Associations
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İngilizce
)
BioImages, the virtual fieldguide, UK tarafından sağlandı
In Great Britain and/or Ireland:
Foodplant / parasite
hypophyllous telium of Chrysomyxa abietis parasitises live leaf of Picea rubens
Remarks: season: 3-5
Comments
(
İngilizce
)
eFloras tarafından sağlandı
Throughout the Appalachians, trees of Picea rubens are dying, possibly as a consequence of environmental pollution. In eastern Canada this species hybridizes to a limited extent with P . mariana (A.G. Gordon 1976).
Red spruce ( Picea rubens ) is the provincial tree of Nova Scotia.
- lisans
- cc-by-nc-sa-3.0
- telif hakkı
- Missouri Botanical Garden, 4344 Shaw Boulevard, St. Louis, MO, 63110 USA
Description
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İngilizce
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eFloras tarafından sağlandı
Trees to 40m; trunk to 1m diam.; crown narrowly conic. Bark gray-brown to reddish brown. Branches horizontally spreading; twigs not pendent, rather stout, yellow-brown, densely pubescent to glabrate. Buds reddish brown, 5--8mm, apex acute. Leaves 0.8--2.5(--3)cm, 4-angled in cross section, somewhat flexuous, yellow-green to dark green, not glaucous, bearing stomates on all surfaces, apex mostly acute to sharp-pointed. Seed cones 2.3--4.5(--5)cm; scales broadly fan-shaped, broadest near apex, 8--12 ´ 8--12mm, stiff, margin at apex entire to irregularly toothed. 2 n =24.
- lisans
- cc-by-nc-sa-3.0
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- Missouri Botanical Garden, 4344 Shaw Boulevard, St. Louis, MO, 63110 USA
Habitat & Distribution
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İngilizce
)
eFloras tarafından sağlandı
Upper montane to subalpine forests; 0--2000m; St. Pierre and Miquelon; N.B., N.S., Ont., P.E.I., Que.; Conn., Maine, Md., Mass., N.H., N.J., N.Y., N.C., Pa., Tenn., Vt., Va., W.Va.
- lisans
- cc-by-nc-sa-3.0
- telif hakkı
- Missouri Botanical Garden, 4344 Shaw Boulevard, St. Louis, MO, 63110 USA
Synonym
(
İngilizce
)
eFloras tarafından sağlandı
Picea australis Small; P. nigra (Aiton) Link var. rubra (DuRoi) Engelmann; P. rubra (DuRoi) Link 1831, not A.Dietrich 1824
- lisans
- cc-by-nc-sa-3.0
- telif hakkı
- Missouri Botanical Garden, 4344 Shaw Boulevard, St. Louis, MO, 63110 USA
Brief Summary
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İngilizce
)
EOL authors tarafından sağlandı
Picea rubens, generally called red spruce, is one of the more important coniferous trees in the northeastern United States and adjacent Canada. Also called yellow spruce, West Virginia spruce, eastern spruce, and he-balsam, it is a medium-size tree that may grow to be more than 400 years old. Red spruce wood is used for making paper, for construction lumber, and for musical stringed instruments. Its many uses rival those of eastern white pine (Pinus strobus). The range of red spruce extends from the Maritime Provinces of Canada west to Maine, southern Quebec, and southeastern Ontario, and south into central New York, eastern Pennsylvania, northern New Jersey, and Massachusetts. It also grows south along the Appalachian Mountains in extreme western Maryland, and eastern West Virginia, and north and west in Virginia, western North Carolina, and eastern Tennessee. Red spruce sometimes occurs in pure stands, and is a major component of several forest types, or it may be found mixed with other conifers and hardwoods. Its shallow root system, thin bark, and flammable needles make trees of all ages very susceptible to fire damage. The acreage of red spruce originally present in the southern Appalachians has been reduced to a fraction of what it once was by fire and clear cutting; it is also susceptible to damage from acid rain. The most important insect enemy of red spruce is the spruce budworm, Choristoneura fumiferana. Although red spruce is much less vulnerable to damage than balsam fir or white spruce, largely due to later bud flushing in the spring, much damage and mortality occur in stands containing large quantities of mature balsam fir. Red spruce wood is light in color and weight, straight-grained, and resilient. The wood of red spruce, white spruce (Picea glauca), and black spruce (Picea mariana) cannot be distinguished with certainty by either gross characteristics or minute anatomy, and all three are usually marketed simply as eastern spruce. Chief uses are for lumber and pulpwood, with limited amounts going into poles piling, boatbuilding stock, and cooperage stock. Flakeboard and plywood have been made from spruce in recent years. It is also the preferred wood for piano sounding boards, guitars, mandolins, organ pipes, and violin bellies. A unique use of red spruce was spruce gum, an exudate that accumulates on trunk wounds. This was the raw material for a flourishing chewing-gum industry in Maine during the last half of the 19th century and early years of this century. Forests with red spruce support diverse wildlife, including many songbirds, ruffed grouse, snowshoe hare, and woodcock, and provide important winter cover for deer and moose. Bird, porcupines, bears, deer, yellow-bellied sapsuckers, and red squirrels all browse or eat various parts of the trees, included twigs, foliage, and vegetative and reproductive buds. Mice and voles consume and store significant amounts of spruce seeds in preference to those of balsam fir, suggesting one reason for the low ratio of spruce to fir seedlings commonly found in naturally regenerated stands. Excerpted and edited from Blum 1990.
- bibliyografik atıf
- Blum, B.M. 1990. <i>Picea rubens</i> Sarg., Red Spruce. In Burns, Russell M., and Barbara H. Honkala, tech. coords. 1990. <i>Silvics of North America: Vol. 1. Conifers</i>. Agriculture Handbook 654. U.S. Department of Agriculture, Forest Service, Washington, DC.
- yazar
- Jacqueline Courteau (Jacqueline Courteau)
Common Names
(
İngilizce
)
Fire Effects Information System Plants tarafından sağlandı
red spruce
yellow spruce
West Virginia spruce
eastern spruce
he-balsam
blue spruce
- bibliyografik atıf
- Sullivan, Janet. 1993. Picea rubens. In: Fire Effects Information System, [Online]. U.S. Department of Agriculture, Forest Service, Rocky Mountain Research Station, Fire Sciences Laboratory (Producer). Available: http://www.fs.fed.us/database/feis/
Cover Value
(
İngilizce
)
Fire Effects Information System Plants tarafından sağlandı
More info for the term:
coverRed spruce provides thermal and loafing cover for spruce grouse in
winter [
62].
- bibliyografik atıf
- Sullivan, Janet. 1993. Picea rubens. In: Fire Effects Information System, [Online]. U.S. Department of Agriculture, Forest Service, Rocky Mountain Research Station, Fire Sciences Laboratory (Producer). Available: http://www.fs.fed.us/database/feis/
Description
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İngilizce
)
Fire Effects Information System Plants tarafından sağlandı
More info for the terms:
duff,
treeRed spruce is a native, evergreen conifer. It is a medium-sized tree,
attaining a maximum height of 115 feet (35 m); the average mature height
is 60 to 75 feet (18-23 m). The ovulate cones are 1.3 to 1.5 inches
(3-4 cm) long, with rigid rounded scales that are often slightly toothed
on the edges. Red spruce is very shallow rooted; most of the feeding
roots occur in the duff and top few centimeters of soil. In Maine, the
average depth of roots was 13 inches (33 cm), with a maximum depth of 22
inches (56 cm) [
9]. Red spruce is long-lived, often achieving ages
greater than 350 years [
1].
- bibliyografik atıf
- Sullivan, Janet. 1993. Picea rubens. In: Fire Effects Information System, [Online]. U.S. Department of Agriculture, Forest Service, Rocky Mountain Research Station, Fire Sciences Laboratory (Producer). Available: http://www.fs.fed.us/database/feis/
Distribution
(
İngilizce
)
Fire Effects Information System Plants tarafından sağlandı
Red spruce occurs from Cape Breton Island, Nova Scotia, and New
Brunswick, west to Maine, southern Quebec, and southeastern Ontario, and
south to central New York, northeastern Pennsylvania, northern New
Jersey, and northeastern Massachusetts. Its range extends south in the
Appalachian Mountains of extreme western Maryland, eastern West
Virginia, northern and western Virginia, western North Carolina, and
eastern Tennessee [
48].
- bibliyografik atıf
- Sullivan, Janet. 1993. Picea rubens. In: Fire Effects Information System, [Online]. U.S. Department of Agriculture, Forest Service, Rocky Mountain Research Station, Fire Sciences Laboratory (Producer). Available: http://www.fs.fed.us/database/feis/
Fire Ecology
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İngilizce
)
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More info for the terms:
avoidance,
fire frequency,
fire regime,
forest,
formation,
frequency,
natural,
severity,
treeRed spruce forests persist without fire. Red spruce is easily killed by
fire due to its thin bark, shallow roots, flammable needles, and lack of
self-pruning [
9,
23,
39]. Its slow early growth rate delays the formation
of a corky layer, which increases the fire susceptibility of young trees
[
39]. In a study based on a survey of foresters, Starker [
76] rated the
fire resistance of 22 New England tree species based on fire mortality
and fire avoidance (occurrence in habitat that does not burn very
often). Red spruce was not resistant in terms o fire mortality but
moderately or very resistant in terms of fire avoidance, and was ranked
13th overall.
Red spruce habitat is subject to few fires; fires that occurred in
presettlement times were usually of low severity [
1]. Saunders [
73]
noted that old-timers claimed that forest fires would stop when they
reached the spruce-fir forest boundary. Electrical storms are common in
this area but are usually accompanied by sufficient rain, and fuels are
usually moist [
32]. Severe surface fires probably occurred
infrequently, during periods of prolonged drought, and usually affected
forests that were breaking up due to wind, ice storm damage, or similar
events that generate surface fuels [
25,
32,
60,
61,
87].
The estimated natural fire return intervals for the northeastern United
States and adjacent Canada range from 330 to 3,300 or more years
[
25,
32,
51,
52,
84]. Estimates of natural fire frequency have been
complicated by human activities. Logging in these forests has resulted
in an increase in fire frequency and intensity, particularly in logging
slash [
18,
32,
52]. The catastrophic fires of the 19th and 20th centuries
can be attributed to human activities [
21,
32,
52]. However, even with
the increase in fires due to human activity, most fires are small and
quickly suppressed. There should be sufficient time between fires for
red spruce to regain dominance on most sites unless deliberately and/or
repeatedly burned.
It has been suggested that, in presettlement forests, the increase of
dead fuels following spruce budworm outbreaks increased the likelihood
of fire [
21,
25,
32]. Such outbreaks are more common in
balsam-fir-dominated forests than in red-spruce-dominated forests, but
the two species usually occur together, in varying proportions.
Before settlement by Europeans, forests in northern New England, the
Adirondack Mountains, and the hillier sections of southern New England
and Pennsylvania were not deliberately burned by Native Americans as
were other areas in the northeastern United States [
18].
FIRE REGIMES : Find fire regime information for the plant communities in which this
species may occur by entering the species name in the
FEIS home page under
"Find FIRE REGIMES".
- bibliyografik atıf
- Sullivan, Janet. 1993. Picea rubens. In: Fire Effects Information System, [Online]. U.S. Department of Agriculture, Forest Service, Rocky Mountain Research Station, Fire Sciences Laboratory (Producer). Available: http://www.fs.fed.us/database/feis/
Fire Management Considerations
(
İngilizce
)
Fire Effects Information System Plants tarafından sağlandı
More info for the terms:
competition,
fire exclusion,
fire management,
forest,
fuel,
natural,
prescribed fire,
treeSome managers believe that prescribed fire may be a useful silvicultural
tool for managing red spruce on some sites. On such sites, the exposed
mineral soil must have plentiful moisture, soil temperatures must be
moderate, and competition must be minimal [
65]. In general, however,
fires in red spruce habitat are of little silvicultural value [
87].
Slash burning following logging kills advance reproduction and creates
rank postfire vegetation that delays any new seedling establishment
[
39].
The fire management plan for Acadia National Park, Maine, dictates the
suppression of natural fires. Prescribed fires may be used on occasion
to reduce fuels [
61]. Patterson and others [
60] estimated fuel loadings
for a number of stands in Acadia National Park that contained red
spruce. They concluded that fire exclusion was probably resulting in
increased fuel loads.
Alexander [
4] compiled slash fuel indices for red spruce and compared
actual fire spread, intensity, and slash and organic layer depletions
with those predicted by the Canadian Forest Fire Danger Rating System.
Freeman and others [
24] developed equations to determine average crown
weight per tree as a function of tree height and diameter for use in a
method to predict slash weight after logging red spruce.
- bibliyografik atıf
- Sullivan, Janet. 1993. Picea rubens. In: Fire Effects Information System, [Online]. U.S. Department of Agriculture, Forest Service, Rocky Mountain Research Station, Fire Sciences Laboratory (Producer). Available: http://www.fs.fed.us/database/feis/
Growth Form (according to Raunkiær Life-form classification)
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İngilizce
)
Fire Effects Information System Plants tarafından sağlandı
More info on this topic. More info for the term:
phanerophytePhanerophyte
- bibliyografik atıf
- Sullivan, Janet. 1993. Picea rubens. In: Fire Effects Information System, [Online]. U.S. Department of Agriculture, Forest Service, Rocky Mountain Research Station, Fire Sciences Laboratory (Producer). Available: http://www.fs.fed.us/database/feis/
Habitat characteristics
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İngilizce
)
Fire Effects Information System Plants tarafından sağlandı
More info for the terms:
cover,
organic soilsRed spruce grows in climates with cool, moist summers and cold winters
[
74]. In the northeastern United States, the mean annual precipitation
ranges from 36 to 52 inches (910-1,320 mm) and is often higher in the
mountainous terrain where red spruce occurs, due to fog drip. Snow
cover averages 80 to 160 inches (203-406 cm), with 100 to 140 days of
snow cover per year [
9].
