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Image of Ataenius gracilis (Melsheimer 1845)
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Ataenius gracilis (Melsheimer 1845)

Scientific Names

Preferred Names

Ataenius gracilis (Melsheimer 1845)
Species recognized by EOL Dynamic Hierarchy April 2022
Reference taxon from World Scarabaeidae Database in Species 2000 & ITIS Catalogue of Life
Ataenius gracilis
Species recognized by Costa Rica Species List, Dominican Republic Species List, Nicaragua Species List, Peru Species List, Puerto Rico Species List, Mexico Species List, North Atlantic Species List, North Pacific Species List, and United States Species List
Ataenius gracilis
Recognized by Furui 2020
Ataenius gracilis (Melsheimer 1845)
Species recognized by EOL Dynamic Hierarchy 1.1, EOL Dynamic Hierarchy 1.1, GBIF classification, and Global Register of Introduced and Invasive Species
Ataenius gracilis Melsheimer (1844)
Recognized by IABIN
Ataenius gracilis
Species recognized by wikidata_hierarchy tar gz, wikipedia PT, wikipedia EN, Global Biotic Interactions, iNaturalist data coverage, and GGBN data coverage
Ataenius gracilis
Recognized by BHL data coverage
Ataenius gracilis (Melsheimer 1845)
Species recognized by Catalogue of Life 26 March 2024, iNaturalist, Integrated Taxonomic Information System (ITIS) Taxonomic Hierarchy, GBIF data coverage, Dynamic Hierarchy Version 2.2.3, and EOL Dynamic Hierarchy Version 2.2.4
Ataenius gracilis (Melsheimer 1846)
Species recognized by TreatmentBank
Ataenius gracilis (Melsheimer)
Recognized by Smithsonian Contributions to Zoology
Saprosites nocturnus Nomura
Recognized by NMNH type specimens

Alternative Names

Saprosites nocturnus Nomura 1943
Not accepted according to EOL Dynamic Hierarchy April 2022
Reference taxon from World Scarabaeidae Database in Species 2000 & ITIS Catalogue of Life
Saprosites nocturnus Nomura 1943
Synonym according to EOL Dynamic Hierarchy 1.1 and EOL Dynamic Hierarchy 1.1
Oxyomus gracilis Melsheimer 1845
Synonym according to Catalogue of Life 26 March 2024
Saprosites nocturnus Nomura 1943
Invalid according to Integrated Taxonomic Information System (ITIS) Taxonomic Hierarchy
Saprosites nocturnus Nomura 1943
Not accepted according to Dynamic Hierarchy Version 2.2.3 and EOL Dynamic Hierarchy Version 2.2.4
Saprosites nocturnus Nomura 1943
Synonym according to Catalogue of Life 26 March 2024

Common Names

Japanese

コツツマグソコガネ preferred
Recognized by Furui 2020

Curated hierarchies for Ataenius gracilis (Melsheimer 1845)

