Eleocharis compressa Sull.

Eleocharis compressa is extremely variable. After E. bifida and E. occulta are segregated from E. compressa (type from Ohio) and E. acutisquamata (type from Texas), the remaining plants intergrade completely and are best treated as varieties of E. compressa (S. G. Smith 2001). Because some plants of E. compressa approach E. elliptica some authors have treated E. compressa as a synonym of E. elliptica. Putative E. compressa × E. erythropoda hybrids from southeastern Ontario have been described (P. M. Catling 1994), and I have observed similar putative hybrids in the field in southeastern Wisconsin. Introgression from E. erythropoda is suggested by the presence in many plants of E. compressa of some 2-fid styles and biconvex achenes and some or rarely all entire floral scales, including the type. Some collections of E. compressa from the District of Columbia region and Tennessee have distinctly cancellate achenes as is common in E. tenuis var. verrucosa. The earlier name E. acuminata (Muhlenberg) Nees, name confused (H. K. Svenson 1932), is often given as a synonym of E. compressa (S. G. Smith 2001).

Both 2n = 24 and 2n = 26 cytotypes for E. compressa from south-central United States have been reported (L. J. Harms 1972). Fertile artificial hybrids between these two cytotypes were also reported, as well as artificial E. compressa (2n = 36) × E. elliptica (2n = 38) hybrids with meiotic pairing and pollen stainability more than 90 percent. I have not seen voucher specimens for those artificial hybrids or for any of the published chromosone number reports for E. compressa, except for a count of 2n = 24 (presumably made by L. J. Harms) that is given on a herbarium specimen label of E. compressa var. acutisquamata from Kansas (S. G. Smith 2001).

Plants perennial, mat-forming; rhizomes evident, short to rather long, 2–3 mm thick, firm, cortex persistent, internodes very short to 2 mm, scales often decaying to coarse fibers, 5–12 mm, papery. Culms subterete to greatly compressed, to 5 times as wide as thick, often with 2–12 ridges, 8–45 cm × (0.2–)0.5–1.8 mm, firm to hard, spongy. Leaves: distal leaf sheaths persistent, not splitting, proximally red, distally green to stramineous, papery, apex usually red-brown, broadly obtuse to sub-truncate, callose, tooth absent. Spikelets ovoid, 4–8 × 2–4 mm, apex acute; proximal scale clasping 3/4 of culm to amplexicaulous, entire; subproximal scale empty or with flower; floral scales spreading in fruit, 20–60, 6 per mm of rachilla, medium brown to very dark brown, sometimes stramineous or colorless, midrib region often paler, ovate, 2–3(–4) × 1–1.5 mm, apex acute to acuminate, the apical colorless hyaline region mostly longer than wide and 0.6–1.2 mm, in proximal part of spikelet 2-fid to shallowly cut, rarely entire, in distal part often entire, carinate in distal part of spikelet. Flowers: perianth bristles 0–5, stramineous, length variable, to equaling achene, obscurely to clearly retrorsely spinulose; stamens 3; anthers yellow- to orange-brown, 0.7–2 mm; styles 3-fid or some 2-fid. Achenes falling with scales or some persistent, yellow-brown or dark brown, obovoid to obpyriform, nearly equilaterally to compressed-trigonous or usually some biconvex, angles evident or obscure, 0.8–1.1 × 0.6–0.8 mm, neck very short, finely rugulose and sometimes finely (or coarsely) cancellate at 10–20X with 14–20 blunt horizontal ridges in a vertical series. Tubercles brown (to whitish), pyramidal, usually depressed, often rudimentary, 0.15–0.3 × 0.2–0.35 mm.

Eleocharis elliptica Kunth var. compressa (Sullivant) Drepalik & Mohlenbrock

Eleocharis compressa Sull. Am. Jour. Sci 42 : 50. 1842.
?Scirpus acuminatus Muhl. Descr. Gram. 27. 1817. (Pennsylvania.)
Eleocharis acuminata Nees, Linnaea 9 : 294. 1834.
Eleocharis compressa var. atrata Svenson, Rhodora 34: 218. 1932. (Pennsylvania.)
Culms strongly flattened, 1-2 mm. in width, striate, erect from creeping rootstocks sometimes nearly 1 cm. thick; upper sheaths 2-9 cm. high, reddish-brown to stramineous, truncate, faintly to prominently toothed; spikelets 5-12 mm. long, oblong-ovate to ovate, acute or obtuse; scales ovate-lanceolate, chestnut-brown (or black in var. atrata), the whitened tips attenuate and commonly bifid; style trifid; achene 1-1.5 mm. long, obovate-pyriform, golden-yellow to brown, bluntly trigonous to nearly terete, granular-roughened or reticulate under magnification, the raised margins of the cells often forming undulating lines; stylebase depressedto globose-conic, usually acute; bristles 1-5, fugacious, usually shorter than the achene.
(Sullivlnt)^°^^^^^^ ' ^^* ^^^^^^ '" ^^^ ^^^^^ ^^^'"®' ^^ '"'^^^ ^^^* °^ Columbus, Ohio
Distribution :_ Ontario to central New York, western New Jersey, Washington, D. C, west to

Eleocharis compressa là loài thực vật có hoa trong họ Cói. Loài này được Sull. mô tả khoa học đầu tiên năm 1842.