Pupilla is a genus of minute air-breathing land snails, terrestrial pulmonate gastropod mollusks or micromollusks in the subfamily Pupillinae of the family Pupillidae. [2]
Shells of Pupilla species are known from terrestrial Cenozoic strata dating back to the Oligocene until the Holocene period.[3]
Distribution
This genus occurs in Eurasia, northern Africa and North America.[4]
Species
Species within the genus Pupilla include:
- † Pupilla aeoli (Hilber, 1884)
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Pupilla afghanicum Solem, 1979
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Pupilla alabiella Schileyko, 1984
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Pupilla alaskensis Nekola & Coles, 2014
- † Pupilla almuhambetovae Prysjazhnjuk, 2017
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Pupilla alpicola (Charpentier, 1837)[4]
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Pupilla annandalei Pilsbry[5]
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Pupilla anzobica Izzatullaev, 1970
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Pupilla armeniaca (Issel, 1865)
- † Pupilla bargi Prysiazhniuk, 2017
- † Pupilla belokrysi Steklov, 1966
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Pupilla bigranata (Rossmässler 1839)[4] (taxon inquirendum)
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Pupilla bipatulata Akramowski, 1943
- † Pupilla bituberculata H.-J. Wang & S.-Y. Guo, 1991
- † Pupilla blainvilleana (Dupuy, 1850)
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Pupilla blandii E. S. Morse, 1865 - Rocky Mountain column
- † Pupilla bogdanovkaense Prysiazhniuk, 2017
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Pupilla brevicostis (Benson, 1849)
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Pupilla calacharicus (Boettger, 1886)
- † Pupilla capitani (de Morgan, 1920)
- † Pupilla crossei (Michaud, 1862)
- † Pupilla cupella O. Boettger, 1889
- † Pupilla diezi (Flach, 1890)
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Pupilla diopsis (Benson, 1863)
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Pupilla duplicata (Preston, 1911)
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Pupilla ficulnea (Tate, 1894)[6]
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Pupilla gallae Tzvetkov, 1940
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Pupilla goniodon Pilsbry, 1927
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Pupilla guadalupensis Pilsbry, 1927
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Pupilla gutta (Benson, 1864)
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Pupilla hebes (Ancey 1881) - crestless column
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Pupilla hokkaidoensis Nekola, Coles & S. Chiba, 2014
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Pupilla hudsoniana Nekola & Coles, 2014
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Pupilla iheringi Suter, 1900
- † Pupilla impressa (F. Sandberger, 1862)
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Pupilla inequidenta Schileyko & Almuhambetova, 1979
- † Pupilla inermis Russell, 1931
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Pupilla inops (Reinhardt, 1877)
- † Pupilla iratiana (Dupuy, 1850)
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Pupilla khunjerabica Pokryszko, Auffenberg, Hlaváč & Naggs, 2009[5]
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Pupilla kyrostriata Walther & Hausdorf, 2014
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Pupilla limata Schileyko, 1984
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Pupilla loessica Ložek, 1954[7]
- † Pupilla michaudi Wenz, 1919
- † Pupilla mlomnickii (Friedberg, 1905)
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Pupilla muscorum (Linnaeus, 1758)[4] - widespread column - type species
- † Pupilla mutabilis Steklov, 1966
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Pupilla obliquicosta Smith, 1892 - extinct[8]
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Pupilla paraturcmenica Pokryszko, Auffenberg, Hlaváč & Naggs, 2009[5]
- † Pupilla parvula (Deshayes, 1863)
- † Pupilla perlabiata Gottschick & Wenz, 1919
- † Pupilla poltavica O. Boettger, 1889
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Pupilla profundotuberculata H.-J. Wang & S.-Y. Guo, 1991
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Pupilla pupula[9]
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Pupilla quxuensis D.-N. Chen & G.-Q. Zhang, 2001
- † Pupilla retusa (F. Sandberger, 1858)
- † Pupilla roberti Prysiazhniuk, 2017
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Pupilla salemensis (W. T. Blanford & H. F. Blanford, 1861)
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Pupilla satparanica Pokryszko, Auffenberg, Hlaváč & Naggs, 2009[5]
- † Pupilla selecta (Thomä, 1845)
- † Pupilla selectiformis (Simionescu & Barbu, 1940)
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Pupilla seriola (Benson, 1861)
- † Pupilla shantungensis Yen, 1969
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Pupilla simplexa Y.-T. Li, 1985
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Pupilla sonorana (Sterki, 1899) - three-tooth column
- † Pupilla staszicii (Łomnicki, 1886)
- † Pupilla steinheimensis (K. Miller, 1900)
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Pupilla sterkiana (Pilsbry, 1890)
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Pupilla sterrii (Forster & Voith, 1840)[4]
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Pupilla striopolita Schileyko, 1984
- † Pupilla subquadridens (A. d'Orbigny, 1852)
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Pupilla syngenes (Pilsbry, 1890) - top-heavy column
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Pupilla tetrodus (Boettger, 1870)
