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Creatures » … » Animal » … » Vertebrates » … » Amphibians » Frogs And Toads » … » Shield Toads » …

Shield Toad

Brachycephalus pernix Pombal, Wistuba & Bornschein 1998

Scientific Names

Preferred Names

Brachycephalus pernix Pombal, Wistuba & Bornschein 1998
Species recognized by EOL Dynamic Hierarchy April 2022
Reference taxon from Integrated Taxonomic Information System (ITIS)
Brachycephalus pernix
Species recognized by wikipedia VI, wikipedia CA, wikipedia EU, NCBI, and Wildfinder (WWF)
Brachycephalus pernix
Recognized by Arkive
Brachycephalus pernix Pombal, Wistuba & Bornschein 1998
Species recognized by EOL Dynamic Hierarchy 1.1, EOL Dynamic Hierarchy 1.1, Frost Darrel R 2018 Amphibian Species of the World, and GBIF classification
Brachycephalus pernix Pombal, Wistuba & Bornschein 1998
Recognized by IABIN
Brachycephalus pernix
Species recognized by wikidata_hierarchy tar gz, wikipedia ES, wikipedia PT, wikipedia FR, wikipedia EN, iNaturalist data coverage, and Wikipedia (inferred records)
Brachycephalus pernix
Recognized by Animal Diversity Web Traits and images, AmphibiaWeb articles, and BHL data coverage
Brachycephalus pernix Pombal, Wistuba & Bornschein 1998
Species recognized by IUCN, Catalogue of Life 26 March 2024, Integrated Taxonomic Information System (ITIS) Taxonomic Hierarchy, GBIF data coverage, Dynamic Hierarchy Version 2.2.3, and EOL Dynamic Hierarchy Version 2.2.4

Common Names

English

Shield toad preferred
Recognized by Arkive

Curated hierarchies for Brachycephalus pernix Pombal, Wistuba & Bornschein 1998

EOL Dynamic Hierarchy April 2022

Life
Cellular Organisms
Eukaryota
Opisthokonta
Metazoa
Bilateria
Deuterostomia
Chordata
Vertebrata
Gnathostomata
Osteichthyes
Sarcopterygii
Tetrapoda
Lissamphibia
Anura
Brachycephaloidea Günther 1858
Brachycephalidae Günther 1858
Brachycephalus Fitzinger 1826
Brachycephalus pernix Pombal, Wistuba & Bornschein 1998 (this page)
Brachycephalus actaeus Monteiro, Condez, Garcia, Comitti, Amaral & Haddad 2018
Brachycephalus albolineatus Bornschein, Ribeiro, Blackburn, Stanley & Pie 2016
Brachycephalus alipioi Pombal & Gasparini 2006
Brachycephalus atelopoide Miranda-Ribeiro 1920
Brachycephalus auroguttatus Ribeiro, Firkowski, Bornschein & Pie 2015
Brachycephalus boticario Pie, Bornschein, Firkowski, Belmonte-Lopes & Ribeiro 2015
Brachycephalus brunneus Ribeiro, Alves, Haddad & Reis 2005
Brachycephalus bufonoides Miranda-Ribeiro 1920
Brachycephalus coloratus Ribeiro, Blackburn, Stanley, Pie & Bornschein 2017
Brachycephalus crispus Condez, Clemente-Carvalho, Haddad & Reis 2014
Brachycephalus curupira Ribeiro, Blackburn, Stanley, Pie & Bornschein 2017
Brachycephalus darkside Guimarães, Luz, Rocha & Feio 2017
Brachycephalus didactylus (Izecksohn 1971)
Brachycephalus ephippium (Spix 1824)
Brachycephalus ferruginus Alves, Ribeiro, Haddad & Reis 2006
Brachycephalus fuscolineatus Pie, Bornschein, Firkowski, Belmonte-Lopes & Ribeiro 2015
Brachycephalus garbeanus Miranda-Ribeiro 1920
Brachycephalus guarani Clemente-Carvalho, Giaretta, Condez, Haddad & Reis 2012
Brachycephalus hermogenesi (Giaretta & Sawaya 1998)
Brachycephalus izecksohni Ribeiro, Alves, Haddad & Reis 2005
Brachycephalus leopardus Ribeiro, Firkowski & Pie 2015
Brachycephalus margaritatus Pombal & Izecksohn 2011
Brachycephalus mariaeterezae Bornschein, Morato, Firkowski, Ribeiro & Pie 2015
Brachycephalus mirissimus Pie, Ribeiro, Confetti, Nadaline & Bornschein 2018
Brachycephalus nodoterga Miranda-Ribeiro 1920
Brachycephalus olivaceus Bornschein, Morato, Firkowski, Ribeiro & Pie 2015
Brachycephalus pitanga Alves, Sawaya, Reis & Haddad 2009
Brachycephalus pombali Alves, Ribeiro, Haddad & Reis 2006
Brachycephalus pulex Napoli, Caramaschi, Cruz & Dias 2011
Brachycephalus quiririensis Pie & Ribeiro 2015
Brachycephalus sulfuratus Condez, Monteiro, Comitti, Garcia, Amaral & Haddad 2016
Brachycephalus toby Haddad, Alves, Clemente-Carvalho & Reis 2010
Brachycephalus tridactylus Garey, Lima, Hartmann & Haddad 2012
Brachycephalus verrucosus Ribeiro, Firkowski, Bornschein & Pie 2015
Brachycephalus vertebralis Pombal 2001