EOL Dynamic Hierarchy April 2022

Life
Cellular Organisms
Eukaryota
Opisthokonta
Metazoa
Bilateria
Protostomia
Ecdysozoa
Arthropoda
Pancrustacea
Hexapoda
Insecta
Pterygota
Neoptera
Endopterygota
Coleoptera
Polyphaga
Scarabaeiformia
Scarabaeoidea
Aphodiidae
Ataenius
Ataenius gracilis (Melsheimer 1845) (this page)
Ataenius aequalis Harold 1880
Ataenius africanus Endrödi 1967
Ataenius ambaritae Stebnicka 1988
Ataenius angulatus Petrovitz 1964
Ataenius apicalis Hinton 1936
Ataenius arrowi Hinton 1936
Ataenius asper Petrovitz 1975
Ataenius atramentarius (Erichson 1847)
Ataenius barberi Cartwright 1974
Ataenius beattyi Chapin 1940
Ataenius brouni (Sharp 1876)
Ataenius buenavistae Stebnicka 2001
Ataenius cartago Stebnicka 2007
Ataenius castaniellus Bates 1887
Ataenius catarinaensis Stebnicka 2007
Ataenius clavatus Schmidt 1916
Ataenius confertus Fall 1909
Ataenius coriarius Schmidt 1912
Ataenius corporaali Boucomont 1924
Ataenius costulifer Balthasar 1941
Ataenius crenatipennis (MacLeay 1871)
Ataenius dentatus Koshantschikov 1916
Ataenius ecruensis Stebnicka & Lago 2005
Ataenius elegans Harold 1868
Ataenius elongatus (Palisot de Beauvois 1811)
Ataenius eurynotus Lea 1923
Ataenius excisicollis Petrovitz 1961
Ataenius fattigi Cartwright 1948
Ataenius floreanae Cook & Peck 2000
Ataenius gammonensis Stebnicka & Howden 1997
Ataenius garamas Peyerimhoff 1929
Ataenius gilesi Stebnicka & Howden 1997
Ataenius glaseri Cartwright 1974
Ataenius gruneri Chalumeau 1979
Ataenius guayasi Stebnicka 2001
Ataenius guriensis Stebnicka 2005
Ataenius heinekeni (Wollaston 1854)
Ataenius hesperius Cartwright 1974
Ataenius horticola Harold 1869
Ataenius howdeni Chalumeau 1978
Ataenius indutus Petrovitz 1961
Ataenius inquisitus Horn 1887
Ataenius insulae Chalumeau & Gruner 1974
Ataenius integricollis Lea 1923
Ataenius jamaicensis Chapin 1940
Ataenius jardinensis Stebnicka 2002
Ataenius kapalgaensis Stebnicka & Howden 1997
Ataenius koebelei Blackburn 1904
Ataenius koelleri Balthasar 1963
Ataenius lobatus Horn 1871
Ataenius luctuosus (Burmeister 1877)
Ataenius luteomargo Chapin 1940
Ataenius martinezi Petrovitz 1973
Ataenius miamii Cartwright 1934
Ataenius millstreamae Stebnicka & Howden 1997
Ataenius monticola Paulian 1940
Ataenius montreuili Bordat & Théry 2012
Ataenius nakpandurii Endrödi 1973
Ataenius nigricans Paulian 1933
Ataenius noronhai Stebnicka 2007
Ataenius nugator Harold 1880
Ataenius oaxacaensis Stebnicka & Lago 2005
Ataenius ocumarensis Stebnicka 2001
Ataenius onkonensis Stebnicka 2005
Ataenius ovatulus Horn 1871
Ataenius palmaritoensis Stebnicka 2007
Ataenius palustris (Montrouzier 1860)
Ataenius panamensis Hinton 1936
Ataenius peregrinator Harold 1877
Ataenius pereirai Petrovitz 1970
Ataenius picinus Harold 1867
Ataenius plaumanni Petrovitz 1973
Ataenius pseudoclavatus Stebnicka 2005
Ataenius pseudostercorator Stebnicka 2003
Ataenius punctatohirsutus Schmidt 1909
Ataenius raccurti Chalumeau 1978
Ataenius raslani Bordat & Théry 2012
Ataenius rubrotessellatus (Blanchard 1846)
Ataenius saltae Stebnicka 2007
Ataenius scabrellus Schmidt 1909
Ataenius scutellaris Harold 1867
Ataenius siminasus Petrovitz 1973
Ataenius skelleyi Stebnicka 2007
Ataenius sparsicollis Blackburn 1904
Ataenius spretulus (Haldeman 1848)
Ataenius stebnickae Bordat & Théry 2012
Ataenius steinheili Harold 1874
Ataenius stephani Cartwright 1974
Ataenius stercorator (Fabricius 1775)
Ataenius strigatus (Say 1823)
Ataenius strigifrons Schmidt 1920
Ataenius suineata Deloya 2012
Ataenius synnotensis Stebnicka & Howden 1997
Ataenius tarumensis Stebnicka 2007
Ataenius texanus Harold 1874
Ataenius tindalensis Stebnicka & Howden 1997
Ataenius tomentosus Petrovitz 1969
Ataenius usingeri Hinton 1936
Ataenius warisensis Stebnicka 1998
Ataenius windjanae Stebnicka & Howden 1997
182 additional siblings truncated for brevity.
See the resource file for a full list.