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Pupilla triplicata (Studer, 1820)[4]
- † Pupilla triplicatoidea Steklov, 1966
- † Pupilla tschumakovi Prysjazhnjuk, 2017
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Pupilla turcmenica (O. Boettger)[5]
- † Pupilla zeisslerae Schlickum, 1975
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Pupilla ziaratana Pokryszko, Auffenberg, Hlaváč & Naggs, 2009[5]
Synonyms
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Pupilla barrackporensis Gude, 1914: synonym of Insulipupa malayana (Issel, 1874) (junior synonym)
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Pupilla cinghalensis Gude, 1914: synonym of Indopupa cinghalensis (Gude, 1914) (original combination)
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Pupilla cryptodon (Heude, 1880): synonym of Gibbulinopsis cryptodon (Heude, 1880) (unaccepted combination)
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Pupilla cupa (Jan, 1832): synonym of Pupilla muscorum (Linnaeus, 1758) (junior synonym)
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Pupilla draparnaudi Leach [in Turton], 1831: synonym of Lauria cylindracea (Da Costa, 1778)
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Pupilla floridana Dall, 1885: synonym of Gastrocopta pentodon (Say, 1822) (junior synonym)
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Pupilla fontana (Krauss, 1848): synonym of Gibbulinopsis fontana (F. Krauss, 1848) (superseded combination)
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Pupilla gracilis Izzatullaev, 1970: synonym of Gibbulinopsis gracilis (Izzatullaev, 1970) (original combination)
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Pupilla hudsonianum [sic]: synonym of Pupilla hudsoniana Nekola & Coles, 2014 (wrong gender agreement of specific epithet)
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Pupilla muscerda (Benson, 1853): synonym of Microstele muscerda (Benson, 1853) (unaccepted combination)
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Pupilla pratensis (Clessin, 1871):[4] synonym of Pupilla alpicola (Charpentier, 1837) (unaccepted> junior subjective synonym)
- † Pupilla quadrigranata (F. Sandberger, 1858): synonym of † Pupilla selecta (Thomä, 1845) (junior synonym)
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Pupilla reibischi Dall & Ochsner, 1928: synonym of Gastrocopta reibischi (Dall & Ochsner, 1928) (original name)
- Pupilla salmensis (W. T. Blanford & H. F. Blanford, 1861): synonym of Pupilla salemensis (W. T. Blanford & H. F. Blanford, 1861) (misspelling of original name.)
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Pupilla signata (Mousson, 1873):[5] synonym of Gibbulinopsis signata (Mousson, 1873)
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Pupilla stoneri Chamberlin & Jones, 1929: synonym of Gastrocopta pilsbryana (Sterki, 1890)
- † Pupilla submuscorum Gottschick & Wenz, 1919: synonym of † Gibbulinopsis submuscorum (Gottschick & Wenz, 1919)
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Pupilla wolfae Tzvetkov, 1940: synonym of Pupilla sterrii (Voith, 1840) (junior synonym)
References
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^ Fleming J. (1828). A history of British animals, exhibiting the descriptive characters and systematic arrangement of the genera and species of quadrupeds, birds, reptiles, fishes, Mollusca, and Radiata of the United Kingdom; including the indigenous, extirpated, and extinct kinds, together with periodical and occasional visitants. pp. i-xxxii [= 1-23], 1-565, [1]. Edinburgh. (Bell & Bradfute).
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^ -MolluscaBase eds. (2022). MolluscaBase. Pupilla J. Fleming, 1828. Accessed through: World Register of Marine Species at: https://www.marinespecies.org/aphia.php?p=taxdetails&id=426395 on 2022-03-05
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^ (in Czech) Ivanov M., Hrdličková, S. & Gregorová, R. (2001). Encyklopedie zkamenělin. Rebo Productions, Dobřejovice, 1. vydání, 312 pp., page 125.
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^ a b c d e f g "Genus summary for Pupilla". AnimalBase. Last modified 02-09-2006, accessed 30 July 2010.
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^ a b c d e f g Pokryszko B. M., Auffenberg K., Hlaváč J. Č. & Naggs F. (2009). "Pupilloidea of Pakistan (Gastropoda: Pulmonata): Truncatellininae, Vertigininae, Gastrocoptinae, Pupillinae (In Part)". Annales Zoologici 59(4): 423-458. doi:10.3161/000345409X484847.
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^ Kessner V. (1996). Pupilla ficulnea. 2006 IUCN Red List of Threatened Species. Downloaded on 7 August 2007.
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^ (in Czech) Ložek V. (1954). "Noví měkkýši československého pleistocénu (Neue Mollusken aus dem tschechoslowakischen Pleistozän). Vertigo pseudosubstriata sp. n., Pupilla muscorum densegyrata ssp. n. a Pupilla loessica sp. n." Anthropozoikum 3(1953): 327–342, Tab. 1.
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^ Mollusc Specialist Group (1996). Pupilla obliquicosta. 2006 IUCN Red List of Threatened Species. Downloaded on 7 August 2007.
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^ Mollusc Specialist Group (1996). Pupilla pupula. 2006 IUCN Red List of Threatened Species. Downloaded on 7 August 2007.
- Flach, K. (1890). Palaeontologische Beiträge. Verhandlungen der Physikalisch-Medicinischen Gesellschaft zu Würzburg, Neue Folge. 24(3): 49-59, pl. 1.
- Schileyko, A. A. (1984). Nazemnye molljuski podotrjada Pupillina fauny SSSR (Gastropoda, Pulmonata, Geophila). In:. Fauna SSSR, Molljuski. III, 3
- Held, F. (1837-1838). Notizen über die Weichthiere Bayerns. Isis (Oken), 30 (4): 303-309 (1837); 30 (12): 901-919 (1838). Leipzig