Catalogue of Life 26 March 2024

Biota
Animalia
Chordata
Vertebrata
Gnathostomata
Osteichthyes
Sarcopterygii
Tetrapoda
Amphibia
Anura
Brachycephaloidea Günther 1858
Brachycephalidae Günther 1858
Brachycephalus Fitzinger 1826
Brachycephalus pernix Pombal, Wistuba & Bornschein 1998 (this page)
Brachycephalus actaeus Monteiro, Condez, Garcia, Comitti, Amaral & Haddad 2018
Brachycephalus albolineatus Bornschein, Ribeiro, Blackburn, Stanley & Pie 2016
Brachycephalus alipioi Pombal & Gasparini 2006
Brachycephalus atelopoide Miranda-Ribeiro 1920
Brachycephalus auroguttatus Ribeiro, Firkowski, Bornschein & Pie 2015
Brachycephalus boticario Pie, Bornschein, Firkowski, Belmonte-Lopes & Ribeiro 2015
Brachycephalus brunneus Ribeiro, Alves, Haddad & Reis 2005
Brachycephalus bufonoides Miranda-Ribeiro 1920
Brachycephalus coloratus Ribeiro, Blackburn, Stanley, Pie & Bornschein 2017
Brachycephalus crispus Condez, Clemente-Carvalho, Haddad & Reis 2014
Brachycephalus curupira Ribeiro, Blackburn, Stanley, Pie & Bornschein 2017
Brachycephalus darkside Guimarães, Luz, Rocha & Feio 2017
Brachycephalus didactylus (Izecksohn 1971)
Brachycephalus ephippium (Spix 1824)
Brachycephalus ferruginus Alves, Ribeiro, Haddad & Reis 2006
Brachycephalus fuscolineatus Pie, Bornschein, Firkowski, Belmonte-Lopes & Ribeiro 2015
Brachycephalus garbeanus Miranda-Ribeiro 1920
Brachycephalus guarani Clemente-Carvalho, Giaretta, Condez, Haddad & Reis 2012
Brachycephalus hermogenesi (Giaretta & Sawaya 1998)
Brachycephalus izecksohni Ribeiro, Alves, Haddad & Reis 2005
Brachycephalus leopardus Ribeiro, Firkowski & Pie 2015
Brachycephalus margaritatus Pombal & Izecksohn 2011
Brachycephalus mariaeterezae Bornschein, Morato, Firkowski, Ribeiro & Pie 2015
Brachycephalus mirissimus Pie, Ribeiro, Confetti, Nadaline & Bornschein 2018
Brachycephalus nodoterga Miranda-Ribeiro 1920
Brachycephalus olivaceus Bornschein, Morato, Firkowski, Ribeiro & Pie 2015
Brachycephalus pitanga Alves, Sawaya, Reis & Haddad 2009
Brachycephalus pombali Alves, Ribeiro, Haddad & Reis 2006
Brachycephalus pulex Napoli, Caramaschi, Cruz & Dias 2011
Brachycephalus quiririensis Pie & Ribeiro 2015
Brachycephalus sulfuratus Condez, Monteiro, Comitti, Garcia, Amaral & Haddad 2016
Brachycephalus toby Haddad, Alves, Clemente-Carvalho & Reis 2010
Brachycephalus tridactylus Garey, Lima, Hartmann & Haddad 2012
Brachycephalus verrucosus Ribeiro, Firkowski, Bornschein & Pie 2015
Brachycephalus vertebralis Pombal 2001

