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Bigyra ( Catalan; Valensian )

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Bigyra és un agrupament d'Heterokonts.[1]

Inclou Bicosoecida, Blastocystis i Labyrinthulida.[1]

També s'ha descrit com contenent Opalozoa, Bicoecia, i Sagenista.[2]

Referències

  1. 1,0 1,1 Riisberg I, Orr RJ, Kluge R, et al. «Seven gene phylogeny of heterokonts». Protist, 160, 2, maig 2009, pàg. 191–204. DOI: 10.1016/j.protis.2008.11.004. PMID: 19213601.
  2. Cavalier-Smith T, Chao EE «Phylogeny and megasystematics of phagotrophic heterokonts (kingdom Chromista)». J. Mol. Evol., 62, 4, abril 2006, pàg. 388–420. DOI: 10.1007/s00239-004-0353-8. PMID: 16557340.
 src= A Wikimedia Commons hi ha contingut multimèdia relatiu a: Bigyra Modifica l'enllaç a Wikidata
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Bigyra: Brief Summary ( Catalan; Valensian )

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Bigyra és un agrupament d'Heterokonts.

Inclou Bicosoecida, Blastocystis i Labyrinthulida.

També s'ha descrit com contenent Opalozoa, Bicoecia, i Sagenista.

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Bigyra ( Frisian setentrional )

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Amrum.pngTekst üüb Öömrang
 src=
Aplanochytrium sp. oner't elektroonenmikroskoop

Bigyra san en stam faan Chromista uun di auerstam faan a Heterokonta. Det san miast algen.

Iindialang

Kwelen

  • Ruggiero, M.A., Gordon, D.P., Orrell, T.M., Bailly, N., Bourgoin, T., Brusca, R.C., Cavalier-Smith, T., Guiry, M. D. & Kirk, P. M. (2015). A Higher Level Classification of All Living Organisms. PLoS ONE 10(4) online

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Bigyra ( Antërlenga (Assossiassion Antërnassional për na Lenga Ausiliaria) )

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Bigyra es un phylo de Heterokonta.

Nota
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Bigyra: Brief Summary ( Frisian setentrional )

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 src= Aplanochytrium sp. oner't elektroonenmikroskoop

Bigyra san en stam faan Chromista uun di auerstam faan a Heterokonta. Det san miast algen.

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Bygira ( Bosnìach )

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Bigyra (Opalozoa) je bazni kladus heterokontnih organizama. Preci Bigyra i vjerojatno svih heterokonta bili su slobodnoživeći fagotrofni organizmi. Bicosoecea i Placididea su jedine grupe u ovom redu koje imaju pradavni način ishrane, putem fagotrofije. Nakon toga, Opalinata i Blastocystea su izmijenile fagotrofgnost predaka sekundarnom osmotrofijom kada su postali životinje ili paraziti životinja.

S druge strane, Labyrinthulea su vjerovatno neovisno izgubiile sposobnost fagotrofije, kada su se prilagodile fungoidnom saprobnom načinu života.[1][2]

Filogenija

Bigyra je parafiletska bazna grupa Heterokonta i predstavlja evolucijski stepen na kojem je heterotrofni prekursor pseudogljiva i okrofitne grupe algi. Sastoji se od kladusa Sagenista i Opalozoa, uspostavljajući sljedeće odnose:[3]

Heterokonta Sagenista

Labyrinthulomycetes

   

Eogyrea / Amphitremidae (?)

      Opalozoa

Bikosea

  Placidozoa  

Placididea

     

Nanomonadea

     

Opalomonadea

     

Blastocystea

   

Opalinea

               

Pseudofungi

   

Ochrophyta

       

Također pogledajte

Reference

  1. ^ Cavalier-Smith, T., & Chao, E. E. (2006). Phylogeny and megasystematics of phagotrophic heterokonts (kingdom Chromista). Journal of molecular evolution, 62(4), 388-420.
  2. ^ Riisberg I, Orr RJ, Kluge R. "Seven gene phylogeny of heterokonts" (PDF). Protist. 160 (2): 191–204. doi:10.1016/j.protis.2008.11.004. PMID 19213601. Pristupljeno May 1, 2009.CS1 održavanje: koristi se parametar authors (link)
  3. ^ Silar, P. (2016). Protistes Eucaryotes.

