0–3.0 in). Males, which are slightly larger than females, possess a conspicuous proboscis, an elongated structure arising from the middle of the snout and about 1–2 cm (0.4–0.8 in) in length; it appears to be used in courtship displays, as is the dewlap that is present in both sexes.[4] There are multiple colour morphs, with the proboscis ranging from greenish yellow to orange-brown, and variety of patterning over the body, including spots and stripes.[4] Anolis Proboscis bears many resemblances to the Greater Antillean twig anoles, both morphologically, with comparable sexual dimorphism, a slender body, short legs and tail, and in a behavioral sense, residing almost exclusively on narrow surfaces.[4] As such, the species can be classified as a member of the twig ecomorph among Anoles.[4] The species possesses a semi-prehensile tail, which aids them as they climb and forage in the canopy. Although slow moving, they have been known to cover great distances.[4] The species differs from similar species in the region in its comparatively small, whitish dewlap, and a row of enlarged scales that run along the dorsum.[5]

Proboscis

One of the most notable features of the species is its proboscis - a pliable, blunt appendage extending beyond the snout approximately 85% of the head length.[4] Although the exact purpose of the appendage is unknown, male to male combat can be ruled out due to its highly flexible nature. Beyond sexual selection, the proboscis may help males appear larger, which could lend a significant advantage in male to male interactions, despite not acting as an actual weapon.[6] Finally, because of its sexually dimorphic nature, the proboscis' use in thermoregulation or as a lure for hunting prey can also be ruled out.[6] Because the dewlap of A. proboscis is comparatively underdeveloped compared to other Anole species - it is far smaller and less colorful - the appendage could compensate and act as an ornament, as it is prominently displayed during mating rituals.[7] Likely, it serves as an intraspecific communication signal, and could also play a role in territorial displays as a way for males to assert dominance over each other or in mate choice.[6][4]

Distribution and Habitat

The Tumbes-Chocó-Magdalena region of Panama, Colombia, Ecuador, and Peru (from top to bottom).

Based on the few localities known, the species inhabits montane forest habitats in the Chocó, where it keeps to high trees.[1][8] Specimens have predominantly been found in primary and secondary vegetation along a well-traveled dirt road. Only five localities in total are currently known, with a maximum distance of 13 km (8 mi) between the two farthest ones. While the area of occurrence includes pasture land and secondary forest, it is likely that the species also lives in other mid-altitude (1,200–1,650 m or 3,940–5,410 ft) areas in the Ecuadorian Andes.[1] Despite most sightings of them occurring in areas with severe habitat fragmentation and destruction, it can be concluded that they prefer the deep forest where they are far more difficult to spot.[6] The species has been known to occupy all levels of the rainforest, ranging from the groundfloor to the canopy.[4] However, they seem to spend most of their time high off the ground, inhabiting dense vegetation, making them particularly difficult to observe.[6] At night, the lizards roost on small twigs and leaves (less than a centimeter in diameter) between 2.1 and 10 meters off of the ground, which allow them to detect the vibrations of potential predators.[5] Observations in the Mindo area of Ecuador located the species in apple trees and bamboo plants, as well as in disturbed forest habitats.[4]

Conservation

The species is currently classified as Endangered due to its possibly restricted distribution in combination with ongoing threats in the form of habitat loss from logging, human settlement, agriculture, and grazing.[1] Further, because of the species' rarity and unique appearance, it is at risk of poaching for the international pet trade.[5]

The species was feared to be extinct when it was not seen after the original collection in 1953. It was rediscovered in 2004 when an individual was seen and photographed in a cloud forest near Mindo, Ecuador, by a visiting ornithologist.[9] In August 2009, a herpetology expedition from the University of New Mexico located the species in a remote region of Ecuador. In total, they found five individuals including three males and the first two females ever seen and collected.[10] Since 2009, several other expeditions were able to relocate the species in remote regions of Ecuador.[11] However, Anolis proboscis' rarity could be the result of several different factors, including a very niche environment (the canopies of cloud forests), and naturally low populations.[6]

Speciation

Among anoles, only three species have a proboscis, the other being the poorly known and rarely seen Amazonian A. laevis and A. phyllorhinus.[12][8] Despite this similarity and the historical inclusion of all three in the laevis group, A. phyllorhinus does not appear to be closely related to the two others. A. phyllorhinus has a comparatively longer tail and limbs, is green in color, and is not a twig specialist, like A. proboscis.[4] However, these differences do not necessarily suggest a distant relationship, as Anoles are well known for their ecomorphological diversity. Thus, it is likely that the proboscis is not tied to a specific specialization of Anole.[4] However, they do appear to be close relatives and part of the "Phenacosaurus" group. The proboscis is likely the result of convergent evolution.[10][8]

Diet

Butterfly in the Lepidoptera Order in Mindo, Ecuador.