Dynamic Hierarchy Version 2.2.3

Brachycephalus pernix Pombal, Wistuba & Bornschein 1998 (this page)
Brachycephalus actaeus Monteiro, Condez, Garcia, Comitti, Amaral & Haddad 2018
Brachycephalus albolineatus Bornschein, Ribeiro, Blackburn, Stanley & Pie 2016
Brachycephalus alipioi Pombal & Gasparini 2006
Brachycephalus atelopoide Miranda-Ribeiro 1920
Brachycephalus auroguttatus Ribeiro, Firkowski, Bornschein & Pie 2015
Brachycephalus boticario Pie, Bornschein, Firkowski, Belmonte-Lopes & Ribeiro 2015
Brachycephalus brunneus Ribeiro, Alves, Haddad & Reis 2005
Brachycephalus bufonoides Miranda-Ribeiro 1920
Brachycephalus coloratus Ribeiro, Blackburn, Stanley, Pie & Bornschein 2017
Brachycephalus crispus Condez, Clemente-Carvalho, Haddad & Reis 2014
Brachycephalus curupira Ribeiro, Blackburn, Stanley, Pie & Bornschein 2017
Brachycephalus darkside Guimarães, Luz, Rocha & Feio 2017
Brachycephalus didactylus (Izecksohn 1971)
Brachycephalus ephippium (Spix 1824)
Brachycephalus ferruginus Alves, Ribeiro, Haddad & Reis 2006
Brachycephalus fuscolineatus Pie, Bornschein, Firkowski, Belmonte-Lopes & Ribeiro 2015
Brachycephalus garbeanus Miranda-Ribeiro 1920
Brachycephalus guarani Clemente-Carvalho, Giaretta, Condez, Haddad & Reis 2012
Brachycephalus hermogenesi (Giaretta & Sawaya 1998)
Brachycephalus izecksohni Ribeiro, Alves, Haddad & Reis 2005
Brachycephalus leopardus Ribeiro, Firkowski & Pie 2015
Brachycephalus margaritatus Pombal & Izecksohn 2011
Brachycephalus mariaeterezae Bornschein, Morato, Firkowski, Ribeiro & Pie 2015
Brachycephalus mirissimus Pie, Ribeiro, Confetti, Nadaline & Bornschein 2018
Brachycephalus nodoterga Miranda-Ribeiro 1920
Brachycephalus olivaceus Bornschein, Morato, Firkowski, Ribeiro & Pie 2015
Brachycephalus pitanga Alves, Sawaya, Reis & Haddad 2009
Brachycephalus pombali Alves, Ribeiro, Haddad & Reis 2006
Brachycephalus pulex Napoli, Caramaschi, Cruz & Dias 2011
Brachycephalus quiririensis Pie & Ribeiro 2015
Brachycephalus sulfuratus Condez, Monteiro, Comitti, Garcia, Amaral & Haddad 2016
Brachycephalus toby Haddad, Alves, Clemente-Carvalho & Reis 2010
Brachycephalus tridactylus Garey, Lima, Hartmann & Haddad 2012
Brachycephalus verrucosus Ribeiro, Firkowski, Bornschein & Pie 2015
Brachycephalus vertebralis Pombal 2001