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Bygira: Brief Summary ( Bosnìach )

fornì da wikipedia emerging languages

Bigyra (Opalozoa) je bazni kladus heterokontnih organizama. Preci Bigyra i vjerojatno svih heterokonta bili su slobodnoživeći fagotrofni organizmi. Bicosoecea i Placididea su jedine grupe u ovom redu koje imaju pradavni način ishrane, putem fagotrofije. Nakon toga, Opalinata i Blastocystea su izmijenile fagotrofgnost predaka sekundarnom osmotrofijom kada su postali životinje ili paraziti životinja.

S druge strane, Labyrinthulea su vjerovatno neovisno izgubiile sposobnost fagotrofije, kada su se prilagodile fungoidnom saprobnom načinu života.

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Bigyra ( Anglèis )

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Bigyra (from Latin bi- 'twice', and gyrus 'circle')[1] is a possibly paraphyletic phylum of heterotrophic organisms belonging to the Stramenopile lineage.[5][2] It includes the groups Bicosoecida, Opalinata and Labyrinthulea, as well as several small clades that were initially discovered through environmental DNA samples: Nanomonadea, Placididea, Opalomonadea and Eogyrea.[3][6]

Diversity and ecology

Bigyra is a widely distributed group of stramenopile lineages, characterized by an anterior flagellum with tripartite hairs. It contains well-known ecological groups such as the fungi-like slime nets, the flagellate bicosoecids and the opalines.[7]

Slime nets

The slime nets (known as Labyrinthulea, Labyrinthulomycetes or Labyrinthulomycota) are fungus-like heterotrophic, colorless or yellowish protists that absorb nutrients in an osmotrophic or phagotrophic manner, either as free-living amoebae or as networks of anastomosing cytoplasmic threads that extend from a bothrosome. They are typically saprotrophic decomposers of the detrital food web, making organic matter more accessible to grazing protists. Some are parasitic, and others feed on bacteria. They are cosmopolitan, ubiquitous in marine, freshwater and estuarine environments, associated with algae, marine plants and detritus.[7]

Opalines

The opalines (known as Opalinata) are a diverse assemblage of modified unicellular protists, consisting of three closely-related groups: proteromonads, opalinids and Blastocystis. They inhabit the intestines of various animals and can be found in all continents.[7]

Bicosoecids

The bicosoecids (known as Bicosoecida) are a small group of marine or freshwater heterotrophic flagellates that feed on bacteria. Their classification has changed multiple times over the years,[8] and is still an unresolved issue.[9]

History and phylogeny

Origin

Bigyra was first described in 1997 by Thomas Cavalier-Smith, as a phylum within Heterokonta that contained three subphyla: 1) the walled saprotrophic Pseudofungi, 2) the non-phagotrophic gut-symbiotic Opalinata, and 3) the phagotrophic zooflagellate Developayella, which received its own subphylum Bigyromonada. These groups would have originated from a common ancestor that had a double ciliary transition zone helix as its synapomorphy. The common ancestor would have evolved from photosynthetic heterokonts, but would have secondarily lost its plastids, as opposed to the photosynthetic Ochrophyta which retain them. Bigyra was, at the time, postulated as a monophyletic group (or clade), followed by a paraphyletic grade of ochrophyte classes.[1][10]

