Coriaria terminalis Hemsl.

Herbs subshrubby, 0.5-1 m tall; branches few; branchlets tetragonous or slightly narrowly winged, purplish glandular pubescent. Leaves sessile or shortly petiolate; leaf blade: lower ones broadly ovate or almost orbicular, 4-6(-7.5) × 3-5 cm, upper ones or those on lateral branches ovate-lanceolate or oblong-lanceolate, 2.5-4 × ca. 1.5 cm, papery, both surfaces glan- dular pubescent, basally (3-)5-9-veined, base cordate and semiamplexicaul or subrounded, margin glandular ciliate, apex acute or rounded, mucronate at tip. Raceme terminal, 12.5-21 cm; rachis purplish red, white glandular pilose. Flowers small, unisexual and plant monoecious; bracts purple, lanceolate, 3-4 mm. Pedicel 3-6 mm, elongated to 1.2 cm at fruiting, glandular pilose. Sepals broadly ovate, ovate, or ovate-lanceolate, 2.5-3 × 1.3-2.5 mm, outside grayish white puncticulate, margins translucent, apex acuminate or acute. Petals ovate, small, 1-1.5 × ca. 0.6 mm, fleshy, enlarged after anthesis. Filaments linear, ca. 0.8 mm; anthers oblong, 2-2.5 mm. Carpels 5, laterally compressed, ca. 1 mm; styles short; stigma ca. 2 mm. Fruit purplish red to black when mature, 2.5-3 mm in diam. Fl. May-Jun, fr. Jul-Sep. 2n = 40.

W Sichuan, S and SE Xizang, NW Yunnan [Bhutan, India, Nepal].

Thickets on mountain slopes; 1800-3700 m.

Coriaria terminalis var. xanthocarpa Rehder & E. H. Wilson.

Flowering from May to June; fruiting from July to September.

Coriaria terminalis is close relative of Coriaria nepalensis, but differs from the latter in its terminal (vs. axillary) inflorescences, subshrubby herbs (vs. shrubs).

Coriaria terminalis is occurring in W Sichuan, S and SE Xizang, NW Yunnan of China, Bhutan, India, Nepal.

The phylogenetic relationships of 12 Coriaria species collected from the representative disjunct areas were inferred by comparing 2416 bp of the combined data set of rbcL and matK genes. The phylogenetic tree shows that the Chile-Papua New Guinea-New Zealand-Pacific islands species and the Central America-northern South America species form a sister group, and the Eurasian clade is more basal to them (Yokoyama et al., 2000).

Herbs subshrubby, 0.5-1 m tall; branches few; branchlets tetragonous or slightly narrowly winged, purplish glandular pubescent. Leaves sessile or shortly petiolate; leaf blade: lower ones broadly ovate or almost orbicular, 4-6 cm long, 3-5 cm wide, upper ones or those on lateral branches ovate-lanceolate or oblong-lanceolate, 2.5-4 cm long, ca. 1.5 cm wide, papery, both surfaces glan- dular pubescent, basally 5-9-veined, base cordate and semiamplexicaul or subrounded, margin glandular ciliate, apex acute or rounded, mucronate at tip. Raceme terminal, 12.5-21 cm; rachis purplish red, white glandular pilose. Flowers small, unisexual and plant monoecious; bracts purple, lanceolate, 3-4 mm. Pedicel 3-6 mm, elongated to 1.2 cm at fruiting, glandular pilose. Sepals broadly ovate, ovate, or ovate-lanceolate, 2.5-3 mm long, 1.3-2.5 mm wide, outside grayish white puncticulate, margins translucent, apex acuminate or acute. Petals ovate, small, 1-1.5 mm long, ca. 0.6 mm wide, fleshy, enlarged after anthesis. Filaments linear, ca. 0.8 mm; anthers oblong, 2-2.5 mm. Carpels 5, laterally compressed, ca. 1 mm; styles short; stigma ca. 2 mm. Fruit purplish red to black when mature, 2.5-3 mm in diameter.

The chromosomal number of Coriaria terminalis is 2n = 40 (Oginuma, 1993).

Growing in thickets, on mountain slopes; 1800-3700 m.

Coriaria terminalis est une espèce de plantes dicotylédones de la famille des Coriariaceae, originaire de l'Himalaya (Chine, Bhoutan, Inde, Népal). C'est un arbrisseau ou sous-arbrisseau, pouvant atteindre 1 mètre de haut, aux feuilles caduques, opposées ou verticillées, qui produit des faux-fruits charnus contenant des akènes. Toutes les parties de la plante, y compris les graines, sont toxiques du fait de la présence de tutine, toxine qui attaque le système nerveux central.

Coriaria terminalis là một loài thực vật có hoa trong họ Coriariaceae. Loài này được Hemsl. mô tả khoa học đầu tiên năm 1892.

草马桑（学名：Coriaria terminalis）为马桑科马桑属的植物。分布在锡金、印度以及中国大陆的西藏、四川、云南等地，生长于海拔1,900米至3,700米的地区，多生于山坡林下或灌丛中，目前尚未由人工引种栽培。