The species is omnivorous and has a diverse diet. The stomach can hold anywhere between 4-16 prey items. A large array of different insects is preyed on, with the primary sources of food including caterpillars, coleopterans(beetles), hemipterans (bugs), dipterans (flies), and hymenoptera (bees and wasps). However, this species also eats pseudoscorpions, larvae, millipedes, spiders, and assorted plant material including flower petals, pieces of wood, and seeds.[5]

Mating

Pinocchio Anolis (Anolis proboscis) in Ecuador (19312208744).jpg

A number of different behaviours connected to mating behaviour have been recorded. Initially it was assumed that the proboscis played a role in reproduction, but this has not been confirmed.[13] A. proboscis follows a polygynous mating system, with a wide variety of visual cues and movements involved in the entirety of the process.[7]

Courtship

In order to attract a mate both males and females display their dewlaps, employing variations of extension lengths. Males have been seen with their dewlaps extended and their proboscis drooped downward while courting.[4] Males are able to slowly change the stiffness and direction of their horns when courting. Other courting behaviors that have been observed include swinging of the head from side to side, known as "proboscis flourishing," and up and down when approaching a female all while the dewlap is displayed, either half or fully extended.[7] Courtship is deemed successful if and when the females runs past the male and continues in the opposite direction which leads to the second stage of mating called chasing. The female may perform a push-up with her forelimbs along with a head-bob and extension of her dewlap in response to the male's courting.[7]

Chasing progresses when the female prompts the male to follow by running past him in the opposite direction. It involves the male following the female in quick short sprints one to three steps long while frequently bobbing their heads in the same pattern as seen in courtship. Chasing ends with the female arching her neck, or when the male begins to climb on her tail with his forelimbs.[7] If the male cannot successfully follow the female, no mating occurs.[7] Similar behavior has been observed in A. carolinensis, suggesting that is a challenge issued by the female as test for the male before mating can begin.[7]

Next, the stimulation stage begins when the male catches the female and the female arches her neck in response, signifying readiness for copulation. The male must mount the female and find a good position to copulate in. As the male does this, he is constantly rubbing her nape with the tip of his snout in order to stimulate the female and to elevate the female's rostral appendage.[7] Males have also been observed biting the female on the nape, which then stopped after copulation.[7]

Copulation

The copulation stage begins after the male and female have found an appropriate position together. Once this happens the male inserts one of his hemipenes into her cloaca.[7] There is no specific time length for copulation but mating ends when the male and female have separated. There are a variety of displays by the male that may occur after copulation, including head bobbing, extension of the dewlap, which the female may or may not respond to.[7]

Reproduction

Anolis Proboscis reproduces oviparously. In captivity, females lay a single egg, which is incubated for roughly 165 days (5.5 months).[5] Interestingly, males are born with a proboscis already formed, which lengthens as they develop.[14]

Conflict between Males

Male agonistic behaviors can occur when two males attempt to court the same female. Non-violent agonistic behaviors from males include attempts to display at each other, chase the other male away, open their mouths to display teeth, arch their neck, raise their backs and probosces, and keep their heads down.[7] They may also bring their rostral appendages against each other's faces, touching the opposing male with the tip or their snout, or perform slow tail undulations.[7] Further, the males have been observed performing a movement resembling a push-up, whereby he moves his body forward with his head and tail down, and then back, raising his head and tail in the process. During this display, the male's dewlaps will be extended, the tail curled up, and the proboscis straight.[14] These behaviors are intended to scare away the other males, potentially by making the male appear larger. They have also been recorded turning sideways and straightening out their body, extending their tongues, perhaps to accentuate their body length.[6] Violent agonistic behaviors can include biting and pushing of other males.[7] Agonistic behaviors can last as long as it takes for the other male to stop attempting courtship. Unlike many other reptile species with prominent proboscises, A. proboscis does not use the appendage as a weapon, and may even attempt to protect it when fighting.[14] During violent male-male interactions, the proboscis remains lifted, and some specimens bear scars, suggesting the appendage can be injured.[14] In general, the species appears to be less aggressive than other kinds of anoles, which falls in line with the behavior of other twig ecomorphs.[14] Unlike species like A. marcanoi or A. carolinensis, there are far fewer attempts at biting or "jaw locking" during male-male conflict.[14]