Heterokonta Bigyra Pseudofungi

Oomycetes

Hyphochytrea

Bigyromonadea

Opalinata

Proteromonadea

Opalinea

Limnistia

Sarcinochrysia

Dictyochia

Fucistia

Diatomeae

Sagenista

Taxonomic modifications

Posterior phylogenetic analyses that used 18S rRNA genes revealed that Pseudofungi and Bigyromonadea were more closely related to Ochrophyta than they were to Opalinata, meaning that the synapomorphy of a double helix could have been present in the common ancestor of all heterokonts. This rendered Bigyra paraphyletic. Consequently, Bigyra was revised and modified in 2006 to comprise a different set of three subphyla: 1) Opalozoa, a previously polyphyletic diverse phylum that was modified to only include Opalinata and Nucleohelea; 2) Bicoecea, containing the bicosoecids; and 3) Sagenista, containing the osmotrophic Labyrinthulea. The phylogeny of Bigyra, however, could not be resolved, and its monophyly was weakly supported. The weak support was thought to be caused by all three bigyran lineages diverging from each other very soon after the separation from other heterokonts; this deep branching makes it difficult to find the exact branching order of bigyran clades.[2]

Heterokonta Ochrophyta

Khakista

Limnista

Pseudofungi

Bigyromonadea

Hyphochytrea

Oomycetes

Opalozoa

Nucleohelea

Opalinata

Bicoecea

Sagenista

Bigyra

Bigyra was modified again in 2013 after the discovery of several heterokont clades called MAST (‘marine stramenopiles’), recovered through environmental rDNA sequencing. The subphylum Opalozoa assimilated the bicosoecids in an infraphylum Bikosia, while another new infraphylum Placidozoa assimilated the Opalinata and an array of new clades: Placididea, Nanomonadea (MAST-3) and Opalomonadea (MAST-12), all three classified under the paraphyletic taxon Wobblata. The subphylum Sagenista, on the other hand, received a new class Eogyrea that was composed of several MAST lineages not yet described.[3] Later, one of the MAST clades within Eogyrea would be described as Pseudophyllomitus (MAST-6).[6]

Stramenopiles Gyrista

Ochrophyta

Pseudofungi

Opalozoa Placidozoa Opalinata

Opalinea

Blastocystea

Opalomonadea

Nanomonadea

Placididea

Bikosea

Sagenista

Eogyrea

Labyrinthulea

Current phylogeny

Despite modern large-scale phylogenomic analyses that use bigger taxon sampling, the validity of Bigyra remains uncertain. The positions of the deep-branching bigyran clades are not consistent among the published studies, and not all the clades are well-represented by genomic and transcriptomic data.[6] Several studies support the monophyly of Bigyra through multi-gene phylogenetic analysis and a rich taxon sampling.[6][11] There is also support for its paraphyly from more recent phylogenetic studies of the 2020s decade.[12][13]

Taxonomy

The present classification of Bigyra is as follows:[4][11]

