定義： aquatic organisms that construct and inhabit simple unbranched vertical shafts with only one opening to the surface. These are primarily above-sediment filter-feeders, surface deposit-feeders, and subsurface deposit-feeders (head-down, conveyor-belt feeders)
定義： Upward conveyors are vertically oriented species that typically feed head-down at depth in the sediment. Vertically oriented head-down feeders actively select and ingest particles at the deeper sediments and egest these non-locally as faeces in the sediment surface
