Leodice laurillardi (Quatrefages 1866)

Eunice laurillardi Quatrefages, 1866

Eunice laurillardi Quatraefages, 1866:314–315, pl. 10: fig.3—Grube, 1870a:294.

MATERIAL EXAMINED.—Syntype, MNHN, Paris, A.1(R.)- 1868-no. 49b, Nice. Syntype. MNHN-Paris, A.1(R.)-1868-no. 49c, Palermo. Syntype, MNHN-Paris, A.1(R.)-1868-no. 49a, Marseille, coll. M. Barbour (posterior fragment only).

COMMENTS ON MATERIAL EXAMINED.—One additional specimen identified as E. laurillardi, MNHN A1 (R) 1868 no 49d from Palermo, Italy, is a posterior fragment of an arabellid and cannot be further identified. The description is based primarily on the specimen from Nice, except as noted.

DESCRIPTION.—Syntype from Nice complete, of unknown sex, with 189 setigers; total length 150 mm; maximal width 5 mm; length through setiger 10, 9 mm. Syntype from Palermo complete, of unknown sex, with 170 setigers; total length 143 mm; maximal width 4.5 mm; length through setiger 10,9 mm.

Prostomium (Figure 63a) distinctly shorter and narrower than peristomium, less than as deep as peristomium. Prostomial lobes frontally rounded, dorsally slightly flattened; median sulcus deep. Eyes between bases of A-I and A-II. Antennae in transverse row, evenly spaced, similar in thickness. Ceratophores ring-shaped in all antennae, without articulations. Ceratostyles tapering, articulated to bases, with up to 13 articulations in A-III; innermost articulations cylindrical, gradually becoming drop-shaped distally. A-I to setiger 1; A-II to setiger 5; A-III to setiger 8. Peristomium slightly flaring anteriorly. Separation between rings distinct dorsally and ventrally; anterior ring of total peristomial length. Peristomial cirri to middle of prostomium, tapering, with 5 cylindrical articulations.

Maxillary formula 1+1, 6+6, 5+0, 5+8, and 1+1. Left Mx IV short; part of distal arc with Mx III.

Branchiae (Figure 63b, c) present, pectinate, distinctly shorter than notopodial cirri, not reduced in mid-body region, erect. Branchiae from setiger 3 to setiger 184 (five setigers before pygidium). Branchiae present to near posterior end, present on more than 65% of total number of setigers. First branchia single filaments; maximum 10 filaments at about setiger 15. Stems short, flexible. Filaments slender, longer than stems.

Neuropodial acicular lobes rounded; aciculae emerging at midline. Pre- and postsetal lobes low, transverse folds. Anterior ventral cirri tapering, becoming basally inflated by setiger 10. Inflated bases modest, ovate, completely reduced by setiger 35; narrow tips tapering. Posterior ventral cirri broadly attached forming shallow open scoops covering lower edge of neuropodia. Notopodial cirri long, tapering to slender digitiform tips, with 4 to 5 articulations anteriorly and 3 to 4 articulations posteriorly.

Limbate setae narrow, marginally smooth. Pectinate setae (Figure 63e) tapering, flat. One marginal tooth longer than other teeth, with ~15 very slender teeth. Shafts of compound falcigers (Figure 63d, f) tapering, marginally coarsely serrated, internally striated. Appendages narrow, tapering towards small head, bidentate. Proximal teeth very much smaller than distal teeth, nearly button-shaped, directed laterally. Distal teeth tapering, directed obliquely distally. Guards asymmetrically bluntly pointed, marginally smooth; mucros absent. Pseudocompound falcigers and compound spinigers absent. Aciculae mostly paired, dark brown or black, tapering to blunt tips, slightly curved; cross-sections round. Subacicular hooks (Figure 63g) dark brown to black, bidentate. Hooks first present from setiger 32–33, present in all setigers thereafter, always single (except for replacements). Hooks tapering to small head. Proximal teeth very much larger than distal teeth. Both teeth directed distally. Subacicular hooks darker than aciculae.

UNKNOWN MORPHOLOGICAL FEATURES.—Pygidium and anal cirri.

EXPECTED STATES OF SELECTED UNKNOWN FEATURES.—None.

CHARACTERS USED IN PREPARATION OF KEY NOT SCORED.—Inappropriate Characters: 56, 58, 59. Unknown Characters: 4, 6, 74, 78.

ASSUMED STATES FOR PURPOSE OF PREPARING KEY.—None.