References

  1. ^ a b c Cavalier-Smith T (1998). "A revised six-kingdom system of life". Biol Rev Camb Philos Soc. 73 (3): 203–66. doi:10.1111/j.1469-185X.1998.tb00030.x. PMID 9809012. S2CID 6557779.
  2. ^ a b c Cavalier-Smith T, Chao EE (April 2006). "Phylogeny and megasystematics of phagotrophic heterokonts (kingdom Chromista)". J. Mol. Evol. 62 (4): 388–420. doi:10.1007/s00239-004-0353-8. PMID 16557340. S2CID 29567514.
  3. ^ a b c d e Cavalier-Smith, Thomas; Scoble, Josephine Margaret (2013). "Phylogeny of Heterokonta: Incisomonas marina, a uniciliate gliding opalozoan related to Solenicola (Nanomonadea), and evidence that Actinophryida evolved from raphidophytes". European Journal of Protistology. 49 (3): 328–353. doi:10.1016/j.ejop.2012.09.002. PMID 23219323.
  4. ^ a b Adl SM, Bass D, Lane CE, Lukeš J, Schoch CL, Smirnov A, Agatha S, Berney C, Brown MW, Burki F, Cárdenas P, Čepička I, Chistyakova L, del Campo J, Dunthorn M, Edvardsen B, Eglit Y, Guillou L, Hampl V, Heiss AA, Hoppenrath M, James TY, Karnkowska A, Karpov S, Kim E, Kolisko M, Kudryavtsev A, Lahr DJG, Lara E, Le Gall L, Lynn DH, Mann DG, Massana R, Mitchell EAD, Morrow C, Park JS, Pawlowski JW, Powell MJ, Richter DJ, Rueckert S, Shadwick L, Shimano S, Spiegel FW, Torruella G, Youssef N, Zlatogursky V, Zhang Q (2019). "Revisions to the Classification, Nomenclature, and Diversity of Eukaryotes". Journal of Eukaryotic Microbiology. 66 (1): 4–119. doi:10.1111/jeu.12691. PMC 6492006. PMID 30257078.
  5. ^ Riisberg I, Orr RJ, Kluge R, et al. (May 2009). "Seven gene phylogeny of heterokonts". Protist. 160 (2): 191–204. doi:10.1016/j.protis.2008.11.004. PMID 19213601.
  6. ^ a b c d Thakur, Rabindra; Shiratori, Takashi; Ishida, Ken-ichiro (2019). "Taxon-rich Multigene Phylogenetic Analyses Resolve the Phylogenetic Relationship Among Deep-branching Stramenopiles". Protist. 170 (5): 125682. doi:10.1016/j.protis.2019.125682. ISSN 1434-4610.
  7. ^ a b c d e f Archibald, John M.; Simpson, Alastair G. B.; Slamovits, Claudio H., eds. (2017). Handbook of the Protists (2 ed.). Springer International Publishing. pp. ix. ISBN 978-3-319-28147-6.
  8. ^ Karpov SA, Sogin ML, Silberman JD (2001). "Rootlet homology, taxonomy, and phylogeny of bicosoecids based on 18S rRNA gene sequences". Protistology. 2 (1): 34–47.
  9. ^ Schoenle A, Hohlfeld M, Rybarski A, Sachs M, Freches E, Wiechmann K, Nitsche F, Arndt H (2022). "Cafeteria in extreme environments: Investigations on C. burkhardae and three new species from the Atacama Desert and the deep ocean". European Journal of Protistology. 85: 125905. doi:10.1016/j.ejop.2022.125905.
  10. ^ Cavalier-Smith T (1998). "Sagenista and Bigyra, two phyla of heterotrophic heterokont chromists". Archiv für Protistenkunde. 148 (3): 253–267. doi:10.1016/S0003-9365(97)80006-1.
  11. ^ a b Cavalier-Smith, Thomas (2017). "Kingdom Chromista and its eight phyla: a new synthesis emphasising periplastid protein targeting, cytoskeletal and periplastid evolution, and ancient divergences". Protoplasma. 255 (1): 297–357. doi:10.1007/s00709-017-1147-3. PMC 5756292. PMID 28875267.
  12. ^ Tan MH, Loke S, Croft LJ, Gleason FH, Lange L, Pilgaard B, Trevathan-Tackett SM (2021). "First Genome of Labyrinthula sp., an Opportunistic Seagrass Pathogen, Reveals Novel Insight into Marine Protist Phylogeny, Ecology and CAZyme Cell-Wall Degradation". Microbial Ecology. 82: 498–511. doi:10.1007/s00248-020-01647-x.
  13. ^ Cho A, Tikhonenkov DV, Hehenberger E, Karnkowska A, Mylnikov AP, Keeling PJ (2022). "Monophyly of diverse Bigyromonadea and their impact on phylogenomic relationships within stramenopiles" (PDF). Molecular Phylogenetics and Evolution. 171 (107468): 107468. doi:10.1016/j.ympev.2022.107468. ISSN 1055-7903. S2CID 247815732.
  14. ^ Shiratori, Takashi; Thakur, Rabindra; Ishida, Ken-ichiro (2017). "Pseudophyllomitus vesiculosus (Larsen and Patterson 1990) Lee, 2002, a Poorly Studied Phagotrophic Biflagellate is the First Characterized Member of Stramenopile Environmental Clade MAST-6". Protist. 168 (4): 439–451. doi:10.1016/j.protis.2017.06.004. ISSN 1434-4610. PMID 28822908.