References

  1. ^ a b c d Arteaga-Navarro, A.; Mayer, G.C.; Poe, S.; Cisneros-Heredia, D.F. (2019). "Anolis proboscis". IUCN Red List of Threatened Species. 2019: e.T178727A18975861. doi:10.2305/IUCN.UK.2019-2.RLTS.T178727A18975861.en. Retrieved 13 November 2021.
  2. ^ "SICB 2015: Anolis proboscis Display Behavior". Anole Annals. 2015-01-06.
  3. ^ "Efforts to save the rare 'Pinocchio lizard' which was once thought extinct". ScienceDaily.
  4. ^ a b c d e f g h i j k l m Losos, Jonathan B.; Woolley, Melissa L.; Mahler, D. Luke; Torres-Carvajal, Omar; Crandell, Kristen E.; Schaad, Eric W.; Narváez, Andrea E.; Ayala-Varela, Fernando; Herrel, Anthony (2012). "Notes on the Natural History of the Little-Known Ecuadorian Horned Anole, Anolis proboscis". Breviora. 531 (1): 1–17. doi:10.3099/531.1. ISSN 0006-9698. S2CID 51899170.
  5. ^ a b c d e Arteaga A (2022) Pinocchio Anole (Anolis proboscis). In: Arteaga A, Bustamante L, Vieira J, Guayasamin JM (Eds) Reptiles of Ecuador: Life in the middle of the world. Available from: www.reptilesofecuador.com
  6. ^ a b c d e f g Inch, Ivan; Koppetsch, Thore; Bohme, Wolfgang (15 May 2022). "Pinocchio lizards and other lizards bearing rostral appendages − the peculiar habitus of the draconine agamid Harpesaurus tricinctus with highlights on its ecological implications and convergence with its New World equivalent, the dactyloid Anolis proboscis" (PDF). Salamandra. 58 (2): 133 – via ResearchGate.
  7. ^ a b c d e f g h i j k l m n Quirola, Diego R.; Mármol, Andrés; Torres-Carvajal, Omar; Narváez, Andrea E.; Ayala-Varela, Fernando; Moore, Ignacio T. (26 July 2017). "Use of a rostral appendage during social interactions in the Ecuadorian Anolis proboscis". Journal of Natural History. 51 (27–28): 1625–1638. doi:10.1080/00222933.2017.1332790. S2CID 91078845.
  8. ^ a b c Yánez-Muñoz, M.H.; M.A. Urgilés; M. Altamirano Benavides; S.R. Cáceres (2010). "Redescripción de Anolis proboscis Peters & Orcés (Reptilia: Polychrotidae), con el descubrimiento de las hembras de la especie y comentarios sobre su distribución y taxonomía". Avances en Ciencias e Ingeniería. 2 (3): 1–14. doi:10.18272/aci.v2i3.39.
  9. ^ Almendariz, A. C.; Vogt, C. (2007). "Anolis proboscis (Squamata: Polychrotidae), Una lagartija rara pero no extinta" (PDF). Politécnica. 27 (4): Biología 7 133–135.
  10. ^ a b Poe, Steven; Ayala, Fernando; Latella, Ian M.; Kennedy, Thomas L.; Christensen, James A.; Gray, Levi N.; Blea, Natalie J.; Armijo, Beeling M.; Schaad, Eric W. (2012). "Morphology, phylogeny, and behavior of Anolis proboscis". Breviora. 530: 1–11. doi:10.3099/530.1. S2CID 34632653.
  11. ^ Main, D. (2013-10-04). "Once 'Extinct' Pinocchio Lizard Pokes His Nose Out". LiveScience.com. Retrieved 2013-10-08.
  12. ^ Rodrigues, M.T.; V. Xavier; G. Skuk; D. Pavan (2002). "New specimens of Anolis phyllorhinus (Squamata, Polychrotidae): the first female of the species and of proboscid anoles". Papéis Avulsos de Zoologia. 42 (16). doi:10.1590/S0031-10492002001600001.
  13. ^ Poe, Steven; Ayala, Fernando; Latella, Ian M.; Kennedy, Thomas L.; Christensen, James A.; Gray, Levi N.; Blea, Natalie J.; Armijo, Beeling M.; Schaad, Eric W. (2012). "Morphology, Phylogeny, and Behavior of Anolis proboscis". Breviora. 530 (1): 1–11. doi:10.3099/530.1. ISSN 0006-9698. S2CID 34632653.
  14. ^ a b c d e f Rodriguez, Diego (November 2015). "Insights into social interactions in the Ecuadorian horned anole, Anolis proboscis" (PDF). Pontifica Universidad Católica del Ecuador Facultad de Ciencias Exactas y Naturales Escuela de Ciencias Biológicas. 1 (1): 8 – via CORE.