Most of the soils on which red spruce occurs are developed from glacial
deposits. The most productive soils are derived from parent materials
of unsorted glacial drift and till deposited on the midslopes of hills
and mountains. Soils on red spruce sites are usually acid Spodosols,
Inceptisols, and sometimes Histosols with thick mor humus and a
well-defined A2 horizon. Soil pH ranges from 4.0 to 5.5. Red spruce is
often found on sites that are unfavorable for other species, such as
organic soils overlying rocks in mountainous locales, on steep rocky
slopes with thin soils, and in wet bottomlands [
9].
In the northern part of its range, red spruce occurs at elevations
ranging from sea level to 4,500 feet (0-1,370 m), above which it is
usually replaced by balsam fir (Abies balsamea). The elevational
zonation of species is defined as follows [
67]:
up to 1,485 feet (450 m) northern hardwoods (hemlock phase)
1,486 to 2,508 feet (451- 760 m) northern hardwoods (spruce phase)
2,508 to 4,026 feet (761-1,220 m) subalpine (spruce-fir phase)
4,027 to 4,785 feet (1,221-1,450 m) subalpine (fir phase)
In the southern Appalachian Mountains, red spruce occurs at elevations
from about 3,200 feet to 6,200 feet (980-1,890 m); above 6,200 feet
(1,890 m), red spruce tends is usually replaced by Fraser fir (Abies
fraseri) [
59].
- bibliyografik atıf
- Sullivan, Janet. 1993. Picea rubens. In: Fire Effects Information System, [Online]. U.S. Department of Agriculture, Forest Service, Rocky Mountain Research Station, Fire Sciences Laboratory (Producer). Available: http://www.fs.fed.us/database/feis/
Habitat: Cover Types
(
İngilizce
)
Fire Effects Information System Plants tarafından sağlandı
More info on this topic. This species is known to occur in association with the following cover types (as classified by the Society of American Foresters):
5 Balsam fir
12 Black spruce
16 Aspen
17 Pin cherry
18 Paper birch
21 Eastern white pine
22 White pine - hemlock
23 Eastern hemlock
24 Hemlock - yellow birch
25 Sugar maple - beech - yellow birch
27 Sugar maple
30 Red spruce - yellow birch
31 Red spruce - sugar maple - beech
32 Red spruce
33 Red spruce - balsam fir
34 Red spruce - Fraser fir
35 Paper birch - red spruce - balsam fir
37 Northern white-cedar
60 Beech - sugar maple
107 White spruce
108 Red maple
- bibliyografik atıf
- Sullivan, Janet. 1993. Picea rubens. In: Fire Effects Information System, [Online]. U.S. Department of Agriculture, Forest Service, Rocky Mountain Research Station, Fire Sciences Laboratory (Producer). Available: http://www.fs.fed.us/database/feis/
Habitat: Ecosystem
(
İngilizce
)
Fire Effects Information System Plants tarafından sağlandı
More info on this topic. This species is known to occur in the following ecosystem types (as named by the U.S. Forest Service in their Forest and Range Ecosystem [FRES] Type classification):
FRES10 White - red - jack pine
FRES11 Spruce - fir
FRES18 Maple - beech - birch
FRES19 Aspen - birch
- bibliyografik atıf
- Sullivan, Janet. 1993. Picea rubens. In: Fire Effects Information System, [Online]. U.S. Department of Agriculture, Forest Service, Rocky Mountain Research Station, Fire Sciences Laboratory (Producer). Available: http://www.fs.fed.us/database/feis/
Habitat: Plant Associations
(
İngilizce
)
Fire Effects Information System Plants tarafından sağlandı
More info on this topic. This species is known to occur in association with the following plant community types (as classified by Küchler 1964):
More info for the term:
forestK096 Northeastern spruce - fir forest
K097 Southeastern spruce - fir forest
K108 Northern hardwoods - spruce forest
- bibliyografik atıf
- Sullivan, Janet. 1993. Picea rubens. In: Fire Effects Information System, [Online]. U.S. Department of Agriculture, Forest Service, Rocky Mountain Research Station, Fire Sciences Laboratory (Producer). Available: http://www.fs.fed.us/database/feis/
Immediate Effect of Fire
(
İngilizce
)
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More info for the terms:
litter,
treeRed spruce is easily killed by fire [
49]. Surface or ground fires that
consume the litter and organic layers covering the superficial roots of
red spruce are almost certain to severely injure the roots [
39]. Fire
kills mature trees by exposing roots, subjecting the tree to water
stress and/or windthrow, which may result in the eventual death of the
tree [
39,
87].
- bibliyografik atıf
- Sullivan, Janet. 1993. Picea rubens. In: Fire Effects Information System, [Online]. U.S. Department of Agriculture, Forest Service, Rocky Mountain Research Station, Fire Sciences Laboratory (Producer). Available: http://www.fs.fed.us/database/feis/
Importance to Livestock and Wildlife
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İngilizce
)
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Spruce grouse browse the leaves and twigs of red spruce [
62]. Mice and
voles consume and store significant amounts of spruce seeds, preferring
red and white spruce to balsam fir [
2]. Birds (particularly crossbills
or grosbeaks) will clip the terminal buds of young spruce, as will
porcupines, bears, snowshoe hares, and, rarely, deer [
7,
55,
78]. Red
squirrels clip twigs and terminal buds and also eat reproductive and
vegetative buds [
7,
72].
In the southern part of its range, red spruce forests are used by only a
few wildlife species. Many of these species are usually only found
farther north, such as snowshoe hare, wood warblers and other songbirds,
rodents, and salamanders [
79].
- bibliyografik atıf
- Sullivan, Janet. 1993. Picea rubens. In: Fire Effects Information System, [Online]. U.S. Department of Agriculture, Forest Service, Rocky Mountain Research Station, Fire Sciences Laboratory (Producer). Available: http://www.fs.fed.us/database/feis/
Key Plant Community Associations
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İngilizce
)
Fire Effects Information System Plants tarafından sağlandı
More info for the terms:
bryophytes,
codominant,
cover,
fern,
forest,
lichens,
natural,
shrub,
treeRed spruce is a common dominant or codominant in the red spruce and the
spruce-fir forests of the northeastern United States and adjacent
Canada.
Shrub associates of red spruce in the Adirondack Mountains of New York
include red raspberry (Rubus idaeus), dwarfed blackberry (R. pubescens),
hobblebush (Viburnum alnifolium), Canada yew (Taxus canadensis), and
American fly honeysuckle (Lonicera canadensis). Ground layer herbs
include wild sarsaparilla (Aralia nudicaulis), Aster acuminatus, yellow
beadlily (Clintonia borealis), and common wood-sorrel (Oxalis montana).
Common bryophytes found in old-growth red spruce forests in the
Adirondacks include Brotherella recurvans, Schreber's moss (Pleurozium
schreberi), Polytrichum ohioense, mountain fern moss (Hylocomium
splendens), Bazzania trilobata, ptilium (Ptilium crista-castrensis),
Drepanocladus uncinatus, Dicranum scoparium, and D. montanum [
47].
In the southern Appalachian Mountains, arboreal associates include
Fraser fir (Abies fraseri), yellow buckeye (Aesculus octandra), sweet
birch (Betula lenta), and black cherry (Prunus serotina) in addition to
those found in the northern part of its range [
59,
79,
87]. Understory
associates in openings include rhododendrons (Rhododendron spp.),
American mountain-ash (Sorbus americana), and wild raisin (Viburnum
cassinoides). Other understory associates include highbush cranberry
(Viburnum edule), mountain holly (Ilex montana), mountain laurel (Kalmia
latifolia), speckled alder (Alnus rugosa), pin cherry (Prunus
pensylvanica), serviceberry (Amelanchier spp.), raspberries (Rubus
spp.), and blueberries and huckleberries (Vaccinium spp.). In closed
red spruce stands, mosses, lichens, and clubmosses predominate in the
understory along with wood sorrel (Oxalis spp.), trillium (Trillium
spp.), and checkerberry wintergreen (Gaultheria procumbens) [
79].
Publications describing habitat or cover types in which red spruce is
dominant or codominant include:
(1) Proceedings of the Region 9 Land Systems conference on the White
Mountain National Forest [
5]
(2) The Hubbard Brook ecosystem study: composition and dynamics of the
tree stratum [
11]
(3) Ground vegetation patterns of the spruce-fir area of the Great
Smoky Mountains National Park [
14]
(4) Spruce-fir forests of the coast of Maine [
16]
(5) Forest type studies in the Adirondack region [
31]
(6) The classification and evaluation of site for forestry [
33]
(7) The identification and description of forest sites [
34]
(8) Old-growth forests of Adirondack Park, New York [
47]
(9) Vegetation-environment relationships in virgin, middle elevation
forests in the Adirondack Mountains, New York [
68]
(10) Natural ecological communities of New York State [
71]
- bibliyografik atıf
- Sullivan, Janet. 1993. Picea rubens. In: Fire Effects Information System, [Online]. U.S. Department of Agriculture, Forest Service, Rocky Mountain Research Station, Fire Sciences Laboratory (Producer). Available: http://www.fs.fed.us/database/feis/
Life Form
(
İngilizce
)
Fire Effects Information System Plants tarafından sağlandı
More info for the term:
treeTree
- bibliyografik atıf
- Sullivan, Janet. 1993. Picea rubens. In: Fire Effects Information System, [Online]. U.S. Department of Agriculture, Forest Service, Rocky Mountain Research Station, Fire Sciences Laboratory (Producer). Available: http://www.fs.fed.us/database/feis/
Management considerations
(
İngilizce
)
Fire Effects Information System Plants tarafından sağlandı
More info for the terms:
basal area,
litter,
natural,
seed,
seed tree,
selection,
treeSilviculture: Various silvicultural systems may be used to manage red
spruce. Single tree selection, group selection, shelterwood, and strip
clearcut are all practical harvesting methods. Red spruce is subject to
windthrow; partial cuttings are recommended not to exceed half of the
basal area, and a lighter harvest is usually better. Seed tree cuts are
not recommended [
6,
9]. Frank and Blum [
23] recommend a selection
silviculture where net growth is maximized by a 10-year, intensive
selection system. Clearcuts are contraindicated for many soil types and
fertility levels [
35].
Postharvest red spruce regeneration is entirely dependent on advance
reproduction. If seedlings are not present at the time of logging, any
new spruce seedlings will be quickly overtopped and suppressed by faster
growing hardwoods [
17]. Leaf litter may aid in red spruce
regeneration. Harvesting during the dormant season or allowing
harvested trees to dry on site has been recommended to increase litter
[
35]. Loucks [
53] noted that in the Maritime Provinces of Canada, red
spruce regeneration is usually good following partial cuts but may be
lacking in clearcuts.
The extent of red spruce forests has decreased following extensive
logging practices and subsequent fire [
3]. In the mountains of central
West Virginia, it is estimated that approximately 500,000 acres (200,000
ha) of red spruce present in the late 19th century had been reduced to
less than 60,000 acres (24,000 ha) by 1975, and as little as 17,500
acres (7,000 ha) in 1978 [
10,
73].
Management for wildlife: Harvest practices have an effect on the
resulting stand structure, and therefore on the numbers and species of
birds that use red spruce habitats. Crawford and Titterington [
15]
identified five seral stages and the corresponding bird species, and
made associated recommendations for management of spruce-fir stands.
They also determined that spruce budworm infestation increases both the
number and diversity of birds. Dense, young stands of red spruce
support a higher population of birds but with less diversity than in
older forests.
Insects and disease: Red spruce is relatively free from insects and
diseases until it is mature. Mature trees are susceptible to the
following insects: spruce budworm (Choristoneura fumiferana), eastern
spruce beetle (Dendroctonus rufipennis), European spruce sawfly (Diprion
hercyniae), yellowheaded spruce sawfly (Pikonema alaskensis), and
eastern spruce gall adelgid (Adelges abietis) [
9,
22,
23,
30]. Diseases of
red spruce have been detailed [
9,
22,
23,
30,
47].
Red spruce decline: Throughout its range, growth rates of red spruce
have declined and mortality has increased [
36]. This decline is
apparently more severe at higher elevations, in older stands, and on
more exposed sites. This decline is not limited to red spruce; balsam
fir and associated white and black spruce appear to be affected also
[
85]. A number of studies on the causes of red spruce decline have
failed to make a definitive case for any single cause. There may be no
single cause or the complexity of the situation may not lend itself to a
clear cause-effect relationship [
36,
42,
47]. The combination of climatic
stress and atmospheric pollution is probably the major cause of this
decline, according to a number of researchers [
19,
36,
41,
42]. Numerous
other causes have been proposed as well, including a natural cycle of
dieback and recovery [3, 36,]. A survey of the extent and identifiable
causes of mortality and decline was published in 1985 [
85].
- bibliyografik atıf
- Sullivan, Janet. 1993. Picea rubens. In: Fire Effects Information System, [Online]. U.S. Department of Agriculture, Forest Service, Rocky Mountain Research Station, Fire Sciences Laboratory (Producer). Available: http://www.fs.fed.us/database/feis/
Occurrence in North America
(
İngilizce
)
Fire Effects Information System Plants tarafından sağlandı
CT ME MD MA NH NJ NY NC PA TN
VT VA WV NB NS PE PQ
- bibliyografik atıf
- Sullivan, Janet. 1993. Picea rubens. In: Fire Effects Information System, [Online]. U.S. Department of Agriculture, Forest Service, Rocky Mountain Research Station, Fire Sciences Laboratory (Producer). Available: http://www.fs.fed.us/database/feis/
Other uses and values
(
İngilizce
)
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Red spruce gum was formerly collected and processed for chewing gum [
29].