Catalogue of Life 26 March 2024

Biota
Animalia
Arthropoda
Insecta
Coleoptera
Scarabaeoidea
Scarabaeidae Latreille 1802
Aphodiinae Leach 1815
Eupariini Schmidt 1910
Ataenius Harold 1867
Ataenius gracilis (Melsheimer 1845) (this page)
Ataenius abancay Stebnicka 2005
Ataenius abditus (Haldeman 1848)
Ataenius aciculus Hinton 1936
Ataenius acutulus (Schmidt 1914)
Ataenius aequalis Harold 1880
Ataenius aequatorialis Petrovitz 1961
Ataenius africanus Endrödi 1967
Ataenius alternatus (Melsheimer 1845)
Ataenius ambaritae Stebnicka 1988
Ataenius angulatus Petrovitz 1964
Ataenius annanus Stebnicka & Howden 1997
Ataenius aphodiiformis Bordat & Théry 2012
Ataenius apicalis Hinton 1936
Ataenius arenosus Harold 1868
Ataenius arrowi Hinton 1936
Ataenius asper Petrovitz 1975
Ataenius atramentarius (Erichson 1847)
Ataenius attenuator Harold 1874
Ataenius australasiae (Boheman 1858)
Ataenius australis Harold 1875
Ataenius balthasari Petrovitz 1973
Ataenius barberi Cartwright 1974
Ataenius basiceps Lea 1923
Ataenius beattyi Chapin 1940
Ataenius benarabyensis Stebnicka & Howden 1997
Ataenius bicolor Petrovitz 1963
Ataenius biroi (Endrödi 1951)
Ataenius bispinulosus Schmidt 1911
Ataenius blapoides Balthasar 1947
Ataenius bolivarensis Stebnicka 2007
Ataenius bolivari Stebnicka 2001
Ataenius borgmeieri Hinton 1936
Ataenius borjae Stebnicka 2005
Ataenius brevicollis (Wollaston 1854)
Ataenius brevis Fall 1930
Ataenius brevitarsis Petrovitz 1964
Ataenius brouni (Sharp 1876)
Ataenius buenavistae Stebnicka 2001
Ataenius caesoides (Fairmaire 1900)
Ataenius caicarae Stebnicka 2005
Ataenius californicus Horn 1887
Ataenius canoasus Stebnicka 2007
Ataenius carinatipennis Petrovitz 1973
Ataenius carinator Harold 1874
Ataenius cartago Stebnicka 2007
Ataenius cartwrighti Chalumeau & Gruner 1974
Ataenius castaniellus Bates 1887
Ataenius catarinaensis Stebnicka 2007
Ataenius catenulatus (Erichson 1847)
Ataenius cavagnaroi Cartwright 1970
Ataenius ceylonensis Schmidt 1912
Ataenius chapini Hinton 1937
Ataenius chinacotae Stebnicka 2007
Ataenius clavatus Schmidt 1917
Ataenius crenatostriatus (Blanchard 1845)
Ataenius cribrithorax Bates 1887
Ataenius ecruensis Stebnicka & Lago 2005
Ataenius elongatus (Palisot de Beauvois 1811)
Ataenius europaeus Quiel 1910
Ataenius exiguus Brown 1932
Ataenius freyi Petrovitz 1961
Ataenius granocostatus Schmidt 1912
Ataenius hygrophilus Paulian 1947
Ataenius imbricatoides Schmidt 1909
Ataenius impiger Schmidt 1917
Ataenius koebelei Blackburn 1904
Ataenius languidus Schmidt 1910
Ataenius maghribinicus Baraud 1985
Ataenius montanus Schmidt 1910
Ataenius monticola Paulian 1940
Ataenius opatroides (Blanchard 1847)
Ataenius palmaritoensis Stebnicka 2007
Ataenius peregianensis Stebnicka & Howden 1997
Ataenius peruvianus Minkina 2020
Ataenius platensis (Blanchard 1847)
Ataenius restructus (Wickham 1912)
Ataenius rubrotessellatus (Blanchard 1847)
Ataenius schmidti Stebnicka 2003
Ataenius siminasus Petrovitz 1973
Ataenius stephani Cartwright 1974
Ataenius suineata Deloya 2012
Ataenius synnotensis Stebnicka & Howden 1997
Ataenius talpoides Stebnicka 2001
Ataenius tambopatae Stebnicka 2001
Ataenius tarumensis Stebnicka 2007
Ataenius temperei Chalumeau & Gruner 1974
Ataenius terminalis (Chevrolat 1865)
Ataenius texanus Harold 1874
Ataenius thomasi Schnepp & Ashman 2020
Ataenius tindalensis Stebnicka & Howden 1997
Ataenius tomentosus Petrovitz 1969
Ataenius torridus Blackburn 1892
Ataenius tovarensis Stebnicka 2001
Ataenius tuberculatus Schmidt 1911
Ataenius uriarrae Stebnicka & Howden 1997
Ataenius usingeri Hinton 1936
Ataenius utahensis Cartwright 1974
Ataenius versicolor Schmidt 1917
Ataenius walkeri Blackburn 1904
Ataenius windjanae Stebnicka & Howden 1997
189 additional siblings truncated for brevity.
See the resource file for a full list.