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Anolis proboscis: Brief Summary ( 英語 )

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A close-up of a male specimen.

Anolis proboscis, commonly known as the horned anole, Ecuadorian horned anole or Pinocchio lizard, is a small anole lizard in the family Dactyloidae. A single male specimen was discovered in 1953 in Ecuador and formally described by Peters and Orces in 1956, but the species then went unreported until its rediscovery in 2004. Its currently known habitat is a small stretch of vegetation along an Ecuadorian highway. It has been classified as Endangered by the IUCN due to its restricted distribution and ongoing habitat loss.

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Anolis proboscis ( 西班牙、卡斯蒂利亞西班牙語 )

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El anolis cornudo de Mindo[2]​ (Anolis proboscis) es una especie de pequeño lagarto de la familia Dactyloidae. Fue descubierto en 1953 en Ecuador y descrito formalmente por Peters y Orcés en 1956. Su hábitat son bosques nublados entre los 1200 y 1650 m de altitud cerca de Mindo en la provincia de Pichincha, Ecuador. Es una especie arbórea.[1]

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Neonato.

Su cuerpo mide entre 54 y 78 milímetros, y la cola mide algo más que el cuerpo. Los machos presentan un apéndice nasal muy alargado. El saco gular está presente en ambos sexos, aunque en el macho es de mayor tamaño, es de color blanco en las hembras y amarillo blanquecino en los machos.[2][3]​ Su nariz suele medir entre 10 y 20 milímetros.[cita requerida]

El lagarto fue considerado extinto cuando no fue vuelto a encontrar después de las colecciones originales, pero fue redescubierto en 2007.[4]​ Posteriormente, se volvió a describir la especie, se descubrieron más localidades y se profundizó en la comprensión de su historia natural.[3]​ Se encuentra amenazado de extinción debido a la pérdida de su hábitat natural en su reducida área de distribución.[1]

Referencias

  1. a b c Mayer, G.C. & Poe, S. (2013). «Anolis proboscis». Lista Roja de especies amenazadas de la UICN 2022 (en inglés). ISSN 2307-8235. Consultado el 6 de octubre de 2013.
  2. a b Ayala-Varela, F., Carvajal-Campos, A. (2010). «Anolis proboscis.». En O. Torres-Carvajal, D. Salazar-Valenzuela y A. Merino-Viteri, ed. ReptiliaWebEcuador. Version 2013.0. Museo de Zoología QCAZ, Pontificia Universidad Católica del Ecuador. Archivado desde el original el 9 de enero de 2013. Consultado el 24 de marzo de 2016.
  3. a b Poe, Steven; Ayala, Fernando; Latella, Ian M.; Kennedy, Thomas L.; Christensen, James A.; Gray, Levi N.; Blea, Natalie J.; Armijo, Beeling M. et al. (2012). «Morphology, phylogeny, and behavior of Anolis proboscis». Breviora: 1-11.
  4. Almendariz, A. C.; Vogt, C. (2007). «Anolis proboscis (Squamata: Polychrotidae), Una lagartija rara pero no extinta». Politécnica 27(4) Biología 7: 133-135.
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Anolis proboscis: Brief Summary ( 西班牙、卡斯蒂利亞西班牙語 )