- bibliyografik atıf
- Sullivan, Janet. 1993. Picea rubens. In: Fire Effects Information System, [Online]. U.S. Department of Agriculture, Forest Service, Rocky Mountain Research Station, Fire Sciences Laboratory (Producer). Available: http://www.fs.fed.us/database/feis/
Palatability
(
İngilizce
)
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Red spruce is unpalatable to white-tailed deer [
78].
- bibliyografik atıf
- Sullivan, Janet. 1993. Picea rubens. In: Fire Effects Information System, [Online]. U.S. Department of Agriculture, Forest Service, Rocky Mountain Research Station, Fire Sciences Laboratory (Producer). Available: http://www.fs.fed.us/database/feis/
Phenology
(
İngilizce
)
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More info on this topic. Red spruce vegetative buds begin growth from May 26 to June 3 [
8].
Needles are shed early in summer [
12]. Reproductive cones open in late
April to early May [
29,
72]. Red spruce cones mature the first autumn
from mid-September to mid-October [
29,
39]. Dissemination of seeds
begins soon after cones are ripe and continues until March [
29].
- bibliyografik atıf
- Sullivan, Janet. 1993. Picea rubens. In: Fire Effects Information System, [Online]. U.S. Department of Agriculture, Forest Service, Rocky Mountain Research Station, Fire Sciences Laboratory (Producer). Available: http://www.fs.fed.us/database/feis/
Plant Response to Fire
(
İngilizce
)
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More info for the terms:
duff,
ferns,
forest,
heath,
seed,
shrubs,
succession,
treeRed spruce does not sprout. Seed germination is greater on burned areas
with exposed mineral soil than in duff; mortality, however, is also
greater due to increased surface temperature and drought [
63].
Burned red spruce or spruce-fir stands are initially restocked by aspen
(Populus spp.) or birch (Betula spp.) via wind-disseminated seed; paper
birch (Betula papyrifera)-aspen stands are particularly diagnostic of
fire in upland red spruce forests [
52]. Red spruce seedlings appear a
few years after fire, developing as an understory in the aspen-birch
complex, and eventually penetrate the overstory after 50 or 60 years.
Birch and aspen become decadent after 75 to 80 years and red spruce or
red spruce and balsam fir regain dominance if left undisturbed
[
49,
52,
65]. On better sites, northern hardwoods, chiefly sugar maple
and American beech, may replace red spruce, and in some areas, balsam
fir will dominate the late postfire succession. Postharvest/postfire
restocking by red spruce is extremely slow where the organic layers are
destroyed by severe fire (particularly where harvest has been heavy)
[
49].
In Nova Scotia, mature spruce forests have few herbs and shrubs in the
understory. After a fire, herbs increase in the first 6 years and
dominate for 40 or more years while conifers slowly establish [
54].
After fire in the southern Appalachians, blackberry (Rubus ursinus) and
red raspberry colonize the site. Pin cherry and yellow birch follow.
Blackberry and raspberry are too competitive for red spruce and must be
shaded out by the hardwoods before red spruce can establish [
39].
In West Virginia, postlogging and postfire succession in red spruce
forests follows a similar pattern: ferns and raspberry are followed by
other shrubs, then hardwoods (particularly hawthorn [Crataegus spp.]),
and eventually spruce. In many areas, this successional pattern has
been extremely slow; heaths or barrens form that do not appear as if
they will ever return to forest [
13]. Martin [
54] studied
postlogging/postfire succession in Nova Scotia and found that red spruce
was present on most sites after the second postfire year, becoming more
numerous and dominant in the later seres. He concluded that repeated
heavy cuttings and light fires on the poorer soils of the southern
upland of Nova Scotia encourages the invasion of heath plants, which
limits the rate and amount of tree regeneration.
- bibliyografik atıf
- Sullivan, Janet. 1993. Picea rubens. In: Fire Effects Information System, [Online]. U.S. Department of Agriculture, Forest Service, Rocky Mountain Research Station, Fire Sciences Laboratory (Producer). Available: http://www.fs.fed.us/database/feis/
Post-fire Regeneration
(
İngilizce
)
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More info for the terms:
root crown,
secondary colonizerTree without adventitious-bud root crown
Secondary colonizer - off-site seed
- bibliyografik atıf
- Sullivan, Janet. 1993. Picea rubens. In: Fire Effects Information System, [Online]. U.S. Department of Agriculture, Forest Service, Rocky Mountain Research Station, Fire Sciences Laboratory (Producer). Available: http://www.fs.fed.us/database/feis/
Regeneration Processes
(
İngilizce
)
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More info for the terms:
cone,
duff,
forest,
litter,
seed,
treeRed spruce reproduces exclusively by seed. The first cone crop is
usually produced when the crown first reaches direct light [
27,
39].
Therefore, red spruce can bear cones as early as 15 to 20 years of age;
cone production peaks about 15 years later. In dense, even-aged stands,
full cone crops are rare until the trees are 40 to 50 years old [
39].
Good seed crops are produced every 3 to 8 years, with light crops in
intervening years. Cones are dropped shortly after they are mature [
9].
The seeds are wind or rain disseminated. The maximum distance for
dispersal by wind is approximately 201 feet (61 m) [
27]. Seeds do not
exhibit dormancy. Most germinate the spring following dispersal;
occasionally germination will occur in the fall soon after seeds drop
from the tree. Seeds are usually not viable after 1 year. Germination
is largely controlled by moisture availability. Seeds will germinate in
almost any medium except sod. Seeds that germinate in thick duff are
subject to overheating and/or drought mortality. Drought and
frost-heave are the major causes of seedling mortality the first year
[
9].
Successful reproduction appears to depend more on seedling survival than
on germination requirements [
9]. Seedling establishment is usually best
on shallow, less fertile soils that discourage competitive hardwoods
[
87]. The primary roots of red spruce seedlings do not penetrate litter
and forest duff to any depth [
38]. Red spruce seedlings have a root
system of finely branched rootlets and no strong laterals; they depend
entirely on the humus for nutrients and water [
57].
- bibliyografik atıf
- Sullivan, Janet. 1993. Picea rubens. In: Fire Effects Information System, [Online]. U.S. Department of Agriculture, Forest Service, Rocky Mountain Research Station, Fire Sciences Laboratory (Producer). Available: http://www.fs.fed.us/database/feis/
Successional Status
(
İngilizce
)
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More info on this topic. More info for the terms:
climax,
cover,
hardwood,
treeOn shallow, acidic, glacial till soils, red spruce is considered climax.
It is usually subclimax on fertile, well-drained slopes and on abandoned
fields and pastures where is is replaced by shade-tolerant hardwoods
such as sugar maple and beech. Other types, such as red spruce-balsam
fir and red spruce-yellow birch are usually climax [
20].
Red spruce is tolerant of shade. Seedlings of red spruce can establish
in as little as 10 percent of full sunlight, but for optimum growth, at
least 50 percent of full sunlight is needed [
9,
75,
81]. Growth tends to
be suppressed in shade, but such suppression can persist for many years
without killing the tree. For example, suppressed understory
individuals may be 4 to 5 feet (1.2-1.5 m) tall, and be more than 50
years old. In comparison, open-grown red spruce can reach sawtimber
size at 50 years [
9,
29].
Red spruce responds to canopy removal even after many years of
suppression. The taller and older a seedling or sapling is, the greater
is its response to release, up to about 55 years of age after which
response to release starts to decline. However, the amount of response
does not revert to seedling levels until the tree is around 100 years of
age. Umbrella-shaped saplings 40 to 80 years old that have been
suppressed will respond to release after a delay of several years, and
in fact have an advantage because they are taller than smaller,
healthier saplings which respond more quickly to canopy opening. More
than half of mature red spruce second growth arises from larger but
suppressed advance growth, as opposed to having arisen from small
advance growth or new seedlings [
16]. Upon release, 60-year-old red
spruce growth exceeds that of same-age balsam fir and therefore tends to
dominate the canopy [
56].
Leak [
44] defined red spruce in New Hampshire as a dominating climax
species on shallow, dry, wet, or poorly aerated soils; it is a minor
component in young stands but increases markedly over time until it is a
canopy dominant. He estimated that, if undisturbed, red spruce can
reach densities of 70 to 80 percent in a minimum of 250 years. Red
spruce is a long-lived species and, once established, persists as a
dominant for a long time.
Davis [
16] observed young spruce-fir stands in coastal Maine originating
in open sites and as the understory to early seral hardwoods such as
paper birch. The young, open-grown stands may be dominated by white
spruce, red spruce, or balsam fir in any proportions. A spruce-fir
stand originating as understory tends to be dominated by red spruce
and/or balsam fir, though white spruce is often present. Moore [
58]
found red spruce forests to be even-aged in groups, indicating that
establishment and/or canopy achievement tends to occur in openings.
Red spruce and red spruce-fir cover types are self-maintaining. Stand
composition may vary with stand age. Both red spruce and its two fir
associates (balsam and Fraser) are shade tolerant, and both spruce and
fir reproduction are found under spruce-fir canopies [
6,
16]. In the
Catskill Mountains of New York, balsam fir reproduction predominated
under both spruce and balsam fir stands. Both red spruce and balsam fir
reproduction occurred at low densities under hardwood stands (mostly
yellow birch) [
55]. McIntosh and Hurley [
55] do not believe that red
spruce forests form a self-perpetuating climax in this area. Their
conclusion may be biased, however, since balsam fir outcompetes red
spruce in early stages, but is usually overtopped or outcompeted by red
spruce in more mature forests [
16]. Flieger [
21] described 350-year-old
stands of red spruce which were characterized by irregular stocking and
variable crown heights and widths, with at least two age classes
apparent. Most virgin red spruce forests are uneven-aged, indicating
that the forests did no originate following stand-destroying
disturbances, and that red spruce is able to reproduce under its own
canopy [
52].
- bibliyografik atıf
- Sullivan, Janet. 1993. Picea rubens. In: Fire Effects Information System, [Online]. U.S. Department of Agriculture, Forest Service, Rocky Mountain Research Station, Fire Sciences Laboratory (Producer). Available: http://www.fs.fed.us/database/feis/
Synonyms
(
İngilizce
)
Fire Effects Information System Plants tarafından sağlandı
Picea rubra (Du Roi) Link
Picea australis Small
Picea nigra var. rubra Engelmann.
- bibliyografik atıf
- Sullivan, Janet. 1993. Picea rubens. In: Fire Effects Information System, [Online]. U.S. Department of Agriculture, Forest Service, Rocky Mountain Research Station, Fire Sciences Laboratory (Producer). Available: http://www.fs.fed.us/database/feis/
Taxonomy
(
İngilizce
)
Fire Effects Information System Plants tarafından sağlandı
More info for the term:
naturalThe accepted scientific name for red spruce is Picea rubens Sarg. There
are no subspecies, varieties, or forms [
48,
64].
Natural hybrids with black spruce (P. mariana) have been reported
[
9,
48].
- bibliyografik atıf
- Sullivan, Janet. 1993. Picea rubens. In: Fire Effects Information System, [Online]. U.S. Department of Agriculture, Forest Service, Rocky Mountain Research Station, Fire Sciences Laboratory (Producer). Available: http://www.fs.fed.us/database/feis/
Value for rehabilitation of disturbed sites
(
İngilizce
)
Fire Effects Information System Plants tarafından sağlandı
Red spruce is occasionally used for revegetation of coal mine sites in
West Virginia, primarily at high elevations, but it is of limited value
for this purpose [
82].
- bibliyografik atıf
- Sullivan, Janet. 1993. Picea rubens. In: Fire Effects Information System, [Online]. U.S. Department of Agriculture, Forest Service, Rocky Mountain Research Station, Fire Sciences Laboratory (Producer). Available: http://www.fs.fed.us/database/feis/
Wood Products Value
(
İngilizce
)
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Red spruce is one of the more important timber species in the
northeastern United States. The wood is light in weight, straight
grained, and resilient. It is used for paper, construction lumber, and
is highly preferred for musical instruments [
9,
29].
- bibliyografik atıf
- Sullivan, Janet. 1993. Picea rubens. In: Fire Effects Information System, [Online]. U.S. Department of Agriculture, Forest Service, Rocky Mountain Research Station, Fire Sciences Laboratory (Producer). Available: http://www.fs.fed.us/database/feis/
Associated Forest Cover
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İngilizce
)
Silvics of North America tarafından sağlandı
Pure stands of red spruce comprise the forest cover type Red Spruce
(Society of American Foresters Type 32). Red spruce is also a major
component in 5 and a minor component in 13 other forest cover types (10):
5 Balsam Fir
12 Black Spruce
16 Aspen
17 Pin Cherry
18 Paper Birch
21 Eastern White Pine
22 White Pine-Hemlock
23 Eastern Hemlock
25 Sugar Maple-Beech-Yellow Birch
30 Red Spruce-Yellow Birch
31 Red Spruce-Sugar Maple-Beech
33 Red Spruce-Balsam Fir
34 Red Spruce-Fraser Fir
35 Paper Birch-Red Spruce-Balsam Fir
37 Northern White-Cedar
60 Beech-Sugar Maple
107 White Spruce
108 Red Maple
Some of the shrubs associated with red spruce are: blueberry (Vaccinium
spp.), hobblebush (Viburnum lantanoides), witherod (V.
cassinoides), rhodora (Rhododendron canadense), lambkill (Kalmia
angustifolia), mountain-holly (Nemopanthus mucronata), speckled
alder (Alnus rugosa), red raspberry (Rubus idaeus var.
strigosus), creeping snowberry (Gaultheria hispidula), wintergreen
(G. procumbens), fly honeysuckle (Lonicera canadensis), gooseberry
(Ribes spp.), witch-hazel (Hamamelis virginiana), downey
serviceberry (Amelanchier arborea), beaked hazel (Corylus
cornuta), and Canada yew (Taxus canadensis).