Dynamic Hierarchy Version 2.2.3

Ataenius gracilis (Melsheimer 1845) (this page)
Ataenius aequalis Harold 1880
Ataenius africanus Endrödi 1967
Ataenius ambaritae Stebnicka 1988
Ataenius angulatus Petrovitz 1964
Ataenius apicalis Hinton 1936
Ataenius arrowi Hinton 1936
Ataenius asper Petrovitz 1975
Ataenius atramentarius (Erichson 1847)
Ataenius barberi Cartwright 1974
Ataenius beattyi Chapin 1940
Ataenius brouni (Sharp 1876)
Ataenius buenavistae Stebnicka 2001
Ataenius cartago Stebnicka 2007
Ataenius castaniellus Bates 1887
Ataenius catarinaensis Stebnicka 2007
Ataenius clavatus Schmidt 1916
Ataenius confertus Fall 1909
Ataenius coriarius Schmidt 1912
Ataenius corporaali Boucomont 1924
Ataenius costulifer Balthasar 1941
Ataenius crenatipennis (MacLeay 1871)
Ataenius dentatus Koshantschikov 1916
Ataenius ecruensis Stebnicka & Lago 2005
Ataenius elegans Harold 1868
Ataenius elongatus (Palisot de Beauvois 1811)
Ataenius eurynotus Lea 1923
Ataenius excisicollis Petrovitz 1961
Ataenius fattigi Cartwright 1948
Ataenius floreanae Cook & Peck 2000
Ataenius gammonensis Stebnicka & Howden 1997
Ataenius garamas Peyerimhoff 1929
Ataenius gilesi Stebnicka & Howden 1997
Ataenius glaseri Cartwright 1974
Ataenius gruneri Chalumeau 1979
Ataenius guayasi Stebnicka 2001
Ataenius guriensis Stebnicka 2005
Ataenius heinekeni (Wollaston 1854)
Ataenius hesperius Cartwright 1974
Ataenius horticola Harold 1869
Ataenius howdeni Chalumeau 1978
Ataenius indutus Petrovitz 1961
Ataenius inquisitus Horn 1887
Ataenius insulae Chalumeau & Gruner 1974
Ataenius integricollis Lea 1923
Ataenius jamaicensis Chapin 1940
Ataenius jardinensis Stebnicka 2002
Ataenius kapalgaensis Stebnicka & Howden 1997
Ataenius koebelei Blackburn 1904
Ataenius koelleri Balthasar 1963
Ataenius lobatus Horn 1871
Ataenius luctuosus (Burmeister 1877)
Ataenius luteomargo Chapin 1940
Ataenius martinezi Petrovitz 1973
Ataenius miamii Cartwright 1934
Ataenius millstreamae Stebnicka & Howden 1997
Ataenius monticola Paulian 1940
Ataenius montreuili Bordat & Théry 2012
Ataenius nakpandurii Endrödi 1973
Ataenius nigricans Paulian 1933
Ataenius noronhai Stebnicka 2007
Ataenius nugator Harold 1880
Ataenius oaxacaensis Stebnicka & Lago 2005
Ataenius ocumarensis Stebnicka 2001
Ataenius onkonensis Stebnicka 2005
Ataenius ovatulus Horn 1871
Ataenius palmaritoensis Stebnicka 2007
Ataenius palustris (Montrouzier 1860)
Ataenius panamensis Hinton 1936
Ataenius peregrinator Harold 1877
Ataenius pereirai Petrovitz 1970
Ataenius picinus Harold 1867
Ataenius plaumanni Petrovitz 1973
Ataenius pseudoclavatus Stebnicka 2005
Ataenius pseudostercorator Stebnicka 2003
Ataenius punctatohirsutus Schmidt 1909
Ataenius raccurti Chalumeau 1978
Ataenius raslani Bordat & Théry 2012
Ataenius rubrotessellatus (Blanchard 1846)
Ataenius saltae Stebnicka 2007
Ataenius scabrellus Schmidt 1909
Ataenius scutellaris Harold 1867
Ataenius siminasus Petrovitz 1973
Ataenius skelleyi Stebnicka 2007
Ataenius sparsicollis Blackburn 1904
Ataenius spretulus (Haldeman 1848)
Ataenius stebnickae Bordat & Théry 2012
Ataenius steinheili Harold 1874
Ataenius stephani Cartwright 1974
Ataenius stercorator (Fabricius 1775)
Ataenius strigatus (Say 1823)
Ataenius strigifrons Schmidt 1920
Ataenius suineata Deloya 2012
Ataenius synnotensis Stebnicka & Howden 1997
Ataenius tarumensis Stebnicka 2007
Ataenius texanus Harold 1874
Ataenius tindalensis Stebnicka & Howden 1997
Ataenius tomentosus Petrovitz 1969
Ataenius usingeri Hinton 1936
Ataenius warisensis Stebnicka 1998
Ataenius windjanae Stebnicka & Howden 1997
182 additional siblings truncated for brevity.
See the resource file for a full list.

