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Cyclophoridae
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Cyclophorus volvulus (O. F. Müller 1774)
topla
genel bakış
veri
makaleler
isimler
Bilimsel İsimler
Tercih Edilen İsimler
Cyclophorus volvulus (O. F. Müller 1774)
Tür
EOL Dynamic Hierarchy April 2022
tarafından tanındı
EOL Mollusca Patch kaynağından referans vergi
Cyclophorus volvulus
Tür
Vietnam Species List
and
NCBI
tarafından tanındı
Cyclophorus volvulus
(O. F. Müller 1774)
Tür
EOL Dynamic Hierarchy 1.1
,
EOL Dynamic Hierarchy 1.1
, and
GBIF classification
tarafından tanındı
Cyclophorus volvulus
Tür
wikidata_hierarchy tar gz
and
wikipedia EN
tarafından tanındı
Cyclophorus volvulus
BHL data coverage
tarafından tanınıyor
Cyclophorus volvulus (Mueller 1774) Mueller 1774
Tür
TreatmentBank
tarafından tanındı
Cyclophorus volvulus (O. F. Müller 1774)
Tür
Catalogue of Life 26 March 2024
,
GBIF data coverage
,
Dynamic Hierarchy Version 2.2.3
, and
EOL Dynamic Hierarchy Version 2.2.4
tarafından tanındı
Alternatif İsimler
Helix volvulus
O. F. Müller 1774
EOL Dynamic Hierarchy 1.1
and
EOL Dynamic Hierarchy 1.1
göre Synonym
Cyclophorus (Cyclophorus) volvulus (O. F. Müller 1774)
Catalogue of Life 26 March 2024
göre Synonym
Helix volvulus O. F. Müller 1774
Catalogue of Life 26 March 2024
göre Synonym
Turbo volvulus (O. F. Müller 1774)
Catalogue of Life 26 March 2024
göre Synonym
Genel Adlar
Bu taksonla ilişkilendirilmiş ortak bir ad yoktur.
Cyclophorus volvulus (O. F. Müller 1774) için küratörlük hiyerarşileri
EOL Dynamic Hierarchy April 2022
Life
Cellular Organisms
Eukaryota
Opisthokonta
Metazoa
Bilateria
Protostomia
Spiralia
Mollusca
Gastropoda Cuvier 1795
Caenogastropoda Cox 1960
Architaenioglossa
Cyclophoridae
Cyclophorus Montfort 1810
Cyclophorus volvulus (O. F. Müller 1774)
(bu sayfa)
Cyclophorus affinis Theobald 1858
Cyclophorus altivagus Benson 1854
Cyclophorus amoenus (L. Pfeiffer 1854)
Cyclophorus aquilus (Sowerby 1843)
Cyclophorus aurantiacus Schumacher 1817
Cyclophorus bensoni (L. Pfeiffer 1854)
Cyclophorus cambodgensis Morlet 1884
Cyclophorus cantori (Benson 1851)
Cyclophorus donghoiensis Thach & F. Huber 2017
Cyclophorus dubius (Morelet 1881)
Cyclophorus franzhuberi Thach 2017
Cyclophorus fulguratus (L. Pfeiffer 1852)
Cyclophorus granum Morelet 1881
Cyclophorus hangmonensis Raheem & S. Schneider 2017
Cyclophorus haughtoni Theobald 1858
Cyclophorus herklotsi Martens 1860
Cyclophorus hirasei Pilsbry 1901
Cyclophorus horridulum (Morelet 1882)
Cyclophorus huberi Thach 2016
Cyclophorus indicus (Deshayes 1832)
Cyclophorus involvulus (O. F. Müller 1774)
Cyclophorus jerdoni (Benson 1851)
Cyclophorus jourdyi Morlet 1886
Cyclophorus khongensis Thach & F. Huber 2017
Cyclophorus kikaiensis Pilsbry 1902
Cyclophorus kubaryi Möllendorff 1895
Cyclophorus microscopicus Morelet 1881
Cyclophorus nagaensis Godwin-Austen & Beddome 1894
Cyclophorus ngheanensis Thach & F. Huber 2018
Cyclophorus nilagiricus (Benson 1852)