A number of mosses and herbs are also found growing in red spruce forest
types. Certain mosses, herbs, and shrubs, however, have been shown to be
related to site quality of red spruce (22). The three main
associations, Hylocomium/Oxalis, Oxalis/Cornus, and Viburnum/0xalis,
in that order, indicate increasing site productivity and increasing
hardwood competition. Similar site types in the higher elevations of the
Appalachian Mountains of North
Carolina include Hylocomium/Oxalis on north-facing slopes above
1520 m (5,000 ft), Oxalis/Dryopteris at high elevations and all
exposures, and the best site type for red spruce and Fraser fir, Viburnum/Vaccinium/Dryopteris
(47).
The Oxalis/Cornus association is considered the best for growing
conditions in the northern part of the range. On these sites the soil is
rich enough for red spruce but not fertile enough for the tolerant
hardwoods to offer serious competition (22).
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Climate
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Silvics of North America tarafından sağlandı
Red spruce grows best in a cool, moist climate. The climate of the
northeastern part of its range can be summarized as follows: annual
precipitation (total), 910 to 1320 mm (36 to 52 in); annual snowfall, 203
to 406 cm (80 to 160 in); days with snow cover, 100 to 140; January
temperature, -7° to -1° C (20° to 30° F) maximum and
-18° to -13° C (0° to 8° F) minimum; July temperature,
21° to 27° C (70° to 80° F) maximum, and 11° to
14° C (52° to 58° F) minimum; frost-free days, 90 to 150
(28). Red spruce attains maximum development in the higher parts
of the southern Appalachian Mountains where the atmosphere is more humid
and the rainfall heavier during the growing season than in other parts of
its range (47). Local extension of the range of red spruce, as along the
southern Maine coast, is related to marine exposure, which provides a cool
growing season and ample moisture supply (8).
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Damaging Agents
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İngilizce
)
Silvics of North America tarafından sağlandı
The shallow root system, thin bark, and
flammable needles of red spruce make trees of all ages very susceptible to
fire damage (11). The acreage of red spruce originally present in the
southern Appalachians has been reduced to a fraction of what it once was
by fire and clearcutting (22). Many former spruce sites are occupied by
inferior tree species, blackberries, and ferns after 20 years (47).
The most important insect enemy of red spruce is the spruce budworm,
Choristoneura fumiferana. Although red spruce is much less
vulnerable to damage than balsam fir or white spruce, largely due to later
bud flushing in the spring (3), much damage and mortality occur in stands
containing large quantities of mature balsam fir. Blum and McLean (4)
suggest that factors such as stand age, species composition, density, and
vigor contribute to the vulnerability of spruce-fir stands to budworm
damage and suggest steps to alleviate damage. Additional, detailed
information may also be found in Sanders, et al. (42) for
spruce-fir stands in the Northeast, the Lake States, and Canada.
The eastern spruce beetle, Dendroctonus rufipennis, damages
mature trees of red spruce. Two species of sawflies, the European spruce
sawfly, Diprion hercyniae, and the native yellowheaded spruce
sawfly, Pikonema alaskensis, have severely defoliated red spruce
in localized areas (22). The eastern spruce gall adelgid, Adelges
abietis, can be a serious pest on spruce when abundant. The pine leaf
adelgid, Pineus pinifoliae, forms unsightly but relatively
harmless conelike galls on red and black spruce (Picea mariana), which
are alternate hosts (46).
Red spruce has few diseases. Needle cast caused by Lirula macrospora
may result in severe defoliation of the lower crown and a subsequent
reduction of growth. Phellinus pini and Phaeolus schweinitzii,
the most destructive of red spruce wood-rotting fungi, are usually
confined to overmature or damaged trees. Climacocystis borealis causes
butt rot in overmature trees (22). Trees are occasionally attacked by Armillaria
mellea and Inonotus tomentosa.
All along the eastern Appalachian mountain chain, from the New England
states to Georgia, growth has declined in high-elevation red spruce since
the 1960's (25). In recent years, this decline has been accompanied by
increased mortality and crown damage in high-elevation red spruce.
Apparently, no significant natural biotic or abiotic causal agents have
been identified, although it has been hypothesized that interaction among
naturally occurring insect and disease factors and anthropogenic air
pollutants, or air pollutants acting alone, are at the root of the
problem. Sulphur dioxide (S02), nitrogen oxides (NOx), and
volatile organic compounds are the pollutants of primary concern;
secondary pollutants such as ozone and nitric and sulfuric acids are also
believed to be important factors (29).
Growth decline and mortality in low-elevation red spruce in northern New
England, while increasing in some areas, appear to be within the normal
ranges for trees and forests of various ages, compositions, and density.
However, some foliar symptoms have been detected in both red spruce and
white pine, particularly from ozone exposure.
Red spruce is occasionally infected with eastern dwarf mistletoe, Arceuthobium
pusillum, a parasite causing growth reduction, tree mortality, and
degradation of wood quality (24).
Mice and voles have been found to consume and store significant amounts
of spruce seeds in preference to those of balsam fir, suggesting one
reason for the low ratio of spruce to fir seedlings commonly found in
naturally regenerated stands (1,23). Wildlife damage to the terminal buds
of young spruce, presumably by birds, also has been noted (2). Some injury
and mortality are also caused occasionally by porcupines, bears, deer, and
yellow-bellied sapsuckers (11). Red squirrels clip twigs and terminals and
eat reproductive and vegetative buds (41).
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Flowering and Fruiting
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İngilizce
)
Silvics of North America tarafından sağlandı
Red spruce is monoecious; male and
female flower buds open in May in axils of the previous year's shoots on
different branches of the same tree. The pendant male flowers are bright
red; female flowers are erect and bright green tinged with purple (21).
Although cone buds differentiate as early as July preceding flowering
in the following spring, they are difficult to distinguish until
September. For experienced workers they provide a possible means of
identifying seed years at that time. The cones mature from about
mid-September to early October, the autumn following flowering (41).
Cones are 3 to 4 cm (1.3 to 1.5 in) long, light reddish brown, with
rigid, rounded scales often slightly toothed on the edges. Cones are
receptive to pollen when fully open, a condition which lasts for only a
few days.
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Genetics
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İngilizce
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Silvics of North America tarafından sağlandı
Successful interspecific crosses with Picea rubens as male or
female parents have been reported or confirmed for P. mariana, P.
omorika, P. glehnii, P. orientalis, and P. koyamai (15); P.
sitchensis (14); P. glauca, P. mexicana (16); P. x
lutzii Little (P. sitchensis x P. glauca), P.
maximowiczii, and P. likiangensis (19).
Crossability of P. rubens with P. omorika is good with
P. mexicana and P. likiangensis moderate; with P.
mariana, P. orientalis, P. maximowiczii, and P. glehnii fair
to poor; and with P. koyamai, P. sitchensis, P. x lutzii,
and P. glauca very poor. Several species fail to cross with P.
rubens (15,16,18,19).
Hybrids between P. rubens and P. mariana occur to some
extent in nature, but parental species remain phenotypically pure in their
characteristic habitats (15,30,31,34,35).
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Growth and Yield
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Silvics of North America tarafından sağlandı
Red spruce is a medium-size tree at maturity,
reaching 30 to 61 cm (12 to 24 in) in d.b.h. and 18 to 23 m (60 to 75 ft)
in height in the Northeast, and up to 35 m (115 ft) in the Appalachian
Mountains. Its maximum age is about 400 years (22). The American Forestry
Association lists a tree 133 cm (52.5 in) in d.b.h. and 33.5 m (110 ft)
tall in Great Smoky National Park in North Carolina as the largest living
red spruce.
The rate of red spruce's growth is strongly influenced by light
conditions. Although trees can live in dense shade for many years, once
they reach sapling to pole stage nearly full sunlight is beneficial.
Understory trees no more than 1.2 or 1.5 m (4 to 5 ft) tall may be more
than 50 years old, whereas trees of the same age in the open may be
approaching small sawtimber size (22).
Under favorable conditions, red spruce may reach an average d.b.h. of 10
cm (4 in) and height of 7 m (23 ft) in 20 years, and be over 23 cm (9 in)
in d.b.h. and 19 m (62 ft) tall in 60 years (22).
Diameter growth of red spruce has been related to vigor, live crown
ratio (ratio of live crown to total height), live crown length, and
initial diameter at breast height (6,32). High vigor red spruce with a
live crown ratio of 0.5 or better averaged 4.3 cm (1.7 in) of diameter
growth in 10 years. Growth rates of trees with smaller crown ratios and
less vigorous trees decreased progressively to an average of 0.8 cm (0.3
in) in 10 years for trees of low vigor or with crown ratios smaller than
0.4 (22). A tree classification for red spruce is shown in table 1 (11).
Table 1- Classification of red spruce trees (11).
Tree class
(rating as
growing stock)
Vigor
Crown class
Live
crown
ratio¹
Average 10-year growth in d.b.h.
cm
in
A, superior
I
Dominant and
Intermediate
0.6+
4.6
1.8
B, good
I
Dominant and
Intermediate
0.3 to 0.5
3.3
1.3
C, acceptable
II
Overtopped
Intermediate
Dominant
0.6+
0.6+
0.6+
2.3
0.9
D, inferior
Intermediate
0.3 to 0.5
1.5
0.6
E, undesirable
III
Intermediate
All others
0.3+
0.3 or less
0.5
0.2
¹Ratio of live crown to total
height.
In one study (40), average net annual growth in softwood stands (66 to
100 percent softwood species) that can be expected from stands receiving
minimal silvicultural input was found to be about 3.5 m³/ha (50 ft³/acre).
In mixed-wood stands (21 to 65 percent softwood species) this dropped to
about 2.8 m³/ha (40 ft³/acre), although the majority of the
growth was contributed by softwoods. A further breakdown of the data shows
the contributions of spruce, most of which was assumed to be red spruce,
to be 51 percent in softwood stands and 39 percent in mixed-wood stands.
Yields per acre, in total volumes of all trees larger than 1.5 cm (0.6
in) in d.b.h. (inside bark and including stump and top but not butt
swell), are given in table 2 (33).
Table 2- Yield of red spruce by age class and site index
(adapted from 33)
Site index¹
Age
12.2 m
or 40 ft
15.2 m
or 50 ft
18.3 m
or 60 ft
21.3 m
or 70 ft
yr
m³/ha
20
6
8
11
14
40
94
132
164
200
60
244
335
422
507
80
308
424
533
640
100
332
456
575
691
yr
ft³/acre
20
80
120
160
200
40
1,350
1,890
2,350
2,850
60
3,490
4,780
6,030
7,240
80
4,400
6,060
7,610
9,150
100
4,740
6,250
8,210
9,870
¹Base age 50 years when age
is measured at d.b.h.- total tree age is estimated to be 65 years at the
time.
These yields are normal yields from even-aged stands growing primarily
on old fields. Therefore, they are higher than yields that might be
expected from more irregular stands such as those developing after cutting
(22).
Site index has not been of great utility in rating the potential
productivity of spruce-fir sites because of the tolerance of the species
and its ability to survive in a suppressed state. Site index at base age
50 years is as good a measure of productivity as any of several growth
functions, however (39). Recently, polymorphic site index curves were
developed for even-aged spruce and fir stands in northern Maine; they
should be valuable for estimating site productivity (20).
Other yield tables for the Northeast (48) take into consideration stand
density, composition, and time since cutting. These tables give
merchantable volume of spruce and fir combined in trees 15.2 cm (6 in) in
d.b.h. and larger from a 0.3 m (1 ft) stump to a 7.6 cm (3 in) top,
diameter inside bark, and are somewhat conservative. Yields of
merchantable volume for different stand densities from 10 to 50 years
after cutting, where 90 percent of the trees are spruce and fir growing on
predominantly softwood sites, are given in table 3.
Table 3- Merchantable yield of red spruce (adapted from
48)
Density index (regional average 100)
Years since cut
50
100
150
m³/ha
10
17.1
24.4
29.5
20
29.8
37.7
43.3
30
43.5
52.0
58.0
40
58.1
67.3
73.4
50
73.8
83.1
89.7
ft³/acre
10
245
349
422
20
425
539
618
30
622
743
828
40
830
961
1,049
50
1,054
1,187
1,281
The development of stand projection growth models that permit computer
simulation of red spruce tree growth for various management practices and
silvicultural treatments over a range of stand conditions has flourished
in recent years. For example, the model FIBER was developed in the
Northeast (43) for spruce-fir, northern hardwood and a range of Mixedwood
forest types between the two. Such models have proved very useful for
forest management planning.
In recent years, interest in total biomass yield and productivity has
increased, and in the future is likely to become more important in
management considerations. As an example, above-ground biomass and
productivity values of typical red spruce stands in Canada are given in
table 4 for stands in a steady state, across a moisture regime catena
(17).
Table 4- Aboveground biomass and annual production of
all tree components and foliage for red spruce at latitude 45° 30' N.