EOL Dynamic Hierarchy 1.1

Life
Cellular Organisms
Eukaryota
Opisthokonta
Metazoa
Bilateria
Protostomia
Ecdysozoa
Panarthropoda
Arthropoda
Pancrustacea
Hexapoda
Insecta
Pterygota
Neoptera
Endopterygota
Coleoptera
Polyphaga
Scarabaeiformia
Scarabaeoidea
Aphodiidae
Ataenius
Ataenius gracilis (Melsheimer 1845) (this page)
Ataenius aequalis Harold 1880
Ataenius africanus Endrödi 1967
Ataenius ambaritae Stebnicka 1988
Ataenius angulatus Petrovitz 1964
Ataenius apicalis Hinton 1936
Ataenius arrowi Hinton 1936
Ataenius asper Petrovitz 1975
Ataenius atramentarius (Erichson 1847)
Ataenius barberi Cartwright 1974
Ataenius beattyi Chapin 1940
Ataenius brouni (Sharp 1876)
Ataenius buenavistae Stebnicka 2001
Ataenius cartago Stebnicka 2007
Ataenius castaniellus Bates 1887
Ataenius catarinaensis Stebnicka 2007
Ataenius clavatus Schmidt 1916
Ataenius confertus Fall 1909
Ataenius coriarius Schmidt 1912
Ataenius corporaali Boucomont 1924
Ataenius costulifer Balthasar 1941
Ataenius crenatipennis (Mac Leay 1871)
Ataenius damzeni Bukejs & Alekseev 2018
Ataenius dentatus Koshantschikov 1916
Ataenius ecruensis Stebnicka & Lago 2005
Ataenius elegans Harold 1868
Ataenius elongatus (Palisot de Beauvois 1811)
Ataenius europaeus Quiel 1910
Ataenius eurynotus Lea 1923
Ataenius excisicollis Petrovitz 1961
Ataenius fattigi Cartwright 1948
Ataenius floreanae Cook & Peck 2000
Ataenius gammonensis Stebnicka & Howden 1997
Ataenius garamas Peyerimhoff 1929
Ataenius gilesi Stebnicka & Howden 1997
Ataenius glaseri Cartwright 1974
Ataenius gruneri Chalumeau 1979
Ataenius guayasi Stebnicka 2001
Ataenius guriensis Stebnicka 2005
Ataenius heinekeni (Wollaston 1854)
Ataenius hesperius Cartwright 1974
Ataenius horticola Harold 1869
Ataenius howdeni Chalumeau 1978
Ataenius indutus Petrovitz 1961
Ataenius inquisitus Horn 1887
Ataenius insulae Chalumeau & Gruner 1974
Ataenius integricollis Lea 1923
Ataenius jamaicensis Chapin 1940
Ataenius jardinensis Stebnicka 2002
Ataenius kapalgaensis Stebnicka & Howden 1997
Ataenius koebelei Blackburn 1904
Ataenius koelleri Balthasar 1963
Ataenius lobatus Horn 1871
Ataenius luteomargo Chapin 1940
Ataenius martinezi Petrovitz 1973
Ataenius miamii Cartwright 1934
Ataenius millstreamae Stebnicka & Howden 1997
Ataenius monticola Paulian 1940
Ataenius montreuili Bordat & Théry 2012
Ataenius nakpandurii Endrödi 1973
Ataenius nigricans Paulian 1933
Ataenius noronhai Stebnicka 2007
Ataenius nugator Harold 1880
Ataenius oaxacaensis Stebnicka & Lago 2005
Ataenius ocumarensis Stebnicka 2001
Ataenius onkonensis Stebnicka 2005
Ataenius ovatulus Horn 1871
Ataenius palmaritoensis Stebnicka 2007
Ataenius palustris (Montrouzier 1860)
Ataenius panamensis Hinton 1936
Ataenius peregrinator Harold 1877
Ataenius pereirai Petrovitz 1970
Ataenius picinus Harold 1867
Ataenius plaumanni Petrovitz 1973
Ataenius pseudoclavatus Stebnicka 2005
Ataenius pseudostercorator Stebnicka 2003
Ataenius raccurti Chalumeau 1978
Ataenius raslani Bordat & Théry 2012
Ataenius rubrotessellatus (Blanchard 1846)
Ataenius saltae Stebnicka 2007
Ataenius scabrellus Schmidt 1909
Ataenius scutellaris Harold 1867
Ataenius siminasus Petrovitz 1973
Ataenius skelleyi Stebnicka 2007
Ataenius sparsicollis Blackburn 1904
Ataenius spretulus (Haldeman 1848)
Ataenius stebnickae Bordat & Théry 2012
Ataenius steinheili Harold 1874
Ataenius stephani Cartwright 1974
Ataenius stercorator (Fabricius 1775)
Ataenius strigatus (Say 1823)
Ataenius strigifrons Schmidt 1920
Ataenius suineata Deloya 2012
Ataenius synnotensis Stebnicka & Howden 1997
Ataenius tarumensis Stebnicka 2007
Ataenius texanus Harold 1874
Ataenius tindalensis Stebnicka & Howden 1997
Ataenius tomentosus Petrovitz 1969
Ataenius usingeri Hinton 1936
Ataenius warisensis Stebnicka 1998
Ataenius windjanae Stebnicka & Howden 1997
186 additional siblings truncated for brevity.
See the resource file for a full list.

