定義： aquatic organisms that construct and inhabit simple unbranched vertical shafts with only one opening to the surface. These are primarily above-sediment filter-feeders, surface deposit-feeders, and subsurface deposit-feeders (head-down, conveyor-belt feeders)
定義： An organism which obtains food primarily by subsurface deposit feeding. Reid WDK, Wigham BD, McGill RAR, Polunin NVC (2012) Elucidating trophic pathways in benthic deep-sea assemblages of the Mid-Atlantic Ridge north and south of the Charlie-Gibbs Fracture Zone. Mar Ecol Prog Ser 463:89-103. doi:10.3354/meps09863