Cyclophorus pyrotrema Benson 1854
Cyclophorus raripilus Mousson 1881
Cyclophorus stenomphalus (L. Pfeiffer 1846)
Cyclophorus stevenabbasorum Thach 2016
Cyclophorus stungtrengensis Thach & F. Huber 2018
Cyclophorus thakhekensis Thach & F. Huber 2018
Cyclophorus turgidus (Pfeiffer 1851)
Cyclophorus validus (G. B. Sowerby I 1842)
Catalogue of Life 26 March 2024
Biota
Animalia
Mollusca
Gastropoda Cuvier 1795
Caenogastropoda L. R. Cox 1960
Architaenioglossa Haller 1892
Cyclophoroidea Gray 1847
Cyclophoridae Gray 1847
Cyclophorinae Gray 1847
Cyclophorini Gray 1847
Cyclophorus Montfort 1810
Cyclophorus volvulus (O. F. Müller 1774)
(bu sayfa)
Cyclophorus abditus Nantarat & Panha 2014
Cyclophorus aborensis Godwin-Austen 1915
Cyclophorus acutimarginatus (G. B. Sowerby I 1842)
Cyclophorus aetarum Möllendorff 1895
Cyclophorus affinis Theobald 1858
Cyclophorus alabastrinus (L. Pfeiffer 1854)
Cyclophorus alabatensis Kobelt 1886
Cyclophorus altivagus W. H. Benson 1854
Cyclophorus amoenus L. Pfeiffer 1852
Cyclophorus appendiculatus L. Pfeiffer 1854
Cyclophorus aquilus (G. B. Sowerby I 1843)
Cyclophorus arthriticus Theobald 1864
Cyclophorus ateribalteiformis D.-N. Chen & G.-Q. Zhang 1998
Cyclophorus atramentarium (G. B. Sowerby I 1843)
Cyclophorus aurantiacus (Schumacher 1817)
Cyclophorus austenianus Preston 1914
Cyclophorus balteatus W. H. Benson 1857
Cyclophorus bankanus E. von Martens 1867
Cyclophorus bapuensis Godwin-Austen 1915
Cyclophorus barandae Hidalgo 1887
Cyclophorus batanicus Quadras & Möllendorff 1894
Cyclophorus beddomeanus Preston 1914
Cyclophorus benguetensis Hidalgo 1888
Cyclophorus bensoni L. Pfeiffer 1852
Cyclophorus bustoi Hidalgo 1888
Cyclophorus cambodgensis Morlet 1885
Cyclophorus canaliferus (G. B. Sowerby I 1842)
Cyclophorus cantori (W. H. Benson 1851)
Cyclophorus caothaii Thach & F. Huber 2020
Cyclophorus ceratodes Möllendorff 1895
Cyclophorus ceylanicus L. Pfeiffer 1852
Cyclophorus charpentieri (Mousson 1849)
Cyclophorus cicatricosus Gredler 1894
Cyclophorus clouthianus Möllendorff 1882
Cyclophorus consociatus E. A. Smith 1893
Cyclophorus cornutus Kobelt 1902
Cyclophorus coronensis Möllendorff 1895
Cyclophorus courbeti Ancey 1888
Cyclophorus crassalabella Godwin-Austen 1888
Cyclophorus crocatus (Born 1778)
Cyclophorus cruentus E. von Martens 1865
Cyclophorus cryptomphalus W. H. Benson 1857
Cyclophorus cucphuongensis K. C. M. von Oheimb 2019
Cyclophorus cucullatus (Gould 1856)
Cyclophorus cybeus (W. H. Benson 1857)
Cyclophorus daraganicus Hidalgo 1888
Cyclophorus delavayanus Heude 1885
Cyclophorus denselineatus L. Pfeiffer 1852
Cyclophorus dilatatus Heude 1886
Cyclophorus diplochilus Möllendorff 1894
Cyclophorus dodrans Mabille 1887
Cyclophorus donghoiensis Thach & F. Huber 2017
Cyclophorus dubius (Morelet 1881)
Cyclophorus ectopoma Möllendorff 1896
Cyclophorus elegans Möllendorff 1881
Cyclophorus eudeli E. A. Smith 1893
Cyclophorus everetti E. A. Smith 1893
Cyclophorus exaltatus (L. Pfeiffer 1855)