EOL Dynamic Hierarchy 1.1

Life
Cellular Organisms
Eukaryota
Opisthokonta
Metazoa
Bilateria
Deuterostomia
Chordata
Vertebrata
Gnathostomata
Osteichthyes
Sarcopterygii
Tetrapoda
Lissamphibia
Anura
Brachycephaloidea Günther 1858
Brachycephalidae Günther 1858
Brachycephalus Fitzinger 1826
Brachycephalus pernix Pombal, Wistuba & Bornschein 1998 (this page)
Brachycephalus actaeus Monteiro, Condez, Garcia, Comitti, Amaral & Haddad 2018
Brachycephalus albolineatus Bornschein, Ribeiro, Blackburn, Stanley & Pie 2016
Brachycephalus alipioi Pombal & Gasparini 2006
Brachycephalus atelopoide Miranda-Ribeiro 1920
Brachycephalus auroguttatus Ribeiro, Firkowski, Bornschein & Pie 2015
Brachycephalus boticario Pie, Bornschein, Firkowski, Belmonte-Lopes & Ribeiro 2015
Brachycephalus brunneus Ribeiro, Alves, Haddad & Reis 2005
Brachycephalus bufonoides Miranda-Ribeiro 1920
Brachycephalus coloratus Ribeiro, Blackburn, Stanley, Pie & Bornschein 2017
Brachycephalus crispus Condez, Clemente-Carvalho, Haddad & Reis 2014
Brachycephalus curupira Ribeiro, Blackburn, Stanley, Pie & Bornschein 2017
Brachycephalus darkside Guimarães, Luz, Rocha & Feio 2017
Brachycephalus didactylus (Izecksohn 1971)
Brachycephalus ephippium (Spix 1824)
Brachycephalus ferruginus Alves, Ribeiro, Haddad & Reis 2006
Brachycephalus fuscolineatus Pie, Bornschein, Firkowski, Belmonte-Lopes & Ribeiro 2015
Brachycephalus garbeanus Miranda-Ribeiro 1920
Brachycephalus guarani Clemente-Carvalho, Giaretta, Condez, Haddad & Reis 2012
Brachycephalus hermogenesi (Giaretta & Sawaya 1998)
Brachycephalus izecksohni Ribeiro, Alves, Haddad & Reis 2005
Brachycephalus leopardus Ribeiro, Firkowski & Pie 2015
Brachycephalus margaritatus Pombal & Izecksohn 2011
Brachycephalus mariaeterezae Bornschein, Morato, Firkowski, Ribeiro & Pie 2015
Brachycephalus mirissimus Pie, Ribeiro, Confetti, Nadaline & Bornschein 2018
Brachycephalus nodoterga Miranda-Ribeiro 1920
Brachycephalus olivaceus Bornschein, Morato, Firkowski, Ribeiro & Pie 2015
Brachycephalus pitanga Alves, Sawaya, Reis & Haddad 2009
Brachycephalus pombali Alves, Ribeiro, Haddad & Reis 2006
Brachycephalus pulex Napoli, Caramaschi, Cruz & Dias 2011
Brachycephalus quiririensis Pie & Ribeiro 2015
Brachycephalus sulfuratus Condez, Monteiro, Comitti, Garcia, Amaral & Haddad 2016
Brachycephalus toby Haddad, Alves, Clemente-Carvalho & Reis 2010
Brachycephalus tridactylus Garey, Lima, Hartmann & Haddad 2012
Brachycephalus verrucosus Ribeiro, Firkowski, Bornschein & Pie 2015
Brachycephalus vertebralis Pombal 2001