(adapted from 17)
Moisture regime
Biomass
Annual
Production
t/ha
tons/acre
t/ha
tons/acre
Dry
121.3
54.1
4.5
2.0
Fresh
263.2
117.4
8.7
3.9
Moist
461.3
205.8
9.9
4.4
Wet
164.1
73.2
3.8
1.7
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Reaction to Competition
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İngilizce
)
Silvics of North America tarafından sağlandı
Red spruce is classified as shade
tolerant in the United States and tolerant or very tolerant in Canada.
Opinions differ as to whether red spruce is more tolerant than balsam fir,
but the relative tolerance may vary with soil fertility and climate (22).
The species' chief competition comes from balsam fir and hardwoods that
produce heavy shade, like beech and maple. Competition from aspen, birch,
and other thin-crowned species is not so severe. Red spruce prunes itself
about as well as most softwoods in dense stands. As much as one-third of
the live crown may be pruned artificially without seriously affecting
radial growth (5).
A number of studies have demonstrated the ability of red spruce to
respond to release after many years of suppression. The vigor of this
response does decline somewhat with age, however, and older trees may
require about 5 years to recover before showing accelerated growth (7).
Reduction of growth to about 2.5 cm (1 in) of diameter in 25 years, for a
duration of 100 years, represents about the limit of suppression for red
spruce. Many of its associated tree species such as balsam fir and hemlock
may outgrow red spruce after release (22).
Red spruce may be grown successfully using even-age silvicultural
prescriptions (11,12). Red spruce is very shallow-rooted, however, making
it subject to windthrow, a major silvicultural constraint in the
management of the species. As a general rule, it is recommended that no
more than one-fourth to one-half of the basal area be removed in the
partial harvest of a spruce-fir stand, depending on site, to avoid
excessive windthrow damage.
Most of the major forest cover types previously listed in which red
spruce is a component are considered either climax or subclimax.
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Rooting Habit
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İngilizce
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Spruce and fir are shallow-rooted, with most of
the feeding roots in the duff and the top few centimeters of mineral soil
(11). The average rooting depth for all sites in Maine was found to be 33
cm (13 in), with a maximum of 56 cm (22 in) (22).
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Seed Production and Dissemination
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İngilizce
)
Silvics of North America tarafından sağlandı
Good seed crops occur every 3
to 8 years, with light crops during intervening years (22). Red
spruce cones number about 140/liter (5,000/bu), which yields 454 to 680 g
(1.0 to 1.5 lb) of seeds. The number of cleaned seeds per kilogram ranges
between 220,000 and 637,000 (100,000 and 289,000/lb), with an average of
about 306,000 (139,000/lb) (41).
Red spruce seeds fall about 1.2 m (4 ft) per second in still air; the
following formula determines distance of travel for wind- disseminated
spruce seeds at various heights (47):
D = Sh (1.47v)
Where D = distance in feet which seed will travel, S = number of
seconds required for seed to fall from a height of h (ft) on a
tree, and v = velocity of the prevailing wind in miles per hour.
Randall (37), in a study of seed dispersal into clearcut areas,
stated that at a distance of 100 m (5 chains or 330 ft) from the timber
edge, the number of spruce seeds trapped were more than adequate for
regeneration in a good seed year and adequate in an average year. Most of
the spruce in the surrounding stands was red spruce.
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Seedling Development
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İngilizce
)
Silvics of North America tarafından sağlandı
Most red spruce seeds germinate the spring
following dispersal; some, however, may germinate in the fall soon after
dropping from the tree. Germination is epigeal. On favorable seedbeds the
usual spring germination period is from late May to early July. On duff,
which is more subject to surface drying than most other seedbed materials,
some seeds may lose viability by midsummer, and some may show delayed
germination well into August (22). Little if any viable seeds remain in
the forest floor beyond 1 year (13).
Adequate moisture is the chief factor controlling germination of red
spruce. Germination takes place on almost any medium (mineral soil, rotten
wood, or shallow duff) except sod. Mineral soil is an excellent seedbed
for germination. Generally ample moisture is available and soil
temperatures are moderate. Litter and humus are poorer seedbeds because
they are likely to be hotter and drier than mineral soil (11). On thicker
duff, germination may be poor also because moisture conditions are less
favorable. Temperatures of 20° to 30° C (68° to 86° F)
are generally favorable for germination. Seeds will not germinate
satisfactorily at temperatures below 20° C (68° F) and are
permanently injured by long exposure to temperatures higher than 33°
C (92° F) (22).
Germination and initial establishment proceed best under cover.
Seedlings can become established under light intensities as low as 10
percent of full sunlight; however, as they develop, they require light
intensities of 50 percent or more for optimum growth. Seedlings starting
in the open undergo heavy mortality when soil surface temperatures reach
46° to 54° C (115° to 130° F) even for a short time
(11). Drought and frost heaving are major causes of mortality the first
year. Crushing by hardwood litter and snow are also causes of seedling
mortality. Winter drying in some years and locations can cause severe
leader damage and dieback.
Natural reproduction depends more on seedling survival than on
requirements for germination. Spruce seedlings have an exceptionally
slow-growing, fibrous, shallow root system. Consequently, a critical
factor in their survival and establishment is the depth of the 01 organic
layers of the soil profile. When the combined thickness of these layers
exceeds 5 cm (2 in), spruce seedlings may not reach mineral soil and the
moisture necessary to carry them through dry periods. Red spruce seedlings
and the commonly associated balsam fir seedlings are similar in many ways
and are controlled by the same factors, but as a rule spruce is the
weaker, slower growing species during the establishment period (22).
Seedlings that have attained a height of about 15 cm (6 in) can be
considered established. Once established, their early growth is determined
largely by the amount and character of overhead competition. Dense growth
of bracken (Pteridium aquilinum), raspberry, and hardwood sprouts
are the chief competition for seedlings on heavily cutover lands; but red
spruce survives as much as 145 years of suppression and still responds to
release (11,39).
Compared to its associates, red spruce is one of the last species to
start height growth in the spring, usually beginning the first week in
June and ending 9 to 11 weeks later. Radial growth usually begins about
the second week of June and continues through August (22).
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Soils and Topography
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İngilizce
)
Silvics of North America tarafından sağlandı
The soils where red spruce and its associates grow are mostly acid
Spodosols, Inceptisols, and sometimes Histosols with a thick mor humus and
a well-defined A2 horizon- characteristics
commonly associated with abundant rainfall, cool climates, and softwood
cover (11). Commonly, the pH of these soils ranges from 4.0 to 5.5. In
northern New England, red spruce is found predominantly on shallow till
soils that average about 46 cm (18 in) to a compact layer. It will grow on
many sites unfavorable for other species, such as organic soils overlying
rocks in mountainous locations, steep rocky slopes, thin soils, and wet
bottomland (26). On poorly drained soils, lack of aeration limits
growth (22).
In the northern part of its range, red spruce grows at elevations from
near sea level to about 1370 m (4,500 ft) (22). In the southern
Appalachian Mountains it comes in at elevations as low as 1370 m (4,500
ft) and from there to about 1520 m (5,000 ft) it is mixed with hardwoods
and eastern hemlock (Tsuga canadensis). At 1520 m (5,000 ft)
balsam fir (Abies balsamea) joins with red spruce to form the
dominant spruce-fir climax type. In West Virginia, spruce-fir stands are
found as low as 980 m (3,200 ft). Above 1890 m (6,200 ft) in the southern
Appalachians, red spruce appears less frequently than Fraser fir (Abies
fraseri) (47). In the White Mountains of New Hampshire, balsam fir is
the predominant species above 1220 m (4,000 ft) but red spruce is well
represented from about 790 to 1010 m (2,600 to 3,300 ft) (27).
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Special Uses
(
İngilizce
)
Silvics of North America tarafından sağlandı
The wood of red spruce, white spruce (Picea glauca), and black
spruce cannot be distinguished with certainty by either gross
characteristics or minute anatomy, and all three are usually marketed
simply as eastern spruce. Chief uses are for lumber and pulpwood, with
limited amounts going into poles piling, boatbuilding stock, and cooperage
stock (36) Flakeboard and plywood have been made from spruce in recent
years. It is also the preferred wood for piano sounding boards, guitars,
mandolins, organ pipes, and violin bellies (21).
Forest cover types that include red spruce support a wide variety of
wildlife. They are particularly important as winter cover for deer and, to
a certain extent, moose. Small game includes ruffed grouse, snowshoe hare,
and woodcock. Many song birds and fur bearers also frequent these forest
types (44).
A unique use of red spruce was spruce gum, an exudate that accumulates
on trunk wounds. This was the raw material for a flourishing chewing-gum
industry in Maine during the last half of the 19th century and early years
of this century (21).
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Vegetative Reproduction
(
İngilizce
)
Silvics of North America tarafından sağlandı
Red spruce rarely, if ever, layers
(15,22,45). Recently developed techniques facilitate propagation from stem
cuttings under controlled conditions, particularly juvenile cuttings
(7,9,38,45).
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Brief Summary
(
İngilizce
)
Silvics of North America tarafından sağlandı
Pinaceae -- Pine family
Barton M. Blum
Red spruce (Picea rubens), also known as yellow spruce, West
Virginia spruce, eastern spruce, and he-balsam, is one of the more
important conifers in the northeastern United States and adjacent Canada.
It is a medium-size tree that may grow to be more than 400 years old. The
wood of red spruce is light in color and weight, straight grained, and
resilient. It is used for making paper, for construction lumber, and for
musical stringed instruments. Its many uses rival those of eastern white
pine (Pinus strobus) (21).
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Distribution
(
İngilizce
)
Silvics of North America tarafından sağlandı
The range of red spruce extends from the Maritime Provinces of Canada
west to Maine, southern Quebec, and southeastern Ontario, and south into
central New York, eastern Pennsylvania, northern New Jersey, and
Massachusetts. It also grows south along the Appalachian Mountains in
extreme western Maryland, and eastern West Virginia, and north and west in
Virginia, western North Carolina, and eastern Tennessee. Discontinuous
stands may also be found in Haliburton Township, in Algonquin Provincial
Park, and near Sturgeon Falls in Nippising Township, and in the
southwestern Parry Sound District in Ontario, Canada.
- The native range of red spruce.
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Physical Description
(
İngilizce
)
USDA PLANTS text tarafından sağlandı
Tree, Evergreen, Monoecious, Habit erect, Trees without or rarely having knees, Tree with bark rough or scaly, Young shoots 3-dimensional, Buds resinous, Buds not resinous, Leaves needle-like, Leaves alternate, Needle-like leaf margins entire (use magnification), Leaf apex acute, Leaves < 5 cm long, Leaves < 10 cm long, Leaves yellow-green above, Leaves yellow-green below, Leaves not blue-green, Needle-like leaves 4-angled, Needle-like leaves not twisted, Needle-like leaf habit erect, Needle-like leaf habit drooping, Needle-like leaves per fascicle mostly 1, Needle-like leaf sheath early deciduous, Needle-like leaf sheath persistent, Twigs pubescent, Twigs densely pubescent, Twigs viscid, Twigs not viscid, Twigs with peg-like projections or large fascicles after needles fall, Berry-like cones orange, Woody seed cones < 5 cm long, Bracts of seed cone included, Seeds brown, Seeds winged, Seeds unequally winged, Seed wings prominent, Seed wings equal to or broader than body.
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- Damon Little
Qırmızı küknar
(
Azerice
)
wikipedia AZ tarafından sağlandı
Vətəni Şimali Amerika nın şimal-şərqidir. Burada iynəyarpaqlı-enliyarpaqlı məşələrdə, təmiz və ya qarışıq əkinlərdə, bataqlıqlarda və dağlarda bitir. Dağlarda 1000–1800 m-dək qalxır. Avropa da 1875-ci ildən becərilir. Hündürlüyü 20–36 m-ə gövdəsinin diametri 60-135 sm-ə çatan, sıx, konusvarı çətirli alçaqboylu ağacdır. Qabığı yarıqlı, bozumtul və ya qırmızımtıl-qonurdur. Gövdəsi qırmızı-qonur qabıqla örtülmüşdür. Zoğları qırmızı-qonur və ya narıncı, parlaq, hamar, sıx, zərif tükcüklüdür. Tumurcuqların uzunluğu 5 mm-ə çatıb, şabalıdı-qonur, ucu biz-yumurtavarı, bir az qatranlı, üst qabıqları uzunsov, bünövrəsi parlaq qırmızıdır. İynəyarpaqların uzunluğu 10–15 mm, eni 1 mm-ə qədər, parlaq yaşıl, ancaq göyümtül deyil, oraqvarı, yuxarı istiqamətlənmiş, cod iynəli, ucu çox vaxt küt, dördtərəfli, hər iki tərəfdən 1-2 məsaməli xətlidir. Zoğları üstdən çox iynəyarpaqlı, alt tərəfdən isə iynəyarpaqları azdır, sürtüldükdə iynəyarpaqlarından xoş meyvə ətri gəlir. Qozaları qısa zoğlarda tək sallanır, bütövlükdə çoxsaylı, uzunluğu 3-5 sm-ə, qalınlığı 1,5-2 sm-ə çatan, yumurtavarı-oval, qatranlı, əvvəlcə yaşılımtıl-bənövşəyi, sonradan qırmızı-qonur rəng alır. Toxum qabıqlarının uzunluğu 13 mm-ə qədər, yumurtavarı, yumru, qabarıq, cod, zolaqlı, qısa tükcüklü, kənarları bütöv, bəzən zəif, xırda dişli, içəridən tünd-qırmızıdır. Kənar qabıqları çox xırda, uzunsov-yumrudur. Toxumların uzunluğu 3 mm-ə qədər olub, qaramtıl və ya boz-qonur rənglidir. 350-400 (450-500) ilədək yaşayır.