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Bigyra: Brief Summary ( Anglèis )

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Bigyra (from Latin bi- 'twice', and gyrus 'circle') is a possibly paraphyletic phylum of heterotrophic organisms belonging to the Stramenopile lineage. It includes the groups Bicosoecida, Opalinata and Labyrinthulea, as well as several small clades that were initially discovered through environmental DNA samples: Nanomonadea, Placididea, Opalomonadea and Eogyrea.

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Bigyra ( Spagneul; Castilian )

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Los bigiros (Bigyra) son un filo parafilético de organismos heterocontos. El antepasado de Bigyra y probablemente de todos los heterocontos era un organismo fagótrofo de vida libre. Bicosoecea y Placididea son los únicos grupos de esta línea que retienen el modo de alimentación ancestral mediante fagotrofia. Posteriormente, Opalinata y Blastocystea reemplazaron la capacidad fagótrofa ancestral por una osmotrofia secundaria cuando se convirtieron en comensales o parásitos de los animales. Por su parte, Labyrinthulea probablemente perdió su capacidad fagotrofa independientemente cuando se adaptó a un modo de vida saprobia de tipo fungoide.[3][4][2]

Filogenia

Bigyra es un grupo parafilético basal de los heterokontos y representa un grado evolutivo heterótrofo precursor del grupo de los pseudohongos y de las algas ocrofitas. Está constituido por los clados Sagenista, Opalozoa y Platysulcea estableciéndose las siguientes relaciones:[2][5][6]

Bigyra

Platysulcea

     

Labyrinthulea

    Opalozoa

Bikosea

  Placidozoa  

Placididea

     

Nanomonadea

     

Opalomonadea

     

Blastocystea

   

Opalinea

               

Pseudofungi

   

Ochrophyta

         

Referencias

  1. https://www.irmng.org/aphia.php?p=taxdetails&id=11912885
  2. a b c Thomas Cavalier-Smith & Josephine Scoble 2012, Phylogeny of Heterokonta: Incisomonas marina, a uniciliate gliding opalozoan related to Solenicola (Nanomonadea), and evidence that Actinophryida evolved from raphidophytes.  European Journal of Protistology 49(3) · December 2012 with 432 DOI: 10.1016/j.ejop.2012.09.002
  3. Cavalier-Smith, T., & Chao, E. E. (2006). Phylogeny and megasystematics of phagotrophic heterokonts (kingdom Chromista). Journal of molecular evolution, 62(4), 388-420.
  4. Riisberg I, Orr RJ, Kluge R, etal (mayo de 2009). «Seven gene phylogeny of heterokonts». Protist 160 (2): 191-204. PMID 19213601. doi:10.1016/j.protis.2008.11.004.
  5. Silar, P. (2016). Protistes Eucaryotes.
  6. Shiratori, Takashi; Nakayama, Takeshi; Ishida, Ken-ichiro (2015). «A New Deep-branching Stramenopile, Platysulcus tardus gen. nov., sp. nov.». Protist 166 (3): 337-348. ISSN 1434-4610. PMID 26070192. doi:10.1016/j.protis.2015.05.001. hdl:2241/00148461.
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Bigyra: Brief Summary ( Spagneul; Castilian )

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Los bigiros (Bigyra) son un filo parafilético de organismos heterocontos. El antepasado de Bigyra y probablemente de todos los heterocontos era un organismo fagótrofo de vida libre. Bicosoecea y Placididea son los únicos grupos de esta línea que retienen el modo de alimentación ancestral mediante fagotrofia. Posteriormente, Opalinata y Blastocystea reemplazaron la capacidad fagótrofa ancestral por una osmotrofia secundaria cuando se convirtieron en comensales o parásitos de los animales. Por su parte, Labyrinthulea probablemente perdió su capacidad fagotrofa independientemente cuando se adaptó a un modo de vida saprobia de tipo fungoide.​​​

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Bigyra ( Fransèis )

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Les Bigyra sont un embranchement d'organismes eucaryotes hétérocontées.