EOL Dynamic Hierarchy Version 2.2.4

Brachycephalus pernix Pombal, Wistuba & Bornschein 1998 (this page)
Brachycephalus actaeus Monteiro, Condez, Garcia, Comitti, Amaral & Haddad 2018
Brachycephalus albolineatus Bornschein, Ribeiro, Blackburn, Stanley & Pie 2016
Brachycephalus alipioi Pombal & Gasparini 2006
Brachycephalus atelopoide Miranda-Ribeiro 1920
Brachycephalus auroguttatus Ribeiro, Firkowski, Bornschein & Pie 2015
Brachycephalus boticario Pie, Bornschein, Firkowski, Belmonte-Lopes & Ribeiro 2015
Brachycephalus brunneus Ribeiro, Alves, Haddad & Reis 2005
Brachycephalus bufonoides Miranda-Ribeiro 1920
Brachycephalus coloratus Ribeiro, Blackburn, Stanley, Pie & Bornschein 2017
Brachycephalus crispus Condez, Clemente-Carvalho, Haddad & Reis 2014
Brachycephalus curupira Ribeiro, Blackburn, Stanley, Pie & Bornschein 2017
Brachycephalus darkside Guimarães, Luz, Rocha & Feio 2017
Brachycephalus didactylus (Izecksohn 1971)
Brachycephalus ephippium (Spix 1824)
Brachycephalus ferruginus Alves, Ribeiro, Haddad & Reis 2006
Brachycephalus fuscolineatus Pie, Bornschein, Firkowski, Belmonte-Lopes & Ribeiro 2015
Brachycephalus garbeanus Miranda-Ribeiro 1920
Brachycephalus guarani Clemente-Carvalho, Giaretta, Condez, Haddad & Reis 2012
Brachycephalus hermogenesi (Giaretta & Sawaya 1998)
Brachycephalus izecksohni Ribeiro, Alves, Haddad & Reis 2005
Brachycephalus leopardus Ribeiro, Firkowski & Pie 2015
Brachycephalus margaritatus Pombal & Izecksohn 2011
Brachycephalus mariaeterezae Bornschein, Morato, Firkowski, Ribeiro & Pie 2015
Brachycephalus mirissimus Pie, Ribeiro, Confetti, Nadaline & Bornschein 2018
Brachycephalus nodoterga Miranda-Ribeiro 1920
Brachycephalus olivaceus Bornschein, Morato, Firkowski, Ribeiro & Pie 2015
Brachycephalus pitanga Alves, Sawaya, Reis & Haddad 2009
Brachycephalus pombali Alves, Ribeiro, Haddad & Reis 2006
Brachycephalus pulex Napoli, Caramaschi, Cruz & Dias 2011
Brachycephalus quiririensis Pie & Ribeiro 2015
Brachycephalus sulfuratus Condez, Monteiro, Comitti, Garcia, Amaral & Haddad 2016
Brachycephalus toby Haddad, Alves, Clemente-Carvalho & Reis 2010
Brachycephalus tridactylus Garey, Lima, Hartmann & Haddad 2012
Brachycephalus verrucosus Ribeiro, Firkowski, Bornschein & Pie 2015
Brachycephalus vertebralis Pombal 2001

