Definition: aquatic organisms that construct and inhabit simple unbranched vertical shafts with only one opening to the surface. These are primarily above-sediment filter-feeders, surface deposit-feeders, and subsurface deposit-feeders (head-down, conveyor-belt feeders)
Definition: Downward conveyors exhibit a feeding strategy opposite to that of upward conveyors. Vertically oriented head-up feeders actively select and ingest particles at the surface and egest these non-locally as faeces in deeper sediment strata