EOL Dynamic Hierarchy Version 2.2.4

Ataenius gracilis (Melsheimer 1845) (this page)
Ataenius aequalis Harold 1880
Ataenius africanus Endrödi 1967
Ataenius ambaritae Stebnicka 1988
Ataenius angulatus Petrovitz 1964
Ataenius apicalis Hinton 1936
Ataenius arrowi Hinton 1936
Ataenius asper Petrovitz 1975
Ataenius atramentarius (Erichson 1847)
Ataenius barberi Cartwright 1974
Ataenius beattyi Chapin 1940
Ataenius brouni (Sharp 1876)
Ataenius buenavistae Stebnicka 2001
Ataenius cartago Stebnicka 2007
Ataenius castaniellus Bates 1887
Ataenius catarinaensis Stebnicka 2007
Ataenius clavatus Schmidt 1916
Ataenius confertus Fall 1909
Ataenius coriarius Schmidt 1912
Ataenius corporaali Boucomont 1924
Ataenius costulifer Balthasar 1941
Ataenius crenatipennis (MacLeay 1871)
Ataenius dentatus Koshantschikov 1916
Ataenius ecruensis Stebnicka & Lago 2005
Ataenius elegans Harold 1868
Ataenius elongatus (Palisot de Beauvois 1811)
Ataenius eurynotus Lea 1923
Ataenius excisicollis Petrovitz 1961
Ataenius fattigi Cartwright 1948
Ataenius floreanae Cook & Peck 2000
Ataenius gammonensis Stebnicka & Howden 1997
Ataenius garamas Peyerimhoff 1929
Ataenius gilesi Stebnicka & Howden 1997
Ataenius glaseri Cartwright 1974
Ataenius gruneri Chalumeau 1979
Ataenius guayasi Stebnicka 2001
Ataenius guriensis Stebnicka 2005
Ataenius heinekeni (Wollaston 1854)
Ataenius hesperius Cartwright 1974
Ataenius horticola Harold 1869
Ataenius howdeni Chalumeau 1978
Ataenius indutus Petrovitz 1961
Ataenius inquisitus Horn 1887
Ataenius insulae Chalumeau & Gruner 1974
Ataenius integricollis Lea 1923
Ataenius jamaicensis Chapin 1940
Ataenius jardinensis Stebnicka 2002
Ataenius kapalgaensis Stebnicka & Howden 1997
Ataenius koebelei Blackburn 1904
Ataenius koelleri Balthasar 1963
Ataenius lobatus Horn 1871
Ataenius luctuosus (Burmeister 1877)
Ataenius luteomargo Chapin 1940
Ataenius martinezi Petrovitz 1973
Ataenius miamii Cartwright 1934
Ataenius millstreamae Stebnicka & Howden 1997
Ataenius monticola Paulian 1940
Ataenius montreuili Bordat & Théry 2012
Ataenius nakpandurii Endrödi 1973
Ataenius nigricans Paulian 1933
Ataenius noronhai Stebnicka 2007
Ataenius nugator Harold 1880
Ataenius oaxacaensis Stebnicka & Lago 2005
Ataenius ocumarensis Stebnicka 2001
Ataenius onkonensis Stebnicka 2005
Ataenius ovatulus Horn 1871
Ataenius palmaritoensis Stebnicka 2007
Ataenius palustris (Montrouzier 1860)
Ataenius panamensis Hinton 1936
Ataenius peregrinator Harold 1877
Ataenius pereirai Petrovitz 1970
Ataenius picinus Harold 1867
Ataenius plaumanni Petrovitz 1973
Ataenius pseudoclavatus Stebnicka 2005
Ataenius pseudostercorator Stebnicka 2003
Ataenius punctatohirsutus Schmidt 1909
Ataenius raccurti Chalumeau 1978
Ataenius raslani Bordat & Théry 2012
Ataenius rubrotessellatus (Blanchard 1846)
Ataenius saltae Stebnicka 2007
Ataenius scabrellus Schmidt 1909
Ataenius scutellaris Harold 1867
Ataenius siminasus Petrovitz 1973
Ataenius skelleyi Stebnicka 2007
Ataenius sparsicollis Blackburn 1904
Ataenius spretulus (Haldeman 1848)
Ataenius stebnickae Bordat & Théry 2012
Ataenius steinheili Harold 1874
Ataenius stephani Cartwright 1974
Ataenius stercorator (Fabricius 1775)
Ataenius strigatus (Say 1823)
Ataenius strigifrons Schmidt 1920
Ataenius suineata Deloya 2012
Ataenius synnotensis Stebnicka & Howden 1997
Ataenius tarumensis Stebnicka 2007
Ataenius texanus Harold 1874
Ataenius tindalensis Stebnicka & Howden 1997
Ataenius tomentosus Petrovitz 1969
Ataenius usingeri Hinton 1936
Ataenius warisensis Stebnicka 1998
Ataenius windjanae Stebnicka & Howden 1997
182 additional siblings truncated for brevity.
See the resource file for a full list.