Çox dekorativdir. Tək əkinlərdə istifadəsi məqsədəuyğundur. Yuxarı zoğlarını sıx örtən tünd-yaşıl iynəyarpaqların fonunda çoxsaylı, qırmızı, xırda qozaları və qırmızı-qonur gövdəsi ağaca xüsusi bəzək verir. Abşeron da, Qəbələ də yaşıllaşdırmada rast gəlinir.
Məlumat mənbələri:
Деревья и кустарники СССР. т.3.1954; Флора Азербайджана. т.5. 1954; Azərbaycanın ağac və kolları. III cild. 1970; Azərbaycanın “Qırmızı” və “Yaşıl Кitabları”na tövsiyə olunan bitki və bitki formasiyaları. 1996; Azərbaycan florasının konspekti. I-III cildlər. 2005; 2006; 2008.
İstinadlar
- “Azərbaycan dendraflorasi” I cild, Baki, “Elm”, 2011, 312 səh.
Mənbə
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Qırmızı küknar: Brief Summary
(
Azerice
)
wikipedia AZ tarafından sağlandı
Vətəni Şimali Amerika nın şimal-şərqidir. Burada iynəyarpaqlı-enliyarpaqlı məşələrdə, təmiz və ya qarışıq əkinlərdə, bataqlıqlarda və dağlarda bitir. Dağlarda 1000–1800 m-dək qalxır. Avropa da 1875-ci ildən becərilir. Hündürlüyü 20–36 m-ə gövdəsinin diametri 60-135 sm-ə çatan, sıx, konusvarı çətirli alçaqboylu ağacdır. Qabığı yarıqlı, bozumtul və ya qırmızımtıl-qonurdur. Gövdəsi qırmızı-qonur qabıqla örtülmüşdür. Zoğları qırmızı-qonur və ya narıncı, parlaq, hamar, sıx, zərif tükcüklüdür. Tumurcuqların uzunluğu 5 mm-ə çatıb, şabalıdı-qonur, ucu biz-yumurtavarı, bir az qatranlı, üst qabıqları uzunsov, bünövrəsi parlaq qırmızıdır. İynəyarpaqların uzunluğu 10–15 mm, eni 1 mm-ə qədər, parlaq yaşıl, ancaq göyümtül deyil, oraqvarı, yuxarı istiqamətlənmiş, cod iynəli, ucu çox vaxt küt, dördtərəfli, hər iki tərəfdən 1-2 məsaməli xətlidir. Zoğları üstdən çox iynəyarpaqlı, alt tərəfdən isə iynəyarpaqları azdır, sürtüldükdə iynəyarpaqlarından xoş meyvə ətri gəlir. Qozaları qısa zoğlarda tək sallanır, bütövlükdə çoxsaylı, uzunluğu 3-5 sm-ə, qalınlığı 1,5-2 sm-ə çatan, yumurtavarı-oval, qatranlı, əvvəlcə yaşılımtıl-bənövşəyi, sonradan qırmızı-qonur rəng alır. Toxum qabıqlarının uzunluğu 13 mm-ə qədər, yumurtavarı, yumru, qabarıq, cod, zolaqlı, qısa tükcüklü, kənarları bütöv, bəzən zəif, xırda dişli, içəridən tünd-qırmızıdır. Kənar qabıqları çox xırda, uzunsov-yumrudur. Toxumların uzunluğu 3 mm-ə qədər olub, qaramtıl və ya boz-qonur rənglidir. 350-400 (450-500) ilədək yaşayır.
Çox dekorativdir. Tək əkinlərdə istifadəsi məqsədəuyğundur. Yuxarı zoğlarını sıx örtən tünd-yaşıl iynəyarpaqların fonunda çoxsaylı, qırmızı, xırda qozaları və qırmızı-qonur gövdəsi ağaca xüsusi bəzək verir. Abşeron da, Qəbələ də yaşıllaşdırmada rast gəlinir.
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Smrk červený
(
Çekçe
)
wikipedia CZ tarafından sağlandı
Smrk červený (Picea rubens) je druh smrku, původem ze Severní Ameriky.
Popis
Smrk červený se samičími šiškami
Jedná se strom dorůstající ve své domovině až 40 m a tloušťky kmene až 1 m v průměru.[2] Může se dožít i více než 400 let.[3] Koruna je úzce kuželovitá, tvar koruny však může být pozměněn díky stanovištním podmínkám (vítr aj.). Borka je šedohnědá až červenohnědá, pupeny jsou červenohnědé.[2] Jehlice jsou na průřezu čtyřhranné, 0,8-2,3 cm (zřídka až 3 cm dlouhé), žlutozelené až tmavě zelené, ale nikoliv sivé.[2] Samičí šišky jsou celkem malé, 2,3-4,5 (zřídka až 5) cm dlouhé, vehčitého tvaru, lesklé, za zralosti oranžovo hnědé Šupiny jsou vějířovitého tvaru, na vrcholu celokrajné nebo nepravidelně zubaté, cca 8-12 mm dlouhé a asi stejně široké. Počet chromozómů je 2n=24.[2]
Rozšíření
Původní areál druhu se rozkládá v jihovýchodní Kanadě a v severovýchodní části USA, jižněji pak jen v Appalačském pohoří.[2][3]
Kříženci
Z volné přírody je znám kříženec s druhem Picea mariana, dochází zde na kontaktu areálů k introgresivní hybridizaci.[2]
Ekologie
Smrk červený roste od mořského pobřeží v Kanadě až po nadmořskou výšku cca 1500 m n. m. v Appalačském pohoří. Roste v oblasti s chladným vlhkým oceanickým klimatem s ročním úhrnem srážek asi 875-2000 mm. Nejčastěji tvoří společenstva se smrkem sivým (Picea glauca) a jedlí balzámovou (Abies balsamea), méně často se smrkem černým (Picea mariana), který roste víc ve vlhku, lokálně také s jedlí Fraserovou (Abies fraseri) či cypřiškem zeravovitým (Chamaecyparis thyoides). Na lepších půdách se přidávají i listnaté dřeviny. Hlavně ve 20. století postihlo některé porosty hromadné hynutí díky kyselým dešťům.[4][5]
Výskyt v České republice
V České republice nepatří k příliš rozšířeným druhům smrků, roste např. v některých arboretech.[4][5]
Odkazy
Reference
-
↑ Červený seznam IUCN 2018.1. 5. července 2018. Dostupné online. [cit. 2018-08-10]
-
↑ a b c d e f Flóra Severní Ameriky [online]. Dostupné online. (anglicky)
-
↑ a b The Gymnosperm Database [online]. Dostupné online. (anglicky)
-
↑ a b Dendrologie online [online]. Dostupné online. (česky)
-
↑ a b Řícmanice [online]. Dostupné online. (česky)
Externí odkazy
Rod smrk (Picea) Evropa
Asie
Amerika
Smrk červený: Brief Summary
(
Çekçe
)
wikipedia CZ tarafından sağlandı
Smrk červený (Picea rubens) je druh smrku, původem ze Severní Ameriky.
Amerikanische Rot-Fichte
(
Almanca
)
wikipedia DE tarafından sağlandı
Die Amerikanische Rot-Fichte (Picea rubens) ist eine Pflanzenart der Gattung Fichten (Picea) in der Familie der Kieferngewächse (Pinaceae).
Beschreibung
Die Amerikanische Rot-Fichte ist ein immergrüner Baum, der Wuchshöhen von bis zu 40 Metern und bis zu 100 cm Stammumfang erreicht. Im Nordosten ihres Verbreitungsgebietes erreicht sie jedoch nur etwa 25 Meter Wuchshöhe und 60 cm Stammumfang.[1] Das höchste bekannte Exemplar mit 46 Meter Höhe steht im Great-Smoky-Mountains-Nationalpark.[2] Die Rinde des Stammes ist kräftig purpurbraun bis dunkelgrau. Sie reißt feinschuppig auf und löst sich bei älteren Bäumen in kleinen konkaven Platten ab. Die Baumkrone ist schmal kegelförmig und lang zugespitzt. Die Äste stehen mehr oder weniger waagrecht ab; die unteren Äste sind etwas abwärts gekrümmt und steigen zu den Spitzen hin wieder an. Die Zweige sind gelbbraun, hellorange oder rotbraun; sie sind teils dicht behaart, teils nackt mit wenigen Haaren in den Furchen. Die eiförmigen Knospen sind rotbraun und 5 bis 8 Millimeter groß. Die Nadeln sind dünn, bis 1 mm breit und 0,8 bis 2,5 cm lang; sie enden spitz. Im ersten Jahr sind die Nadeln grasgrün, später tiefgrün und glänzend. Die zerriebenen Nadeln duften nach Kerzenwachs oder Äpfeln.
Die männlichen Blüten sind hängend und hellrot. Die weiblichen Zapfen sind länglich eiförmig und stehen oft in Büscheln. Sie sind etwa 2,3 bis 4,5 cm lang.
Die Chromosomenzahl ist 2n=24.
Amerikanische Rot-Fichte (
Picea rubens), Zweig mit Zapfen
Verbreitung und Standort
Karte des Verbreitungsgebiets
Die Amerikanische Rot-Fichte ist im östlichen Nordamerika heimisch; ihr Verbreitungsgebiet erstreckt sich im Wesentlichen vom küstennahen östlichen Kanada in südwestlicher Richtung bis in die Appalachen. In Kanada kommt sie in Neuschottland, Neubraunschweig, dem südlichen Quebec und dem südöstlichen Ontario vor; auch auf der Prinz-Edward-Insel und den französisch kontrollierten Inseln Saint-Pierre und Miquelon ist sie verbreitet. In den östlichen USA reichen ihre Vorkommen in die Bundesstaaten Maine, New Hampshire, Vermont, Massachusetts, Connecticut, New York, Pennsylvania, New Jersey, Maryland, Virginia, West Virginia, North Carolina und Tennessee.
Diese Art besiedelt Höhenlagen von 0 bis 2000 m. Sie bevorzugt ein kühl-feuchtes Klima.
In Deutschland wird die Amerikanische Rot-Fichte nur selten gepflanzt.
Nutzung
Die langen Wurzeln wurden früher von einheimischen Indianern ausgegraben, abgeschält und zum Flechten verwendet. Das Harzpech wurde zum Ausstopfen von Löchern verwendet. In Maine wurde in der letzten Hälfte des 19. Jahrhunderts bis zum Anfang des 20. Jahrhunderts aus dem Harz Kaugummi gewonnen. Siedler benutzten die jungen grünen Nadeln zur Aromatisierung beim Bierbrauen.
Das Holz ist hell und von geringer Dichte. Es wird als Bauholz, in der Papierherstellung und bei der Herstellung von Streich- und Zupfinstrumenten verwendet.
Systematik
Der deutsche Botaniker Otto von Münchhausen beschrieb die Amerikanische Rot-Fichte unter dem Taxon Pinus acutissima 1770 in seinem sechsbändigen Werk Der Hausvater (Band 5(1), Seite 225).[3] Die heute gültige Beschreibung der Art als Picea rubens in der Gattung Picea durch den amerikanischen Botaniker Charles Sprague Sargent wurde 1898 veröffentlicht (Silva 12: 33, t. 597, 1898).[3]
Weitere Synonyme für die Art sind:
-
Picea australis Small 1903
-
Picea nigra (Aiton) Link var. rubra (Du Roi) Engelmann
-
Picea rubra (Du Roi) Link 1831 non A.Dietrich 1824.
Sonstiges
Die Bestände der Amerikanischen Rot-Fichte in den Appalachen, die bereits durch Waldbrände und Rodungen auf ein Bruchteil ihrer ursprünglichen Bestände geschrumpft waren,[1] leiden unter durch Luftverschmutzung verursachtem saurem Regen.[2]
Das älteste bekannte Exemplar der Amerikanischen Rot-Fichte war etwa 445 Jahre alt.[2]
Die Amerikanische Rot-Fichte ist der Provinzbaum der kanadischen Provinz Neuschottland.
Quellen
- Alan Mitchell, übersetzt und bearbeitet von Gerd Krüssmann: Die Wald- und Parkbäume Europas: Ein Bestimmungsbuch für Dendrologen und Naturfreunde. Paul Parey, Hamburg und Berlin 1975, ISBN 3-490-05918-2.
-
Christopher J. Earle: Picea rubens. In: The Gymnosperm Database. 20. Januar 2011, abgerufen am 8. November 2011 (englisch).
-
Barton M. Blum: Red Spruce. USDA Forest Service; Northeastern Area State & Private Forestry, abgerufen am 8. November 2011 (englisch).
Einzelnachweise
-
↑ a b Siehe Barton M. Blum.
-
↑ a b c Siehe Eintrag in The Gymnosperm Database.
-
↑ a b Rafaël Govaerts (Hrsg.): Picea. In: World Checklist of Selected Plant Families (WCSP) – The Board of Trustees of the Royal Botanic Gardens, Kew, abgerufen am 14. April 2019.
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Amerikanische Rot-Fichte: Brief Summary
(
Almanca
)
wikipedia DE tarafından sağlandı
Die Amerikanische Rot-Fichte (Picea rubens) ist eine Pflanzenart der Gattung Fichten (Picea) in der Familie der Kieferngewächse (Pinaceae).