Frost Darrel R 2018 Amphibian Species of the World

Amphibia
Anura
Brachycephaloidea Günther 1858
Brachycephalidae Günther, 1858
Brachycephalus Fitzinger 1826
Brachycephalus pernix Pombal, Wistuba, and Bornschein, 1998 (this page)
Brachycephalus actaeus Monteiro, Condez, Garcia, Comitti, Amaral, and Haddad, 2018
Brachycephalus albolineatus Bornschein, Ribeiro, Blackburn, Stanley, and Pie, 2016
Brachycephalus alipioi Pombal and Gasparini, 2006
Brachycephalus atelopoide Miranda-Ribeiro, 1920
Brachycephalus auroguttatus Ribeiro, Firkowski, Bornschein, and Pie, 2015
Brachycephalus boticario Pie, Bornschein, Firkowski, Belmonte-Lopes, and Ribeiro, 2015
Brachycephalus brunneus Ribeiro, Alves, Haddad, and Reis, 2005
Brachycephalus bufonoides Miranda-Ribeiro, 1920
Brachycephalus coloratus Ribeiro, Blackburn, Stanley, Pie, and Bornschein, 2017
Brachycephalus crispus Condez, Clemente-Carvalho, Haddad, and Reis, 2014
Brachycephalus curupira Ribeiro, Blackburn, Stanley, Pie, and Bornschein, 2017
Brachycephalus darkside Guimarães, Luz, Rocha, and Feio, 2017
Brachycephalus didactylus (Izecksohn, 1971)
Brachycephalus ephippium (Spix, 1824)
Brachycephalus ferruginus Alves, Ribeiro, Haddad, and Reis, 2006
Brachycephalus fuscolineatus Pie, Bornschein, Firkowski, Belmonte-Lopes, and Ribeiro, 2015
Brachycephalus garbeanus Miranda-Ribeiro, 1920
Brachycephalus guarani Clemente-Carvalho, Giaretta, Condez, Haddad, and Reis, 2012
Brachycephalus hermogenesi (Giaretta and Sawaya, 1998)
Brachycephalus izecksohni Ribeiro, Alves, Haddad, and Reis, 2005
Brachycephalus leopardus Ribeiro, Firkowski, and Pie, 2015
Brachycephalus margaritatus Pombal and Izecksohn, 2011
Brachycephalus mariaeterezae Bornschein, Morato, Firkowski, Ribeiro and Pie, 2015
Brachycephalus mirissimus Pie, Ribeiro, Confetti, Nadaline, and Bornschein, 2018
Brachycephalus nodoterga Miranda-Ribeiro, 1920
Brachycephalus olivaceus Bornschein, Morato, Firkowski, Ribeiro and Pie, 2015
Brachycephalus pitanga Alves, Sawaya, Reis, and Haddad, 2009
Brachycephalus pombali Alves, Ribeiro, Haddad, and Reis, 2006
Brachycephalus pulex Napoli, Caramaschi, Cruz, and Dias, 2011
Brachycephalus quiririensis Pie and Ribeiro, 2015
Brachycephalus sulfuratus Condez, Monteiro, Comitti, Garcia, Amaral, and Haddad, 2016
Brachycephalus toby Haddad, Alves, Clemente-Carvalho, and Reis, 2010
Brachycephalus tridactylus Garey, Lima, Hartmann, and Haddad, 2012
Brachycephalus verrucosus Ribeiro, Firkowski, Bornschein, and Pie, 2015
Brachycephalus vertebralis Pombal, 2001

GBIF classification

Animalia
Chordata
Amphibia
Anura
Brachycephalidae
Brachycephalus Fitzinger
Brachycephalus pernix Pombal, Wistuba & Bornschein 1998 (this page)
Brachycephalus actaeus Monteiro, Condez, Garcia, Comitti, Amaral & Haddad 2018
Brachycephalus alipioi Pombal & Gasparini 2006
Brachycephalus brunneus Ribeiro, Alves, Haddad & Reis 2005
Brachycephalus darkside
Brachycephalus didactylus (Izecksohn 1971)
Brachycephalus ephippium (Spix 1824)
Brachycephalus ferruginus Alves, Ribeiro, Haddad & Reis 2006
Brachycephalus hermogenesi (Giaretta & Sawaya 1998)
Brachycephalus izecksohni Ribeiro, Alves, Haddad & Reis 2005
Brachycephalus nodoterga Miranda-Ribeiro 1920
Brachycephalus pombali Alves, Ribeiro, Haddad & Reis 2006
Brachycephalus pulex
Brachycephalus sulfuratus Condez, Monteiro, Comitti, Garcia, Amaral & Haddad 2016
Brachycephalus vertebralis Pombal 2001