GBIF classification

Animalia
Arthropoda
Insecta
Coleoptera
Aphodiidae
Ataenius Harold 1867
Ataenius gracilis (Melsheimer 1845) (this page)
Ataenius abditus (Haldeman 1848)
Ataenius aciculus Hinton 1936
Ataenius acutulus (Schmidt 1913)
Ataenius aequalis Harold 1880
Ataenius aequatorialis Petrovitz 1961
Ataenius attenuator Harold 1874
Ataenius australasiae (Boheman 1858)
Ataenius biroi (Endrödi 1951)
Ataenius bispinulosus Schmidt 1911
Ataenius blapoides Balthasar 1947
Ataenius bolivari Stebnicka 2001
Ataenius borgmeieri Hinton 1936
Ataenius brevicollis (Wollaston 1854)
Ataenius brevis Fall 1930
Ataenius brevitarsis Petrovitz 1964
Ataenius buenavistae Stebnicka 2001
Ataenius caesoides (Fairmaire 1900)
Ataenius canoasus Stebnicka 2007
Ataenius catarinaensis Stebnicka 2007
Ataenius confertus Fall 1909
Ataenius coriarius Schmidt 1912
Ataenius corporaali Boucomont 1924
Ataenius corrosus Chapin 1940
Ataenius costulifer Balthasar 1941
Ataenius crenaticollis Petrovitz 1973
Ataenius crenatipennis (Mac Leay 1871)
Ataenius crenator Harold 1876
Ataenius cucutae Stebnicka 2007
Ataenius damzeni Bukejs & Alekseev 2018
Ataenius eringundae Stebnicka & Howden 1997
Ataenius fattigi Cartwright 1948
Ataenius forsteri Balthasar 1960
Ataenius freyi Petrovitz 1961
Ataenius gammonensis Stebnicka & Howden 1997
Ataenius garamas Peyerimhoff 1929
Ataenius gascoyneensis Stebnicka & Howden 1997
Ataenius gibbus Blackburn 1904
Ataenius glabriventris Schmidt 1911
Ataenius guayasi Stebnicka 2001
Ataenius guriensis Stebnicka 2005
Ataenius icanus Balthasar 1941
Ataenius illaetabilis Lea 1923
Ataenius imbricatoides Schmidt 1909
Ataenius imbricatus (Melsheimer 1845)
Ataenius imparilis Blackburn 1904
Ataenius impiger Schmidt 1916
Ataenius impressus (Petrovitz 1963)
Ataenius indutus Petrovitz 1961
Ataenius lenkoi Petrovitz 1973
Ataenius lobatus Horn 1871
Ataenius longiclavus Petrovitz 1970
Ataenius luctuosus (Burmeister 1877)
Ataenius luteomargo Chapin 1940
Ataenius madagassicus Bordat 1990
Ataenius maghribinicus Baraud 1985
Ataenius martinezi Petrovitz 1973
Ataenius miamii Cartwright 1934
Ataenius millstreamae Stebnicka & Howden 1997
Ataenius murchisoni Stebnicka & Howden 1997
Ataenius noques Stebnicka 2007
Ataenius ocumarensis Stebnicka 2001
Ataenius onkonensis Stebnicka 2005
Ataenius ovatulus Horn 1871
Ataenius pacificus (Sharp 1879)
Ataenius palmerstoni Blackburn 1891
Ataenius panamensis Hinton 1936
Ataenius parallelipennis Petrovitz 1973
Ataenius parvus Lea 1923
Ataenius pearlensis Stebnicka 2006
Ataenius peregrinator Harold 1877
Ataenius pereirai Petrovitz 1970
Ataenius pseudimparilis Stebnicka & Howden 1997
Ataenius pseudoclavatus Stebnicka 2005
Ataenius quintanaroo Stebnicka 2006
Ataenius rosinae Balthasar 1967
Ataenius rubrotessellatus (Blanchard 1846)
Ataenius santarosae Stebnicka 2007
Ataenius saulensis Stebnicka 2006
Ataenius scabrelloides Petrovitz 1962
Ataenius scabrellus Schmidt 1909
Ataenius scalptifrons Bates 1887
Ataenius sculptilis Harold 1868
Ataenius sculptor Harold 1868
Ataenius scutellaris Harold 1867
Ataenius spec Martinez 1953
Ataenius stercorator (Fabricius 1775)
Ataenius striatocrenatus (Fairmaire 1889)
Ataenius strigatus (Say 1823)
Ataenius strigicaudus Bates 1887
Ataenius strigifrons Schmidt 1920
Ataenius synnotensis Stebnicka & Howden 1997
Ataenius talpoides Stebnicka 2001
Ataenius tambopatae Stebnicka 2001
Ataenius temperei Chalumeau & Gruner 1974
Ataenius texanus Harold 1874
Ataenius tomentosus Petrovitz 1969
Ataenius uriarrae Stebnicka & Howden 1997
Ataenius versicolor Schmidt 1916
Ataenius vinacoensis Stebnicka 2006
Ataenius yungasus Stebnicka 2001
192 additional siblings truncated for brevity.
See the resource file for a full list.