Cyclophorus excellens L. Pfeiffer 1854
Cyclophorus expansus L. Pfeiffer 1852
Cyclophorus fargesianus Heude 1885
Cyclophorus fernandezi Hidalgo 1888
Cyclophorus ferruginosus Heude 1885
Cyclophorus flavilabris W. H. Benson 1860
Cyclophorus floridus (L. Pfeiffer 1855)
Cyclophorus formosaensis G. Nevill 1881
Cyclophorus franzhuberi Thach 2017
Cyclophorus friesianus Möllendorff 1883
Cyclophorus frinianus Heude 1885
Cyclophorus fruhstorferi Möllendorff 1901
Cyclophorus fulguratus (L. Pfeiffer 1854)
Cyclophorus fultoni Godwin-Austen & R. H. Beddome 1894
Cyclophorus fuscicolor Godwin-Austen 1876
Cyclophorus granum Morelet 1881
Cyclophorus hangmonensis Raheem & S. Schneider 2017
Cyclophorus haughtoni Theobald 1858
Cyclophorus hebereri B. Rensch 1933
Cyclophorus hejingi Thach & F. Huber 2020
Cyclophorus herklotsi E. von Martens 1861
Cyclophorus hirasei Pilsbry 1901
Cyclophorus horridulum (Morelet 1882)
Cyclophorus huberi Thach 2016
Cyclophorus indicus (Deshayes 1832)
Cyclophorus involvulus (O. F. Müller 1774)
Cyclophorus jerdoni (W. H. Benson 1851)
Cyclophorus jourdyi Morlet 1886
Cyclophorus khongensis Thach & F. Huber 2017
Cyclophorus kikaiensis Pilsbry 1902
Cyclophorus microscopicus Morelet 1881
Cyclophorus nagaensis Godwin-Austen & R. H. Beddome 1894
Cyclophorus ngheanensis Thach & F. Huber 2018
Cyclophorus nilagiricus (W. H. Benson 1852)
Cyclophorus pyrotrema W. H. Benson 1854
Cyclophorus raripilus Morelet 1881
Cyclophorus stenomphalus (L. Pfeiffer 1846)
Cyclophorus stevenabbasorum Thach 2016
Cyclophorus stungtrengensis Thach & F. Huber 2018
Cyclophorus thakhekensis Thach & F. Huber 2018
Cyclophorus turgidus L. Pfeiffer 1852
Cyclophorus validus (G. B. Sowerby I 1842)
102 ek kardeş kısalık için kesildi.
Tam liste için
kaynak dosyasına
bakın.
Dynamic Hierarchy Version 2.2.3
Cyclophorus volvulus (O. F. Müller 1774)
(bu sayfa)
Cyclophorus affinis Theobald 1858
Cyclophorus altivagus Benson 1854
Cyclophorus amoenus (L. Pfeiffer 1854)
Cyclophorus aquilus (Sowerby 1843)
Cyclophorus aurantiacus Schumacher 1817
Cyclophorus bensoni (L. Pfeiffer 1854)
Cyclophorus cambodgensis Morlet 1884
Cyclophorus cantori (Benson 1851)
Cyclophorus donghoiensis Thach & F. Huber 2017
Cyclophorus dubius (Morelet 1881)
Cyclophorus franzhuberi Thach 2017
Cyclophorus fulguratus (L. Pfeiffer 1852)
Cyclophorus granum Morelet 1881
Cyclophorus hangmonensis Raheem & S. Schneider 2017
Cyclophorus haughtoni Theobald 1858
Cyclophorus herklotsi Martens 1860
Cyclophorus hirasei Pilsbry 1901
Cyclophorus horridulum (Morelet 1882)
Cyclophorus huberi Thach 2016
Cyclophorus indicus (Deshayes 1832)
Cyclophorus involvulus (O. F. Müller 1774)
Cyclophorus jerdoni (Benson 1851)
Cyclophorus jourdyi Morlet 1886
Cyclophorus khongensis Thach & F. Huber 2017
Cyclophorus kikaiensis Pilsbry 1902
Cyclophorus kubaryi Möllendorff 1895
Cyclophorus microscopicus Morelet 1881
Cyclophorus nagaensis Godwin-Austen & Beddome 1894
Cyclophorus ngheanensis Thach & F. Huber 2018