Integrated Taxonomic Information System (ITIS) Taxonomic Hierarchy

Animalia
Bilateria
Deuterostomia
Chordata
Vertebrata
Gnathostomata
Tetrapoda
Amphibia
Anura
Brachycephaloidea Günther 1858
Brachycephalidae Günther 1858
Brachycephalus Fitzinger 1826
Brachycephalus pernix Pombal, Wistuba & Bornschein 1998 (this page)
Brachycephalus actaeus Monteiro, Condez, Garcia, Comitti, Amaral & Haddad 2018
Brachycephalus albolineatus Bornschein, Ribeiro, Blackburn, Stanley & Pie 2016
Brachycephalus alipioi Pombal & Gasparini 2006
Brachycephalus atelopoide Miranda-Ribeiro 1920
Brachycephalus auroguttatus Ribeiro, Firkowski, Bornschein & Pie 2015
Brachycephalus boticario Pie, Bornschein, Firkowski, Belmonte-Lopes & Ribeiro 2015
Brachycephalus brunneus Ribeiro, Alves, Haddad & Reis 2005
Brachycephalus bufonoides Miranda-Ribeiro 1920
Brachycephalus coloratus Ribeiro, Blackburn, Stanley, Pie & Bornschein 2017
Brachycephalus crispus Condez, Clemente-Carvalho, Haddad & Reis 2014
Brachycephalus curupira Ribeiro, Blackburn, Stanley, Pie & Bornschein 2017
Brachycephalus darkside Guimarães, Luz, Rocha & Feio 2017
Brachycephalus didactylus (Izecksohn 1971)
Brachycephalus ephippium (Spix 1824)
Brachycephalus ferruginus Alves, Ribeiro, Haddad & Reis 2006
Brachycephalus fuscolineatus Pie, Bornschein, Firkowski, Belmonte-Lopes & Ribeiro 2015
Brachycephalus garbeanus Miranda-Ribeiro 1920
Brachycephalus guarani Clemente-Carvalho, Giaretta, Condez, Haddad & Reis 2012
Brachycephalus hermogenesi (Giaretta & Sawaya 1998)
Brachycephalus izecksohni Ribeiro, Alves, Haddad & Reis 2005
Brachycephalus leopardus Ribeiro, Firkowski & Pie 2015
Brachycephalus margaritatus Pombal & Izecksohn 2011
Brachycephalus mariaeterezae Bornschein, Morato, Firkowski, Ribeiro & Pie 2015
Brachycephalus mirissimus Pie, Ribeiro, Confetti, Nadaline & Bornschein 2018
Brachycephalus nodoterga Miranda-Ribeiro 1920
Brachycephalus olivaceus Bornschein, Morato, Firkowski, Ribeiro & Pie 2015
Brachycephalus pitanga Alves, Sawaya, Reis & Haddad 2009
Brachycephalus pombali Alves, Ribeiro, Haddad & Reis 2006
Brachycephalus pulex Napoli, Caramaschi, Cruz & Dias 2011
Brachycephalus quiririensis Pie & Ribeiro 2015
Brachycephalus sulfuratus Condez, Monteiro, Comitti, Garcia, Amaral & Haddad 2016
Brachycephalus toby Haddad, Alves, Clemente-Carvalho & Reis 2010
Brachycephalus tridactylus Garey, Lima, Hartmann & Haddad 2012
Brachycephalus verrucosus Ribeiro, Firkowski, Bornschein & Pie 2015
Brachycephalus vertebralis Pombal 2001

IUCN

Animalia
Chordata
Amphibia
Anura
Brachycephalidae
Brachycephalus pernix Pombal, Wistuba & Bornschein 1998 (this page)
Brachycephalus alipioi Pombal & Gasparini 2006
Brachycephalus brunneus Ribeiro, Alves, Haddad & Dos Reis 2005
Brachycephalus didactylus (Izecksohn 1971)
Brachycephalus ephippium (Spix 1824)
Brachycephalus ferruginus Alves, Ribeiro, Haddad & dos Reis 2006
Brachycephalus hermogenesi (Giaretta & Sawaya 1998)
Brachycephalus izecksohni Ribeiro, Alves, Haddad & Dos Reis 2005
Brachycephalus nodoterga Miranda-Ribeiro 1920
Brachycephalus pombali Alves, Ribeiro, Haddad & dos Reis 2006
Brachycephalus vertebralis Pombal 2001
Ischnocnema bolbodactyla (A. Lutz 1925)
Ischnocnema epipeda (Heyer 1984)
Ischnocnema erythromera (Heyer 1984)
Ischnocnema gehrti (Miranda-Ribeiro 1926)
Ischnocnema gualteri (B. Lutz 1974)
Ischnocnema guentheri (Steindachner 1864)
Ischnocnema henselii (Peters 1870)
Ischnocnema hoehnei (B. Lutz 1959)
Ischnocnema holti (Cochran 1948)
Ischnocnema izecksohni (Caramaschi & Kisteumacher 1989)
Ischnocnema juipoca (Sazima & Cardoso 1978)
Ischnocnema lactea (Miranda-Ribeiro 1923)
Ischnocnema manezinho (Garcia 1996)
Ischnocnema nasuta (A. Lutz 1925)
Ischnocnema nigriventris (Lutz 1925)
Ischnocnema octavioi (Bokermann 1965)
Ischnocnema oea (Heyer 1984)
Ischnocnema paranaensis (Langone & Segalla 1996)
Ischnocnema parva (Girard 1853)
Ischnocnema penaxavantinho Giaretta, Toffoli & Oliveira 2007
Ischnocnema pusilla Bokermann 1967
Ischnocnema randorum (Heyer 1985)
Ischnocnema sambaqui (Castanho & Haddad 2000)
Ischnocnema spanios (Heyer 1985)
Ischnocnema venancioi (B. Lutz 1959)
Ischnocnema verrucosa Reinhardt & Lütken 1862