Integrated Taxonomic Information System (ITIS) Taxonomic Hierarchy

Animalia
Bilateria
Protostomia
Ecdysozoa
Arthropoda
Hexapoda
Insecta
Pterygota
Neoptera
Holometabola
Coleoptera Linnaeus 1758
Polyphaga Emery 1886
Scarabeiformia Crowson 1960
Scarabaeoidea Latreille 1802
Scarabaeidae Latreille 1802
Aphodiinae Leach 1815
Eupariini Schmidt 1910
Ataenius Harold 1867
Ataenius gracilis (Melsheimer 1845) (this page)
Ataenius abditus (Haldeman 1848)
Ataenius aequalis Harold 1880
Ataenius alternatus (Melsheimer 1845)
Ataenius apicalis Hinton 1937
Ataenius barberi Cartwright 1974
Ataenius brevicollis (Wollaston 1854)
Ataenius brevis Fall 1930
Ataenius californicus Horn 1887
Ataenius carinator Harold 1874
Ataenius castaniellus Bates 1887
Ataenius cognatus (Le Conte 1858)
Ataenius confertus Fall 1909
Ataenius crenulatus Schmidt 1910
Ataenius cribrithorax Bates 1887
Ataenius cylindrus Horn 1871
Ataenius desertus Horn 1871
Ataenius ecruensis Stebnicka & Lago 2005
Ataenius erratus Fall 1930
Ataenius exiguus Brown 1932
Ataenius fattigi Cartwright 1948
Ataenius glaseri Cartwright 1974
Ataenius heinekeni (Wollaston 1854)
Ataenius hesperius Cartwright 1974
Ataenius hirsutus Horn 1871
Ataenius imbricatus (Melsheimer 1845)
Ataenius inquisitus Horn 1887
Ataenius insculptus Horn 1887
Ataenius languidus Schmidt 1911
Ataenius liogaster Bates 1887
Ataenius lobatus Horn 1871
Ataenius miamii Cartwright 1934
Ataenius oklahomensis Brown 1930
Ataenius opatrinus Harold 1867
Ataenius ovatulus Horn 1871
Ataenius pacificus (Sharp 1879)
Ataenius peregrinator Harold 1877
Ataenius picinus Harold 1867
Ataenius platensis (Blanchard 1847)
Ataenius puncticollis Le Conte 1868
Ataenius robustus Horn 1871
Ataenius scabrelloides Petrovitz 1962
Ataenius scabrellus Schmidt 1909
Ataenius sculptor Harold 1868
Ataenius setiger Bates 1887
Ataenius spretulus (Haldeman 1848)
Ataenius stephani Cartwright 1974
Ataenius stercorator (Fabricius 1775)
Ataenius strigatus (Say 1823)
Ataenius texanus Harold 1874
Ataenius utahensis Cartwright 1974
Ataenius vandykei Cartwright 1974
Ataenius waltherhorni Balthasar 1938
Ataenius wenzelii Horn 1887

wikidata_hierarchy tar gz

Biota
Eukaryota
Unikonta
Opisthokonta
Holozoa
Filozoa
Apoikozoa
Animalia
Eumetazoa
Bilateria
Protostomia
Ecdysozoa
Arthropoda
Hexapoda
Insecta
Dicondylia
Pterygota
Metapterygota
Neoptera
Eumetabola
Endopterygota
Coleoptera
Polyphaga
Scarabaeiformia
Scarabaeoidea
Scarabaeidae
Ataenius
Ataenius gracilis (this page)
Ataenius abancay
Ataenius abditus
Ataenius aciculus
Ataenius acutulus
Ataenius aequalis
Ataenius aequatorialis
Ataenius africanus
Ataenius alternatus
Ataenius angulatus
Ataenius apicalis
Ataenius arenosus
Ataenius argillaceus
Ataenius arrowi
Ataenius attenuator
Ataenius bolivarensis
Ataenius borjae
Ataenius caicarae
Ataenius canoasus
Ataenius cartago
Ataenius catarinaensis
Ataenius cavagnaroi
Ataenius ceylonensis
Ataenius chapini
Ataenius chinacotae
Ataenius circulusbrumalis
Ataenius clavatus
Ataenius clitellarius
Ataenius cochabambae
Ataenius cognatus
Ataenius corrosus
Ataenius costulifer
Ataenius cristobalensis
Ataenius cucutae
Ataenius ecruensis
Ataenius elisaensis
Ataenius exiguus
Ataenius floreanae
Ataenius forsteri
Ataenius freyi
Ataenius garamas
Ataenius gibbus
Ataenius glabriventris
Ataenius glaseri
Ataenius granocostatus
Ataenius gruneri
Ataenius guanacastae
Ataenius guriensis
Ataenius heinekeni
Ataenius hesperius
Ataenius hirsutus
Ataenius hispaniolae
Ataenius hispidus
Ataenius holopubescens
Ataenius horticola
Ataenius howdeni
Ataenius hrubanti
Ataenius huanus
Ataenius hygrophilus
Ataenius icanus
Ataenius illaetabilis
Ataenius imbricatoides
Ataenius imbricatus
Ataenius imparilis
Ataenius impressus
Ataenius integer
Ataenius intermedius
Ataenius iquitosae
Ataenius isabelae
Ataenius jardinensis
Ataenius lamarensis
Ataenius londrinae
Ataenius luctuosus
Ataenius luteomargo
Ataenius madagassicus
Ataenius maghribinicus
Ataenius martinezi
Ataenius miamii
Ataenius napoensis
Ataenius noques
Ataenius noronhai
Ataenius oaxacaensis
Ataenius onkonensis
Ataenius palmaritoensis
Ataenius pearlensis
Ataenius plaumanni
Ataenius polyglyptus
Ataenius pseudoclavatus
Ataenius pseudostercorator
Ataenius pseudousingeri
Ataenius quintanaroo
Ataenius saltae
Ataenius santarosae
Ataenius saulensis
Ataenius schmidti
Ataenius skelleyi
Ataenius talpoides
Ataenius tambopatae
Ataenius tarumensis
Ataenius vinacoensis
Ataenius walterhorni
208 additional siblings truncated for brevity.
See the resource file for a full list.
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