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El anolis cornudo de Mindo​ (Anolis proboscis) es una especie de pequeño lagarto de la familia Dactyloidae. Fue descubierto en 1953 en Ecuador y descrito formalmente por Peters y Orcés en 1956. Su hábitat son bosques nublados entre los 1200 y 1650 m de altitud cerca de Mindo en la provincia de Pichincha, Ecuador. Es una especie arbórea.​

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Su cuerpo mide entre 54 y 78 milímetros, y la cola mide algo más que el cuerpo. Los machos presentan un apéndice nasal muy alargado. El saco gular está presente en ambos sexos, aunque en el macho es de mayor tamaño, es de color blanco en las hembras y amarillo blanquecino en los machos.​​ Su nariz suele medir entre 10 y 20 milímetros.[cita requerida]

El lagarto fue considerado extinto cuando no fue vuelto a encontrar después de las colecciones originales, pero fue redescubierto en 2007.​ Posteriormente, se volvió a describir la especie, se descubrieron más localidades y se profundizó en la comprensión de su historia natural.​ Se encuentra amenazado de extinción debido a la pérdida de su hábitat natural en su reducida área de distribución.​

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Anolis proboscis ( 巴斯克語 )

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Anolis proboscis: Brief Summary ( 巴斯克語 )

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Anolis proboscis Anolis generoko animalia da. Narrastien barruko Dactyloidae familian sailkatuta dago.

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Anolis proboscis ( 法語 )

由wikipedia FR提供

Anolis proboscis ou lézard Pinocchio est une espèce de sauriens de la famille des Dactyloidae[1].

Répartition

Cette espèce est endémique de la province de Pichincha en Équateur[1]. Sa présence au Pérou est incertaine. Il n'avait été aperçu que 3 fois en cinquante ans[2] avant l'année 2013 quand des photographes animaliers réussirent à photographier plusieurs individus dans la forêt équatorienne[2], le faisant classer comme espèce en danger d'extinction par l'UICN.

Étymologie

Le nom spécifique proboscis vient du latin proboscis, la trompe, en référence à l'appendice souple présent à l'extrémité du museau de cette espèce. Son nom vernaculaire de Pinocchio fait aussi référence à cette excroissance[2].

Publication originale

  • Peters & Orcés, 1956 : A third leaf-nosed species of the lizard genus Anolis from South America. Breviora, no 62, p. 1-8 (texte intégral).

Notes et références

  1. a et b Reptarium Reptile Database, consulté lors d'une mise à jour du lien externe
  2. a b et c « Le retour du lézard Pinocchio », Le Monde, supplément Sciences & Médecine,‎ 16 octobre 2013, p. 6.
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Anolis proboscis: Brief Summary ( 法語 )

由wikipedia FR提供

Anolis proboscis ou lézard Pinocchio est une espèce de sauriens de la famille des Dactyloidae.

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Anolis proboscis ( 義大利語 )

由wikipedia IT提供

Anolis proboscis (Peters & Orces, 1956) è un piccolo sauro della famiglia Dactyloidae[2], endemico dell'Ecuador.

Descrizione

È un piccolo sauro, lungo da 54 a 78 mm[3], caratterizzato da un marcato dimorfismo sessuale: il maschio presenta una caratteristica protuberanza in corrispondenza del naso che gli ha valso, in lingua inglese, la denominazione di "Pinocchio anole".

Distribuzione e habitat

L'areale di questa specie è limitato al versante occidentale delle Ande nei pressi della città di Mindo, ad altitudini comprese tra i 1 200 e 1 650 m s.l.m.). Occupa una superficie di circa 33 km2 con non più di cinque località di avvistamento[3]. L'entità della popolazione non è nota, anche se alcuni zoologi hanno osservato una relativa abbondanza di esemplari in alcuni siti[1].

L'habitat è quello di foresta pluviale montana tipico della regione andina nella provincia del Pichincha.