NCBI

root
cellular organisms
Eukaryota
Opisthokonta
Metazoa
Eumetazoa
Bilateria
Deuterostomia
Chordata
Craniata
Vertebrata
Gnathostomata
Teleostomi
Euteleostomi
Sarcopterygii
Dipnotetrapodomorpha
Tetrapoda
Amphibia
Batrachia
Anura
Neobatrachia
Hyloidea
Brachycephalidae
Brachycephalus
Brachycephalus pernix (this page)
Brachycephalus actaeus
Brachycephalus albolineatus
Brachycephalus alipioi
Brachycephalus auroguttatus
Brachycephalus boticario
Brachycephalus brunneus
Brachycephalus didactylus
Brachycephalus ephippium
Brachycephalus ferruginus
Brachycephalus fuscolineatus
Brachycephalus garbeanus
Brachycephalus hermogenesi
Brachycephalus izecksohni
Brachycephalus leopardus
Brachycephalus mariaeterezae
Brachycephalus nodoterga
Brachycephalus olivaceus
Brachycephalus pitanga
Brachycephalus pombali
Brachycephalus quiririensis
Brachycephalus sulfuratus
Brachycephalus toby
Brachycephalus tridactylus
Brachycephalus verrucosus
Brachycephalus vertebralis
unclassified Brachycephalus

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Biota
Eukaryota
Unikonta
Opisthokonta
Holozoa
Filozoa
Apoikozoa
Animalia
Eumetazoa
Bilateria
Deuterostomia
Chordata
Olfactores
Craniata
Vertebrata
Gnathostomata
Osteichthyes
Sarcopterygii
Dipnotetrapodomorpha
Tetrapodomorpha
Tetrapoda
Amphibia
Lissamphibia
Anura
Neobatrachia
Brachycephalidae
Brachycephalus
Brachycephalus pernix (this page)
Brachycephalus actaeus
Brachycephalus albolineatus
Brachycephalus alipioi
Brachycephalus atelopoide
Brachycephalus auroguttatus
Brachycephalus boticario
Brachycephalus brunneus
Brachycephalus bufonoides
Brachycephalus coloratus
Brachycephalus crispus
Brachycephalus curupira
Brachycephalus darkside
Brachycephalus didactylus
Brachycephalus ephippium
Brachycephalus ferruginus
Brachycephalus fuscolineatus
Brachycephalus garbeanus
Brachycephalus guarani
Brachycephalus hermogenesi
Brachycephalus izecksohni
Brachycephalus leopardus
Brachycephalus margaritatus
Brachycephalus mariaeterezae
Brachycephalus mirissimus
Brachycephalus nodoterga
Brachycephalus olivaceus
Brachycephalus pitanga
Brachycephalus pombali
Brachycephalus pulex
Brachycephalus quiririensis
Brachycephalus rotenbergae
Brachycephalus rotenbergae
Brachycephalus sulfuratus
Brachycephalus toby
Brachycephalus tridactylus
Brachycephalus verrucosus
Brachycephalus vertebralis
Bracycephalus tridactylus
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