Cyclophorus nilagiricus (Benson 1852)
Cyclophorus pyrotrema Benson 1854
Cyclophorus raripilus Mousson 1881
Cyclophorus stenomphalus (L. Pfeiffer 1846)
Cyclophorus stevenabbasorum Thach 2016
Cyclophorus stungtrengensis Thach & F. Huber 2018
Cyclophorus thakhekensis Thach & F. Huber 2018
Cyclophorus turgidus (Pfeiffer 1851)
Cyclophorus validus (G. B. Sowerby I 1842)
EOL Dynamic Hierarchy 1.1
Life
Cellular Organisms
Eukaryota
Opisthokonta
Metazoa
Bilateria
Protostomia
Spiralia
Mollusca
Gastropoda
Architaenioglossa
Cyclophoroidea
Cyclophoridae
Cyclophorus
Cyclophorus volvulus (O. F. Müller 1774)
(bu sayfa)
Cyclophorus altivagus Benson 1854
Cyclophorus donghoiensis Thach & F. Huber 2017
Cyclophorus franzhuberi Thach 2017
Cyclophorus fulguratus (L. Pfeiffer 1852)
Cyclophorus hangmonensis Raheem & S. Schneider 2017
Cyclophorus haughtoni Theobald 1858
Cyclophorus huberi Thach 2016
Cyclophorus indicus (Deshayes 1832)
Cyclophorus involvulus (O. F. Müller 1774)
Cyclophorus jerdoni (Benson 1851)
Cyclophorus khongensis Thach & F. Huber 2017
Cyclophorus nagaensis Godwin-Austen & Beddome 1894
Cyclophorus ngheanensis Thach & F. Huber 2018
Cyclophorus nilagiricus (Benson 1852)
Cyclophorus pyrotrema Benson 1854
Cyclophorus stenomphalus (L. Pfeiffer 1846)
Cyclophorus stevenabbasorum Thach 2016
Cyclophorus stungtrengensis Thach & F. Huber 2018
Cyclophorus thakhekensis Thach & F. Huber 2018
Cyclophorus turgidus (Pfeiffer 1851)
Cyclophorus validum (G. B. Sowerby I 1842)
EOL Dynamic Hierarchy Version 2.2.4
Cyclophorus volvulus (O. F. Müller 1774)
(bu sayfa)
Cyclophorus affinis Theobald 1858
Cyclophorus altivagus Benson 1854
Cyclophorus amoenus (L. Pfeiffer 1854)
Cyclophorus aquilus (Sowerby 1843)
Cyclophorus aurantiacus Schumacher 1817
Cyclophorus bensoni (L. Pfeiffer 1854)
Cyclophorus cambodgensis Morlet 1884
Cyclophorus cantori (Benson 1851)
Cyclophorus donghoiensis Thach & F. Huber 2017
Cyclophorus dubius (Morelet 1881)
Cyclophorus franzhuberi Thach 2017
Cyclophorus fulguratus (L. Pfeiffer 1852)
Cyclophorus granum Morelet 1881
Cyclophorus hangmonensis Raheem & S. Schneider 2017
Cyclophorus haughtoni Theobald 1858
Cyclophorus herklotsi Martens 1860
Cyclophorus hirasei Pilsbry 1901
Cyclophorus horridulum (Morelet 1882)
Cyclophorus huberi Thach 2016
Cyclophorus indicus (Deshayes 1832)
Cyclophorus involvulus (O. F. Müller 1774)
Cyclophorus jerdoni (Benson 1851)
Cyclophorus jourdyi Morlet 1886
Cyclophorus khongensis Thach & F. Huber 2017
Cyclophorus kikaiensis Pilsbry 1902
Cyclophorus kubaryi Möllendorff 1895
Cyclophorus microscopicus Morelet 1881
Cyclophorus nagaensis Godwin-Austen & Beddome 1894
Cyclophorus ngheanensis Thach & F. Huber 2018
Cyclophorus nilagiricus (Benson 1852)
Cyclophorus pyrotrema Benson 1854
Cyclophorus raripilus Mousson 1881
Cyclophorus stenomphalus (L. Pfeiffer 1846)
Cyclophorus stevenabbasorum Thach 2016
Cyclophorus stungtrengensis Thach & F. Huber 2018
Cyclophorus thakhekensis Thach & F. Huber 2018