Conservazione

La specie fu scoperta nel 1953, nella provincia del Pichincha, ma non fu più vista dagli anni 1960 al 2005, tanto da farla considerare estinta per un lungo periodo di tempo.[4] Al 2011, secondo lo IUCN, il suo stato di conservazione rimane comunque problematico: la specie, infatti, è classificata in pericolo di estinzione a causa dell'areale ridotto e della distruzione dell'habitat dovute all' attività di disboscamento operata dall'uomo[1].

Note

  1. ^ a b c (EN) Mayer, G.C. & Poe, S. 2011, Anolis proboscis, su IUCN Red List of Threatened Species, Versione 2020.2, IUCN, 2020. URL consultato il 10 febbraio 2016.
  2. ^ (EN) Anolis proboscis, in The Reptile Database. URL consultato il 10 febbraio 2016.
  3. ^ a b (ES) Yánez-Muñoz M.H., Urgilés M.A., Altamirano B.M. and Cáceres S.S.R., Redescripción de Anolis proboscis Peters & Orcés (Reptilia: Polychrotidae), con el descubrimiento de las hembras de la especie y comentarios sobre su distribución y taxonomía, in Avances en Ciencias e Ingenierías, vol. 2, n. 3, 2010, pp. B7-B15.
  4. ^ (EN) Rediscovery Of The Enigmatic Ecuadorian Horned Anole, su Anole Annals - Your source for the latest on Anolis lizards. URL consultato il 10 febbraio 2016.

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Anolis proboscis: Brief Summary ( 義大利語 )

由wikipedia IT提供

Anolis proboscis (Peters & Orces, 1956) è un piccolo sauro della famiglia Dactyloidae, endemico dell'Ecuador.

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Anolis proboscis ( 波蘭語 )

由wikipedia POL提供

Anolis proboscisgatunek jaszczurki z rodziny Dactyloidae charakteryzującej się wyrostkiem na pysku, występującym tylko u samców.

Systematyka i morfologia

Gad ten zalicza się do rodzaju Anolis, należącego do rodziny Dactyloidae[3]. Dawniejsza systematyka zaliczała go do licznej w gatunki rodziny legwanowatych (Iguanidae)[4][5].

Epitet gatunkowy odnosi się do przypominającego trąbę wyrostka sterczącego z pyska jaszczurki. Odgrywa on rolę w dobieraniu się w pary[3].

Rozmieszczenie geograficzne

Anolis ten występuje na średniej wysokości zachodnich stoków Andów w Ekwadorze. Zamieszkuje tereny leżące nie niżej niż 1200 i nie wyżej niż 1650 m n.p.m.[3].

Jaszczurki te żyją w górskich lasach, znajdywano je jednak też w okolicach pastewnych, lasach wtórnych i przy drogach[3].

Zagrożenia i ochrona

W niektórych okolicach gatunek występuje bardzo obficie[3].

Przypisy

  1. Anolis proboscis, w: Integrated Taxonomic Information System (ang.).
  2. J.A. Peters & V.G. Orces. A third leaf-nosed species of the lizard genus Anolis from South America. „Breviora”. 62, 1956 (ang.).
  3. a b c d e f Anolis proboscis. Czerwona księga gatunków zagrożonych (IUCN Red List of Threatened Species) (ang.).
  4. Gady łuskonośne − jaszczurki, amfisbeny i węże. W: Hanna Dobrowolska: gady. Warszawa: Państwowe Wydawnictwo Naukowe, 1981, s. 279, seria: zwierzęta świata. ISBN 83-01-00957-8.
  5. Systematyka gadów. W: Hanna Dobrowolska: gady. Warszawa: Państwowe Wydawnictwo Naukowe, 1981, s. 475, seria: zwierzęta świata. ISBN 83-01-00957-8.
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Anolis proboscis: Brief Summary ( 波蘭語 )

由wikipedia POL提供

Anolis proboscis – gatunek jaszczurki z rodziny Dactyloidae charakteryzującej się wyrostkiem na pysku, występującym tylko u samców.