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Amerikanesch Routfiicht
(
Lüksemburgca
)
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Amerikanesch Routfiicht: Brief Summary
(
Lüksemburgca
)
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D'Amerikanesch Routfiicht (Picea rubens) ass e Bam aus der Gattung vu de Fiichten (Picea) an der Famill vun de Pinaceae. De Bam gëtt ongeféier 27 Meter grouss an huet eng kräfteg purpurbrong Schuel.
D'Verbreedungsgebitt vun der Aart läit an Neischottland am Oste vu Kanada, an am Nordoste vun den USA.
Se gouf 1898 vum US-amerikanesche Biolog Charles Sprague Sargent beschriwwen.
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Picea rubens
(
İngilizce
)
wikipedia EN tarafından sağlandı
Picea rubens, commonly known as red spruce, is a species of spruce native to eastern North America, ranging from eastern Quebec and Nova Scotia, west to the Adirondack Mountains and south through New England along the Appalachians to western North Carolina.[3][4][5] This species is also known as yellow spruce, West Virginia spruce, eastern spruce, and he-balsam.[6][7] Red spruce is the provincial tree of Nova Scotia.[4]
Description
Red spruce is a perennial,[8] shade-tolerant, late successional[9] coniferous tree that under optimal conditions grows to 18–40 m (59–131 ft) tall with a trunk diameter of about 60 cm (24 in), though exceptional specimens can reach 46 m (151 ft) tall and 100 cm (39 inches) in diameter. It has a narrow conical crown. The leaves are needle-like, yellow-green, 12–15 mm (15⁄32–19⁄32 in) long, four-sided, curved, with a sharp point, and extend from all sides of the twig. The bark is gray-brown on the surface and red-brown on the inside, thin, and scaly. The wood is light, soft, has narrow rings, and has a slight red tinge.[10] The cones are cylindrical, 3–5 cm (1+1⁄4–2 in) long, with a glossy red-brown color and stiff scales. The cones hang down from branches.[3][4][5][11]
Habitat
Red spruce grows at a slow to moderate rate, lives for 250 to 450+ years, and is very shade-tolerant when young.[12] It is often found in pure stands or forests mixed with eastern white pine, balsam fir, or black spruce. Along with Fraser fir, red spruce is one of two primary tree types in the southern Appalachian spruce-fir forest, a distinct ecosystem found only in the highest elevations of the Southern Appalachian Mountains.[13] Its habitat is moist but well-drained sandy loam, often at high altitudes. Red spruce can be easily damaged by windthrow and acid rain.
Notable red spruce forests can be seen at Gaudineer Scenic Area, a virgin red spruce forest located in West Virginia, the Canaan Valley, Roaring Plains West Wilderness, Dolly Sods Wilderness, Spruce Mountain and Spruce Knob all also in West Virginia and all sites of former extensive red spruce forest. Some areas of this forest, particularly in Roaring Plains West Wilderness, Dolly Sods Wilderness as well as areas of Spruce Mountain are making a rather substantial recovery.
Related species
It is closely related to black spruce, and hybrids between the two are frequent where their ranges meet.[3][4][5] Genetic data suggests that the red spruce peripatrically speciated from the black spruce during the Pleistocene due to glaciation.[14][15]
Uses
Red spruce is used for Christmas trees and is an important wood used in making paper pulp. It is also an excellent tonewood, and is used in many higher-end acoustic guitars and violins, as well as musical soundboard. The sap can be used to make spruce gum.[11] Leafy red spruce twigs are boiled with sugar and flavoring to make spruce beer.[16] Also it can be made into spruce pudding. It can also be used as construction lumber and is good for millwork and for crates.[17]
Red spruce cones from the Pisgah National Forest
Damaging factors
Like most trees, red spruce is subject to insect parasitism. Their insect enemy is the spruce budworm, although it is a bigger problem for white spruce and balsam fir.[18] Other issues that have been damaging red spruce has been the increase in acid rain and current climate change.[19]
One of the consequences of acid rain deposition is the decrease of soil exchangeable calcium and increase of aluminum. This is because acid precipitation disrupts cation and nutrition cycling in forest ecosystems. Components of acid rain such as H+, NO3−, and SO42- limit the uptake of calcium by trees and can increase aluminum availability.[20]
Calcium concentration is important for red spruce for physiological processes such as dark respiration and cold tolerance, as well as disease resistance, signal transduction, membrane and cell wall synthesis and function, and regulation of stomata. Conversely, dissolved aluminum can be toxic or can interfere with root uptake of calcium and other nutrients. At the ecosystem and community levels, Calcium availability is associated with community composition, mature tree growth, and ecosystem productivity. One study testing the effects of added aluminum to soil, found that P. rubens mortality rate increased under these conditions.[21]
During the 1980s, increased acid deposition contributed to a loss of high-elevation red spruce trees, due to leached calcium and decreased freezing tolerance.[22] Additionally, the structure of the spruce needle enhances the capture of water and particles, which has been shown to add to soil acidification, nutrient leaching, and forest decline.[23]
However, more recently, reductions in acid deposition have contributed to red spruce resurgence in some mountain areas in the northeastern United States. This increase in red spruce growth has been associated with an increase in rainfall pH, which reduces bulk acidic deposition. This suggests that policies aiming to reduce atmospheric pollution in this area have been effective, although other species sensitive to soil acidification, such as sugar maple, are still continuing to decline.[22]
Conservation
The Central Appalachian Spruce Restoration Initiative (CASRI)[24] seeks to unite diverse partners with the goal of restoring historic red spruce ecosystems across the high-elevation landscapes of central Appalachians. The partners that make up this diverse group are Appalachian Mountain Joint Venture, Appalachian Regional Reforestation Initiative, Canaan Valley National Wildlife Refuge, Natural Resources Conservation Service, The Mountain Institute, The Nature Conservancy, Trout Unlimited, U.S. Forest Service Northern Research Station, U.S. Forest Service Monongahela National Forest, West Virginia Division of Natural Resources, West Virginia Division of Forestry, West Virginia Highlands Conservancy, West Virginia State Parks, and West Virginia University.[25]
Prior to the late 1800s, 600,000 hectares (1,500,000 acres) of red spruce were in West Virginia. In the late 1800s and early 1900s, a vast amount of logging began in the state and the number of red spruce dwindled to 12,000 hectares (30,000 acres). Silviculture is being used to help restore the population of the lost red spruce.[26]
Significant efforts have been made to increase the growth of red spruce trees in western North Carolina. Most notably by Molly Tartt on behalf of the Daughters of the American Revolution. Tartt, a resident of Brevard North Carolina, embarked on a mission to find the lost red spruce Pisgah Forest that had been planted by the DAR as a memorial to the lives lost during the American Revolution. The forest, consisting of 50,000 trees was dedicated in 1940 and had until recently been forgotten until Tartt located and identified the forest.[27]
References
-
^ Farjon, A. (2013). "Picea rubens". IUCN Red List of Threatened Species. 2013: e.T42335A2973542. doi:10.2305/IUCN.UK.2013-1.RLTS.T42335A2973542.en. Retrieved 19 November 2021.
-
^ "Picea rubens Sarg.". World Checklist of Selected Plant Families. Royal Botanic Gardens, Kew – via The Plant List. Note that this website has been superseded by World Flora Online
-
^ a b c Farjon, A. (1990). Pinaceae. Drawings and Descriptions of the Genera. Koeltz Scientific Books ISBN 3-87429-298-3.
-
^ a b c d Taylor, Ronald J. (1993). "Picea rubens". In Flora of North America Editorial Committee (ed.). Flora of North America North of Mexico (FNA). Vol. 2. New York and Oxford – via eFloras.org, Missouri Botanical Garden, St. Louis, MO & Harvard University Herbaria, Cambridge, MA.
-
^ a b c Earle, Christopher J., ed. (2018). "Picea rubens". The Gymnosperm Database.
-
^ Blum, Barton M. (1990). "Picea rubens". In Burns, Russell M.; Honkala, Barbara H. (eds.). Conifers. Silvics of North America. Washington, D.C.: United States Forest Service (USFS), United States Department of Agriculture (USDA). Vol. 1 – via Southern Research Station.
-
^ Peattie, Donald Culross (1948-01-01). A Natural History of Trees of Eastern and Central North America. Houghton Mifflin Harcourt. p. 51. ISBN 0-395-58174-5.
-
^ "Red Spruce (Rubens)". Garden Guides. Retrieved 27 February 2014.
-
^ Dumais, D; Prevost, M (June 2007). "Management for red spruce conservation in Quebec: The importance of some physiological and ecological characteristics – A review". Forestry Chronicle. 83 (3): 378–392. doi:10.5558/tfc83378-3. ProQuest 294760995.
-
^ "Red Spruce" (PDF). USDA NRCS. Retrieved 26 February 2014.
-
^ a b "Red Spruce". A handbook of Maritime trees. Atlantic Forestry Centre. Archived from the original on 2008-08-18.
-
^ "Eastern OLDLIST: A database of maximum tree ages for Eastern North America".
-
^ White, Peter (2006). "Boreal Forest". Encyclopedia of Appalachia. University of Tennessee Press. pp. 49–50.
-
^ Juan P. Jaramillo-Correa & Jean Bousquet (2003), "New evidence from mitochondrial DNA of a progenitor-derivative species relationship between black and red spruce (Pinaceae)", American Journal of Botany, 90 (12): 1801–1806, doi:10.3732/ajb.90.12.1801, PMID 21653356
-
^ Isabelle Gamache, Juan P. Jaramillo-Correa, Sergey Payette, & Jean Bousquet (2003), "Diverging patterns of mitochondrial and nuclear DNA diversity in subarctic black spruce: imprint of a founder effect associated with postglacial colonization", Molecular Ecology, 12 (4): 891–901, doi:10.1046/j.1365-294x.2003.01800.x, PMID 12753210, S2CID 20234158
{{citation}}
: CS1 maint: multiple names: authors list (link) -
^ Little, Elbert L. (1980). The Audubon Society Field Guide to North American Trees: Eastern Region. New York: Knopf. p. 285. ISBN 0-394-50760-6.
-
^ "Red Spruce". The Wood Database. Retrieved 27 February 2014.
-
^ "Red Spruce" at the Encyclopedia of Life
-
^ Houle, Daniel (2012). "Compositional vegetation changes and increased red spruce abundance during the Little Ice Age in a sugar maple forest of north-eastern North America". Plant Ecology. 213 (6): 1027–1035. doi:10.1007/s11258-012-0062-0. S2CID 15104515.
-
^ Schaberg, P. G.; DeHayes, D. H.; Hawley, G. J.; Strimbeck, G. R.; Cumming, J. R.; Murakami, P. F.; Borer, C. H. (2000-01-01). "Acid mist and soil Ca and Al alter the mineral nutrition and physiology of red spruce". Tree Physiology. 20 (2): 73–85. doi:10.1093/treephys/20.2.73. ISSN 0829-318X. PMID 12651475.
-
^ Kobe, Richard K; Likens, Gene E; Eagar, Christopher (2002-06-01). "Tree seedling growth and mortality responses to manipulations of calcium and aluminum in a northern hardwood forest". Canadian Journal of Forest Research. 32 (6): 954–966. doi:10.1139/x02-018. ISSN 0045-5067.
-
^ a b Wason, Jay W.; Beier, Colin M.; Battles, John J.; Dovciak, Martin (2019). "Acidic Deposition and Climate Warming as Drivers of Tree Growth in High-Elevation Spruce-Fir Forests of the Northeastern US". Frontiers in Forests and Global Change. 2. doi:10.3389/ffgc.2019.00063. ISSN 2624-893X.
-
^ Pierret, Marie-Claire; Viville, Daniel; Dambrine, Etienne; Cotel, Solenn; Probst, Anne (2019-04-01). "Twenty-five year record of chemicals in open field precipitation and throughfall from a medium-altitude forest catchment (Strengbach - NE France): An obvious response to atmospheric pollution trends". Atmospheric Environment. 202: 296–314. Bibcode:2019AtmEn.202..296P. doi:10.1016/j.atmosenv.2018.12.026. ISSN 1352-2310.
-
^ Burks, Evan (Dec 2010). "Return of the Red Spruce". Wonderful West Virginia. 74 (12): 6–11.
-
^ Bove, Jennifer. "Appalachian Red Spruce Forest". Retrieved 27 February 2014.
-
^ Rentch, James; T. Schuler; M. Ford; G. Nowacki (September 2007). "Red Spruce Stand Dynamics, Simulations, and Restoration Opportunities in the Central Appalachians". Restoration Ecology. 15 (3): 440–452. doi:10.1111/j.1526-100x.2007.00240.x. S2CID 85913515. ProQuest 289371889.
-
^ "Daughters of the American Revolution brings forgotten forest to light".
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Picea rubens: Brief Summary
(
İngilizce
)
wikipedia EN tarafından sağlandı
Picea rubens, commonly known as red spruce, is a species of spruce native to eastern North America, ranging from eastern Quebec and Nova Scotia, west to the Adirondack Mountains and south through New England along the Appalachians to western North Carolina. This species is also known as yellow spruce, West Virginia spruce, eastern spruce, and he-balsam. Red spruce is the provincial tree of Nova Scotia.
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Picea rubens
(
İspanyolca; Kastilyaca
)
wikipedia ES tarafından sağlandı
Picea rubens, la pícea roja, es una especie de conífera perteneciente a la familia Pinaceae nativa del este de Norteamérica, desde Quebec a Nueva Escocia, y desde el sur de Nueva Inglaterra en las montañas Adirondack y los Apalaches al oeste de Carolina del Norte.