Taxinomie

Références

  • (en) Cavalier-Smith T., 1997. Sagenista and Bigyra, two phyla of heterotrophic heterokont chromists. Archiv für Protistenkunde. 148 (3): 253–267, .
  • (en) Cavalier-Smith T., 1998. A revised six-kingdom system of life. Biol Rev Camb Philos Soc. 73 (3): 203–66, , PMID 9809012.

Références taxinomiques

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Bigyra: Brief Summary ( Fransèis )

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Les Bigyra sont un embranchement d'organismes eucaryotes hétérocontées.

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Bigyra ( polonèis )

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ilustracja Systematyka Domena Eukaryota Klad Bikonta Klad SAR Klad Halvaria Klad Heterokonta Klad Bigyra
Cavalier-Smith, 1998[1]

Bigyratakson heterotroficznych organizmów jądrowych w grupie Heterokonta[2].

Należą tu Bicosoecida, Blastocystis oraz Labyrinthulida[2].

Bywa także opisywany jako takson zawierający Opalozoa, Bicoecia oraz Sagenista, wchodzący w skład królestwa Chromista[3].

Przypisy

  1. M. D. Guiry, G.M. Guiry: AlgaeBase. World-wide electronic publication, National University of Ireland, Galway. 2013. [dostęp 2013-10-15].
  2. a b I. Riisberg, R. J. Orr, R. Kluge et al.. Seven gene phylogeny of heterokonts. „Protist”. 160 (2), s. 191–204, maj 2009. DOI: 10.1016/j.protis.2008.11.004. PMID: 19213601.
  3. T. Cavalier-Smith, E. E. Chao. Phylogeny and megasystematics of phagotrophic heterokonts (kingdom Chromista). „J. Mol. Evol.”. 62 (4), s. 388–420, kwiecień 2006. DOI: 10.1007/s00239-004-0353-8. PMID: 16557340.
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Bigyra: Brief Summary ( polonèis )

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Bigyra – takson heterotroficznych organizmów jądrowych w grupie Heterokonta.

Należą tu Bicosoecida, Blastocystis oraz Labyrinthulida.

Bywa także opisywany jako takson zawierający Opalozoa, Bicoecia oraz Sagenista, wchodzący w skład królestwa Chromista.

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Bigyra ( portughèis )

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Bigyra, ou Opalozoa, é um cladode organismos heterocontes considerado o grupo basal dos Heterokontophyta.

Descrição

O antepassado de Bigyra, e provavelmente de todos os heterocontes, era um organismo fagotrófico de vida livre. Bicosoecida e Placididea são os únicos grupos desta linhagem que retiveram o modo de alimentação ancestral fagotrófica. Posteriormente, Opalinata e Blastocystea substituíram a capacidade fagotrófica ancestral por uma osmotrofia secundária quando se converteram em comensais ou parasitas dos animais. Por seu lado, os Labyrinthulea provavelmente perderam a sua capacidade fagotrófica independentemente quando se adaptaram a um modo de vida sapróbio do tipo fungoide.[3][4][5]

Filogenia

Como inicialmente definido, Bigyra era um grupo parafilético basal dos heterocontes que representa um grado evolutivo heterotrófico precursor do grupo dos Pseudofungi e das algas ocrófitas. O grupo é constituído pelos clados Sagenista e Opalozoa, estabelecendo-se as seguintes relações:[2][6]

Heterokonta Sagenista

Labyrinthulomycetes



Eogyrea / Amphitremidae (?)