Cyclophorus turgidus (Pfeiffer 1851)
Cyclophorus validus (G. B. Sowerby I 1842)
GBIF classification
Animalia
Mollusca
Gastropoda
Architaenioglossa
Cyclophoridae
Cyclophorus Monfort 1810
Cyclophorus volvulus (O. F. Müller 1774)
(bu sayfa)
Cyclophorus aetarum Moellendorff 1895
Cyclophorus appendiculatus Pfeiffer 1852
Cyclophorus aurantiacus Schumacher 1817
Cyclophorus aurora (Benson 1851)
Cyclophorus austenianus Preston 1914
Cyclophorus bellus von Martens 1873
Cyclophorus binoyae Sykes
Cyclophorus cambodgensis Morlet 1884
Cyclophorus canaliferus (Sowerby 1842)
Cyclophorus cantori (Benson 1851)
Cyclophorus ceylanicus
Cyclophorus charpentieri (Mousson 1849)
Cyclophorus clathratus Heude
Cyclophorus consociatus E. A. Smith 1893
Cyclophorus courbeti Ancey 1888
Cyclophorus cratera Benson
Cyclophorus crocatus (Born 1778)
Cyclophorus cruentatus von Martens 1865
Cyclophorus cybeus (Benson 1867)
Cyclophorus denselineatus (L. Pfeiffer 1852)
Cyclophorus dilatatus Heude 1886
Cyclophorus dodrans Mabille 1887
Cyclophorus dubius (Morelet 1881)
Cyclophorus egregius von Martens 1900
Cyclophorus elegans Moellendorff 1881
Cyclophorus eximius
Cyclophorus exul Benson 1854
Cyclophorus fargesianus Heude 1885
Cyclophorus ferrugineus Heude
Cyclophorus floridus (Pfeiffer 1855)
Cyclophorus formosaensis Nevill 1881
Cyclophorus formosensis Nevill 1882
Cyclophorus friesianus Möllendorff 1883
Cyclophorus granum Morelet 1881
Cyclophorus haematomma Pfeiffer 1863
Cyclophorus herberi Rensch
Cyclophorus horridulum (Morelet 1882)
Cyclophorus huberi Thach 2016
Cyclophorus implicatus Bavay & Dautzenberg 1908
Cyclophorus indicus (Deshayes 1832)
Cyclophorus jerdoni (Benson 1851)
Cyclophorus jourdyi Morlet 1886
Cyclophorus khasiensis Nevill 1850
Cyclophorus kikaiensis Pilsbry 1902
Cyclophorus koboensis Godwin-Austen 1915
Cyclophorus kubaryi Möllendorff 1895
Cyclophorus latus Kuroda 1941
Cyclophorus leai Tryon 1869
Cyclophorus leucostomus
Cyclophorus loloensis Heude
Cyclophorus lubanicus Clench & Archer 1931
Cyclophorus mansuyi Dautzenberg & Fischer 1908
Cyclophorus martensianus Möllendorff 1874
Cyclophorus massiei Morlet 1891
Cyclophorus microscopicus Morelet 1881
Cyclophorus miyakoensis Pilsbry & Hirase 1905
Cyclophorus moellendorffi Schmacker & Böttger 1891
Cyclophorus muspratti Godwin-Austen & Beddome 1894
Cyclophorus ngheanensis Thach & F. Huber 2018
Cyclophorus nicobaricus L. Pfeiffer 1865
Cyclophorus nilafiricus (Benson 1852)
Cyclophorus nivicola Godwin-Austen 1876
Cyclophorus oakesi Godwin-Austen 1915
Cyclophorus occultus Nantarat & Panha 2019
Cyclophorus orthostylus Möllendorff 1898
Cyclophorus othostylus Moellendorff 1898
Cyclophorus pallens Heude
Cyclophorus papilio L. Pfeiffer 1861
Cyclophorus pealianus Nevill 1877
Cyclophorus pearsoni (Benson 1851)
Cyclophorus phlegeton Godwin-Austen 1889
Cyclophorus polynema (L. Pfeiffer 1854)
Cyclophorus pterocyclus Moellendorff 1895
Cyclophorus punctatulus Heude
Cyclophorus pyrostoma Mlldf.
