Eugenia gandhii N. Snow

Shrubs to trees, 3–12 m. Trunk 3–8 cm dbh; outer bark of main bole reddish, papery to flakey and peeling. Foliage and reproductive parts glabrous except where noted. Branchlets terete to laterally compressed, reddish when fresh but drying brownish or grayish, glabrous but prominently glandular, the epidermis smooth but soon peeling (dried specimens) and often thin and narrow grayish strips or even threads. Leaves opposite, medium green (fresh), slightly to strongly discolorous, venation brochidodromous, thinly coriaceous (easily cracking when dried), surfaces matte. Axillary colleters lacking. Petioles 2.8–6 mm, slightly sulcate distally. Leaf blades (5.5–)9–16 × 2.5–6.0 cm, elliptic to occasionally obovate, base cuneate, apex acute to mostly acuminate or caudate, margins flat; adaxial surface glabrous, densely punctate (glands small and best seen with magnification); abaxial surface similar, lateral veins indistinct to prominent; intramarginal vein 1–3.5 mm from margin at midpoint of blade. Inflorescence terminal, axillary, or ramiflorous, of soliltary monads or in fascicles of 2–3 flowers. Pedicels 9–29 mm long × 0.8–1.2 mm thick, stiff, moderately glandular, ascending to erect, green. Bracteoles narrowly ovate to broadly rounded, 0.5–1.3 × 1.0–1.2 mm, stiff but thin, glabrous to minutely and sparsely sericeous on margin, often persistent in fruit. Hypanthium 4–5 mm long, cupuliform, densely glandular (some glands larger proximally). Calyx lobes 4, 6–9 × up to 11 mm, broadly elliptic to rounded, cream-colored (fresh), reflexed in flower, persistent and often crowning the fruit. Petals 4, 11–14 × up to 12 mm, widely elliptic to widely obovate (narrowly greatly at base), sparsely and minute ciliate marginally, densely glandular (glands of variable size but typically large proximally). Stamens 250–300, multiseriate, exserted; staminal disk short-hairy (trichomes slightly ferrugineous); filaments 8–15 mm, white; anthers globose, 0.3–0.5 mm, brownish, bearing a single large apical gland. Ovary apex glabrous but somewhat glandular. Style 8–10 mm; stigma narrow (scarcely if at all swollen). Berries 2.5–4.0 × 3.0–4.0 cm, subglobose to globose, purplish or violet (drying nearly black). Locules 2; placentation axile. Seeds up to 20 mm long and wide (available material possibly not fully mature), up to 6 per fruit, evidently often somewhat flattened, cotyledons and hypocotyl barely if at all differentiated.

Endemic to northeast Madagascar, recorded from the Island of Nosy Mangabe, the Masoala Peninsula, and in and around the Anjaniharibe Reserve (Figure 11).