Opalozoa

Bikosea


Placidozoa

Placididea




Nanomonadea




Opalomonadea




Blastocystea



Opalinea









Pseudofungi



Ochrophyta





Notas

  1. Adl, S.M. et al. (2012). The revised classification of eukaryotes. Journal of Eukaryotic Microbiology, 59(5), 429-514
  2. a b Thomas Cavalier-Smith & Josephine Scoble 2012, Phylogeny of Heterokonta: Incisomonas marina, a uniciliate gliding opalozoan related to Solenicola (Nanomonadea), and evidence that Actinophryida evolved from raphidophytes.  European Journal of Protistology 49(3) · December 2012 with 432 DOI: 10.1016/j.ejop.2012.09.002
  3. Cavalier-Smith, T., & Chao, E. E. (2006). Phylogeny and megasystematics of phagotrophic heterokonts (kingdom Chromista). Journal of molecular evolution, 62(4), 388-420.
  4. Riisberg I, Orr RJ, Kluge R et al. (maio de 2009). «Seven gene phylogeny of heterokonts» (PDF). Protist. 160 (2): 191–204. PMID 19213601. doi:10.1016/j.protis.2008.11.004
  5. Cavalier-Smith, Thomas, and Josephine Margaret Scoble. Phylogeny of Heterokonta: Incisomonas marina, a uniciliate gliding opalozoan related to Solenicola (Nanomonadea), and evidence that Actinophryida evolved from raphidophytes. European journal of protistology 49.3 (2013): 328-353.
  6. Silar, P. (2016). Protistes Eucaryotes.
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Bigyra: Brief Summary ( portughèis )

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Bigyra, ou Opalozoa, é um cladode organismos heterocontes considerado o grupo basal dos Heterokontophyta.

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双环总门 ( cinèis )

fornì da wikipedia 中文维基百科

见正文

双环总门(Bigyra)是一个不等鞭毛类的子类群[1]

包括BicosoeceaLabyrinthulomycetes[1]OpalineaProteromonadea以及芽囊原虫纲(Blastocystis)[2]

参考文献

  1. ^ 1.0 1.1 Riisberg I, Orr RJ, Kluge R; 等. Seven gene phylogeny of heterokonts. Protist. May 2009, 160 (2): 191–204. PMID 19213601. doi:10.1016/j.protis.2008.11.004. 引文格式1维护:显式使用等标签 (link)
  2. ^ Cavalier-Smith T, Chao EE. Phylogeny and megasystematics of phagotrophic heterokonts (kingdom Chromista). J. Mol. Evol. April 2006, 62 (4): 388–420. PMID 16557340. doi:10.1007/s00239-004-0353-8.
双鞭毛生物多貌生物AH原始色素体生物(A),或广义植物 定鞭/隐藻生物(H)(有争议) SAR不等鞭毛/囊泡生物不等鞭毛类(S) 囊泡虫类(A) 有孔虫界(R) 古虫界 无根虫门英语Apusozoa 单鞭毛生物变形虫界 后鞭毛生物动物总界Mesomycetozoea英语Mesomycetozoea 蜷丝生物英语Filozoa蜷丝球虫纲英语Filasterea
领鞭毛虫纲 动物界或
“后生动物”
菌物总界真菌界 核形虫类


小作品圖示这是一篇與生物相關的小作品。你可以通过编辑或修订扩充其内容。
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双环总门: Brief Summary ( cinèis )

fornì da wikipedia 中文维基百科

双环总门(Bigyra)是一个不等鞭毛类的子类群 。

包括BicosoeceaLabyrinthulomycetesOpalineaProteromonadea以及芽囊原虫纲(Blastocystis) 。

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비기라 ( Corean )

fornì da wikipedia 한국어 위키백과

비기라(Bigyra)는 부등편모조류 분류의 일종이다.[1] 비코소에카류, 블라스토키스티스류, 망형충류를 포함하고 있다.[1] 또한 오팔충류, 사게니스타류를 포함하는 것으로 기술되기도 한다.[2]

하위 분류

각주

  1. Riisberg I, Orr RJ, Kluge R; 외. (2009년 5월). “Seven gene phylogeny of heterokonts”. 《Protist》 160 (2): 191–204. doi:10.1016/j.protis.2008.11.004. PMID 19213601.
  2. Cavalier-Smith T, Chao EE (2006년 4월). “Phylogeny and megasystematics of phagotrophic heterokonts (kingdom Chromista)”. 《J. Mol. Evol.》 62 (4): 388–420. doi:10.1007/s00239-004-0353-8. PMID 16557340.
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