Cyclophorus quadrasi Hidalgo 1888
Cyclophorus rangunensis
Cyclophorus raripilus Mousson 1881
Cyclophorus rebleri Fulton
Cyclophorus semperi Kobelt 1886
Cyclophorus sericatus (Ancey 1888)
Cyclophorus sericinus Guadras & Moellendorff 1894
Cyclophorus sexfilaris Heude
Cyclophorus sidiensis Godwin-Austen 1915
Cyclophorus smithi
Cyclophorus sowerbyi Hidalgo 1888
Cyclophorus speciosus (Philippi 1847)
Cyclophorus stenomphalus (L. Pfeiffer 1846)
Cyclophorus stevenabbasorum Thach 2016
Cyclophorus stungtrengensis Thach & F. Huber 2018
Cyclophorus subcarinatus Moellendorff 1882
Cyclophorus szechwanensis Yen 1939
Cyclophorus thakhekensis Thach & F. Huber 2018
Cyclophorus theobaldianus Benson 1857
Cyclophorus tigrinus
Cyclophorus trouiensis Wattebled 1886
Cyclophorus tryblium Benson 1854
Cyclophorus turgidus (Pfeiffer 1851)
Cyclophorus vescus Sykes
Cyclostoma cereum L. Pfeiffer 1855
86 ek kardeş kısalık için kesildi.
Tam liste için
kaynak dosyasına
bakın.
NCBI
root
cellular organisms
Eukaryota
Opisthokonta
Metazoa
Eumetazoa
Bilateria
Protostomia
Spiralia
Lophotrochozoa
Mollusca
Gastropoda
Caenogastropoda
Architaenioglossa
Cyclophoroidea
Cyclophoridae
Cyclophorus
Cyclophorus volvulus
(bu sayfa)
Cyclophorus abditus
Cyclophorus affinis
Cyclophorus amoenus
Cyclophorus aurantiacus
Cyclophorus bensoni
Cyclophorus cantori
Cyclophorus consociatus
Cyclophorus courbeti
Cyclophorus cryptomphalus
Cyclophorus cucphuongensis
Cyclophorus diplochilus
Cyclophorus expansus
Cyclophorus fasciatus
Cyclophorus formosaensis
Cyclophorus friesianus
Cyclophorus fulguratus
Cyclophorus haughtoni
Cyclophorus herklotsi
Cyclophorus hirasei
Cyclophorus jourdyi
Cyclophorus labiosus
Cyclophorus latus
Cyclophorus malayanus
Cyclophorus martensianus
Cyclophorus moellendorffi
Cyclophorus paracucphuongensis
Cyclophorus perdix
Cyclophorus pfeifferi
Cyclophorus phongnhakebangensis
Cyclophorus pyrostoma
Cyclophorus rangunensis
Cyclophorus saturnus
Cyclophorus semisulcatus
Cyclophorus songmaensis
Cyclophorus speciosus
Cyclophorus subcarinatus
Cyclophorus subfloridus
Cyclophorus takumisaitoi
Cyclophorus turgidus
Cyclophorus zebrinus
unclassified Cyclophorus
wikidata_hierarchy tar gz
Biota
Eukaryota
Unikonta
Opisthokonta
Holozoa
Filozoa
Apoikozoa
Animalia
Eumetazoa
Bilateria
Protostomia
Lophotrochozoa
Mollusca
Gastropoda
Orthogastropoda
Caenogastropoda
Architaenioglossa
Cyclophoroidea
Cyclophoridae
Cyclophorus volvulus
(bu sayfa)
Acroptychia
Adelopoma
Alycaeinae
Alyceus
Anceyiella
Anosycolus
Aulopoma
Aulopomatinae
Awalycaeus
Boucardicus
Chamalychaeus
Chondrocyclus
Cipangocharax
Craspedopoma
Craspedopomatinae
Craspedotropis
Cyathopoma
Cyclophorinae
Cyclophorus
Cyclophorus bensoni
Cyclophorus cantori
Cyclophorus expansus
Cyclophorus malayanus
Cyclosurus
Diaspira
Dicharax
Dioryx
Ditropis
Ditropopsis
Dominamaria
Ettemona
Japonia
Lagocheillinae
Leptopoma
Madecataulus
Madgeaconcha
Micraulax
Millotorbis
Mychopoma
Naggsiaconcha
Nobuea
Otopoma
Owengriffithsius
Pholeoteras
Pterocyclus
Rhisostoma
Spirostomatinae
Theobaldius
Ventriculus