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Anolis proboscis ( 摩爾多瓦語 )

由wikipedia RO提供

Anolis proboscis[2] este o specie de șopârle din genul Anolis, familia Polychrotidae, descrisă de Wilhelm Peters și Orces 1956.[3][4] A fost clasificată de IUCN ca specie în pericol.[1] Conform Catalogue of Life specia Anolis proboscis nu are subspecii cunoscute.[3]

Referințe

  1. ^ a b Anolis proboscis. Lista roșie a speciilor periclitate IUCN. Versiunea 2012.2. International Union for Conservation of Nature. 2011. Accesat în 24 octombrie 2012.
  2. ^ Peters,J.A. & Orces V,G. (1956) A third leaf-nosed species of the lizard genus Anolis from South America., Breviora 62: 1-8
  3. ^ a b Bisby F.A., Roskov Y.R., Orrell T.M., Nicolson D., Paglinawan L.E., Bailly N., Kirk P.M., Bourgoin T., Baillargeon G., Ouvrard D. (red.) (2011). „Species 2000 & ITIS Catalogue of Life: 2011 Annual Checklist”. Species 2000: Reading, UK. Accesat în 24 september 2012. Verificați datele pentru: |access-date= (ajutor)Mentenanță CS1: Nume multiple: lista autorilor (link)
  4. ^ TIGR Reptile Database . Uetz P. , 2007-10-02
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Anolis proboscis: Brief Summary ( 摩爾多瓦語 )

由wikipedia RO提供

Anolis proboscis este o specie de șopârle din genul Anolis, familia Polychrotidae, descrisă de Wilhelm Peters și Orces 1956. A fost clasificată de IUCN ca specie în pericol. Conform Catalogue of Life specia Anolis proboscis nu are subspecii cunoscute.

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匹诺曹蜥蜴 ( 漢語 )

由wikipedia 中文维基百科提供
二名法 Anolis proboscis
Peters and Orces, 1956

匹诺曹蜥蜴Pinocchio lizardPinocchio anole,學名:Anolis proboscis)是一种小蜥蜴安乐蜥属安乐蜥科。仅在南美洲厄瓜多尔境内小镇Mindo附近发现过。

外形特征

雄性蜥蜴有长鼻子,因此叫匹诺曹蜥蜴,鼻子不是笨重而坚硬的结构,而是相当灵活。

种群历史

1953年前一直未对其正式描述。生物学家当时仅采集到6只雄性个体,后来数年内又多次发现,后再未发现。不过在过去的15年间民间有少数几次目击记录。哈佛大学进化生物学家和爬虫研究专家乔纳森·洛索斯(Jonathan Losos)曾对该生物进行过研究。他说:“在长达40年的时间里,没有任何人报告见到过这种蜥蜴。因此在当时我们曾经认为这种蜥蜴已经灭绝了。”[2]

重新发现

2005年,一群观鸟人在小镇Mindo附近偶然发现一只外表怪异的蜥蜴横穿一条马路。他们回家后将照片上传于互联网,爬虫学家们认出是匹诺曹安乐蜥。数个考察组前往当地再次搜索,其中一个由美国新墨西哥大学史蒂夫·波尔(Steve Poe)教授与另外一位专家领衔的考察小组发现了他们。发现的雌性个体没有长鼻子。[3]目击者阿里詹德罗·安提瓜(Alejandro Arteaga)是一名摄影师,他说:“很难描述发现这种蜥蜴时的感受。发现这条‘匹诺曹’变色龙就像是发现了一个秘密,一个深藏不露的秘密。很长时期以来我们都一直认为这是一种传说中的动物。”

习性

匹诺曹蜥蜴喜欢在夜间爬到枝根部睡觉,此时身体的颜色会转为灰白色,色彩伪装效果极好,平时生活在树冠层顶端,移动速度非常缓慢,几乎难以察觉。有考察队表示:“我们发现这种蜥蜴生活的环境与文献描述有很大的出入。比如这种蜥蜴还从未被在远离开阔区域的密林深处被发现过,其它的几次目击事件也都是在靠近森林边缘的位置。”

科学家此前在巴西境内发现过与之相似的蜥蜴,但是这两种蜥蜴是在相互独立的条件下进化出各自的长鼻子的。至于长鼻子作用,还没有确切知道。[4]

参考

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匹诺曹蜥蜴: Brief Summary ( 漢語 )

由wikipedia 中文维基百科提供

匹诺曹蜥蜴(Pinocchio lizard或Pinocchio anole,學名:Anolis proboscis)是一种小蜥蜴安乐蜥属安乐蜥科。仅在南美洲厄瓜多尔境内小镇Mindo附近发现过。

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