Descripción
Alcanza un tamaño de 18-30 metros de altura y un tronco con 60 cm de diámetro, aunque algunos ejemplares llegan a 46 m y 130 cm de diámetro. Tiene una estrecha corona cónica. crece lentamente y vive de 250 a 350 años. Las hojas son como agujas, rígidas, erectas amarillo-verdosa de 12-15 mm de longitud. Tiene una piña cilíndrica de 3-5 cm de longitud, siendo de color rojo-marrón y cuelgan de las ramas.
Usos
Se utiliza comercialmente como árbol de Navidad y por su pulpa para hacer papel o para la construcción de guitarras.
Taxonomía
Picea rubens fue descrita por Charles Sprague Sargent y publicado en The Silva of North America 12: 33, pl. 597. 1898.[2]
- Etimología
Picea; nombre genérico que es tomado directamente del Latín pix = "brea", nombre clásico dado a un pino que producía esta sustancia[3]
rubens: epíteto latíno que significa "de color rojo".[4]
- Sinonimia
-
Abies americana K.Koch
-
Abies rubra (Du Roi) Poir.
-
Picea americana Suringar
-
Picea australis Small
-
Picea nigra var. rubra (Du Roi) Engelm.
-
Picea rubra (Du Roi) Link
-
Picea rubra var. pendula Carrière
-
Picea rubra var. pusilla Peck
-
Picea rubra var. virgata (Rehder) Rehder
-
Picea rubra f. virgata Rehder
-
Pinus abies var. acutissima Münchh.
-
Pinus americana Gaertn.
-
Pinus canadensis var. rubescens Weston
-
Pinus mariana var. rubra Du Roi
-
Pinus rubra (Du Roi) D.Don
-
Pinus rubra var. violacea Endl.[5]
Referencias
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Picea rubens: Brief Summary
(
İspanyolca; Kastilyaca
)
wikipedia ES tarafından sağlandı
Picea rubens, la pícea roja, es una especie de conífera perteneciente a la familia Pinaceae nativa del este de Norteamérica, desde Quebec a Nueva Escocia, y desde el sur de Nueva Inglaterra en las montañas Adirondack y los Apalaches al oeste de Carolina del Norte.
Vista del árbol
Detalle de las hojas
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Punane kuusk
(
Estonyaca
)
wikipedia ET tarafından sağlandı
Punane kuusk (Picea rubens) on igihaljas okaspuuliik kuuse perekonnast. Selle looduslik levila Põhja-Ameerika kirdeosas, kuid 18. sajandi keskel toodi seda ka Euroopasse ning vähesel määral kasvatatakse seda ka Soomes ja Eestis.
Kirjeldus
Puu kasvab keskmiselt 18–23, maksimaalselt 35 m kõrguseks.[3] Puu eluiga võib soodsates oludes küündida kuni 400 aastani.[4]
Tüve keskmine läbimõõt on 30–60, maksimaalne 130 cm.[4] Tüve koor on hallikas või punakaspruun, vanematel puudel pikisuunas lõhenev. Võra on laikuhikja kujuga.[5]
Juurestik on maapinnalähedane, üsna aeglaselt arenev, ulatub keskmiselt 35 (maksimaalselt 55) cm sügavusele, mistõttu esineb tihti tormiheidet.[3]
Võrsed on punakaspruunid, karvased, peened.[6]
Okkad on 1,0–1,5 cm pikkused, teravad, läikivad, tume- või helerohelised, tipust pisut kõverad, neljatahulised, pealmiste külgede mõlemal tahul 6, alumistel tahkudel 3 õhulõherida, hõõrumisel lõhnavad aromaatselt[6]. Okkad katavad võrset igast küljest.
Puukoor on pealtpoolt hallikaspruun ja seestpoolt punakaspruun, õhuke ja soomuseline.
Pungad on munajad, nõrgalt vaigused, tumepruunid, punakaspruunide pikenenud pungasoomustega [5]. Käbid ripuvad okstelt allapoole.
Käbid on 3–4 (6) cm pikkused ja läbimõõduga kuni 2,0 cm, vaigused, läikivad, valminult punakaspruunid. Seemnesoomused on väga jäigad, kumerad, pehmekarvalised, terveservalised või veidi hambulise servaga.[6] Seemned on 2–3 mm pikkused, tumepruunid, helepruunide 6–9 mm pikkuste tiivakestega.[5] 1000 seemne mass on vahemikus 1,6–4,5 g[4].
Levila ja ökoloogia
Punase kuuse levila paikneb Põhja-Ameerika kirdeosas võrdlemisi väiksel alal. Selle põhjaosas kasvab punane kuusk 0–1370 m kõrgusel merepinnast, Apalatšide lõunaosas 1370–1520 m kõrgusel segus lehtpuude ja kanada tsuugaga, 1520 meetrist alates peamiselt koos palsamnuluga. Kõrgemal kui 1900 m asendab teda puistus järk-järgult Fraseri nulg.[4]
Punase kuuse puistud on suhteliselt tundlikud tormiheite ja happevihmade suhtes.[viide?]
Kliima ja kasvupinnas
Punase kuuse areaali kliima on niiske ja jahe. Aasta keskmine sademete hulk on 910–1320 mm. Lund sajab aastas keskmiselt 200–400 cm ning lumikate esineb aastas 100–140 päeval. Jaanuari keskmised minimaalsed õhutemperatuurid on vahemikus –13...–18 °C, maksimaalsed aga –1...–7 °C. Puu talub talvel külma kuni –30...–40 °C[7]. Külmavaba perioodi pikkus aastas on 90–150 ööpäeva. Juuli keskmised maksimaalsed õhutemperatuurid on +21...+27 °C, minimaalsed +11...+14 °C. Parimat kasvu näitavad Apalatšides kasvavad kuusikud, kuna suhteline õhuniiskus ja sademete hulk on seal kõrgemad.[4]
Punane kuusk kasvab peamiselt viljakatel happelistel muldadel, mille pH on vahemikus 4,0–5,5. Samuti kasvab nii soodes kui ka õhukese mullakihiga kaljudel, mägedes tõuseb kuni 2000 m kõrgusele merepinnast. Soostunud aladel jääb kasv kiduraks.[4]
Kaasliigid
Punane kuusk kasvab kas puhtpuistuna või segamini koos järgmiste puuliikidega: palsamnulg (Abies balsamea), must kuusk (Picea mariana), ameerika haab (Populus tremuloides), pensilvaania kirsipuu (Prunus pensylvanica), paberikask (Betula papyrifera), valge mänd (Pinus strobus), kanada tsuuga (Tsuga canadensis), kollane kask (Betula alleghaniensis), suhkruvaher (Acer saccharum), Fraseri nulg (Abies fraseri), harilik elupuu (Thuja occidentalis), punane vaher (Acer rubrum) jt.[4]
Punane ja must kuusk on lähedased sugulased ja kohtades, kus nende levila kattub, esineb tihti nende omavahelisi hübriide.[4]
Punane kuusk toiduallikana
Kuusepüüd toituvad punase kuuse võrsetest ja okastest
Ameerika suurnokk-vint toitub punase kuuse seemnetest ja aitab neid levitada
Kuusepüüd (Falcipennis canadensis) toituvad punase kuuse võrsetest ja okastest. Hiired koguvad suurel hulgal mahalangenud punase kuuse seemneid, eelistades punase kuuse seemneid palsamnulu seemnetele. Linnud, eriti kuuse-käbilind (Loxia curvirostra) ja ameerika suurnokk-vint (Hesperiphona vespertina), ursonid (Erethizon dorsatum), baribalid (Ursus americanus) ja ameerika jänesed (Lepus americanus) toituvad noorte kuuskede pungadest.[3]
Paljunemine ja kasv
Punane kuusk on ühekojaline taim, paljuneb seemnetega, mida levitavad peamiselt tuuled. Vähemal määral aitavad kaasa väikenärilised ja linnud. Viljakandvus algab valgusküllases kasvukohas 15–20 aasta vanuselt ja saavutab oma maksimumi umbes 15 aastat hiljem. Tihedamas ühevanuses puistus aga hakkavad puud reeglina käbisid kandma alles 40–50 aasta vanuselt. Õitseb mais, isasõisikud on erepunased, emasõisikud on ererohelised, asetsevad eelmise aasta võrsete tipul ja on õitsemise ajal püsti. Seemned valmivad sama aasta septembri keskelt kuni oktoobri alguseni. Käbid avanevad peatselt pärast seemnete valmimist. Seemned kanduvad tuulega üldjuhul kuni 60 m kaugusele. Head seemneaastad korduvad 3–8 aasta järel. Esimesel eluaastal hukkuvad noored seemikud peamiselt põua ja külmakahjustuste tõttu. Ellujäämisprotsent on suurem õhematel ja väheviljakatel muldadel, kus kiiremakasvulistele lehtpuudele eriti kasvada ei meeldi.[3]
Kuigi punase kuuse seemikud eelistavad noores eas varjus kasvamist, muutub edaspidi valgus väga oluliseks. Teiste puude varjus kasvav puu võib ka 145 aasta vanusena soodsatesse valgustingimustesse sattudes oluliselt kiiremini kasvama hakata. Täisvarjus kasvav puu võib 50 aasta vanuselt olla vaid 1,2–1,5 m kõrgune. Soodsates tingimustes kasvav puu võib kasvada 20-aastaselt kuni 7 ja 60-aastaselt 19 meetri kõrguseks.[4]
Kasutamine
Punane kuusk on USA kirdeosas tähtis majanduslikult kasutatav puuliik. Puit on helepruun või kreemikas, kergelt töödeldav, elastne, keskmiste tugevusomadustega, paralleelsete kiududega. Eriti hinnatud on punase kuuse puit muusikainstrumentide puitosade valmistamisel (nt klaverid, kitarrid, mandoliinid ja viiulid). Puitu kasutatakse veel ehitusmaterjalina, vineeritööstuses, paberitööstuses tselluloosi saamiseks, jõulupuuna jne.[3]
Punase kuuse puit on väärtuslik tooraine muusikainstrumentide valmistamisel
Puidu füüsikalised ja mehaanilised omadused on esitatud tabelis.[8][9]
Omadus Väärtus Ühik
Tihedus, õhukuiv puit* 448 kg/m
3 Erikaal, õhukuiv puit* 0,40 -
Elastsusmoodul, värske puit / õhukuiv puit* 9200 / 11400 MPa
Paindetugevus, värske puit / õhukuiv puit* 41,0 / 74,0 MPa
Survetugevus, (õhukuiv puit*) pikikiudu / ristikiudu 38,2 / 3,8 MPa
Nihketugevus, värske puit / õhukuiv puit* 5,2 / 8,9 MPa
Ruumala kahanemine kuivamisel, ahjukuiv puit** 11,8 % * – niiskusesisaldus 12%; ** – niiskusesisaldus 0%.
Punase kuuse vaiku närisid juba indiaanlased. 1848 hakkas John Curtis kuusevaigust närimiskummi valmistama[10] ja seda müüakse tänapäevani, harilikult piparmündi- või puuviljamaitseliste lisanditega.
Värskeid kuusekasve keedetakse kuuseõlle valmistamisel.[11]
Linnade haljastuses ja tööstuspiirkondades on punase kuuse kasutamine raskendatud, kuna ta ei talu tahma ja heitgaase.[6] Samuti ei talu ta mulla kuivust.[5]
Punane kuusk on Kanadas asuva Nova Scotia provintsi rahvuspuu.[12]
Kasvatamine Eestis
Punane kuusk toodi Euroopasse juba 1750. aastal. Eestisse jõudis ta 19. sajandil, kuid on meil vähe levinud. Tema esinemist on registreeritud Tallinnas, Tartus, Vana-Antslas, Orul, Porkunis, Olustveres ning Saaremaal Kuusenõmmel ja J. Alase dendraariumis.[5] Praegu võib neid näha Tallinna Botaanikaaias ja Luua arboreetumis.[13] Eestis kipub ta olema külmahell, samuti kannatab ta kuivadel kasvupinnastel, kuna koduses levilas on sademete hulk pea kaks korda suurem. Külmakindlaid seemikuid peaks saama areaali põhjaosast kogutud seemnete abil.[5] Soomes kasvatatakse punast kuuske üksikute rühmadena Mustilas ja Punkaharjus ning seal on puud külmakindlad ja kannavad ka käbisid.[6]
Viited
-
↑ "Conifer database: "Picea rubens".". Catalogue of Life: 2010 Annual Checklist. Vaadatud 18.11.2010. Inglise.
-
↑ Conifer Specialist Group (1998). Picea rubens. IUCNi punase nimistu ohustatud liigid. IUCN 2010.
-
↑ 3,0 3,1 3,2 3,3 3,4 "Picea rubens". www.fs.fed.us. Vaadatud 18.11.2010. Inglise.
-
↑ 4,0 4,1 4,2 4,3 4,4 4,5 4,6 4,7 4,8 "Red Spruce". www.na.fs.fed.us. Vaadatud 18.11.2010. Inglise.
-
↑ 5,0 5,1 5,2 5,3 5,4 5,5 V. Hainla, U. Valk. "Eestis kasvavad kuused", Tartu, 1961.
-
↑ 6,0 6,1 6,2 6,3 6,4 Endel Laas. "Dendroloogia", Tallinn: Valgus, 1987.
-
↑ L. J. Sheppard, R. I. Smith and M. G. R. Cannell. "Frost hardiness of Picea rubens growing in spruce decline regions of the Appalachians". Tree Physiology 5, 25-37, 1989. Victoria, Canada: Heron Publishing. Failitüüp: PDF. Vaadatud 18.11.2010. Inglise.
-
↑ Samuel V. Glass, Samuel L. Zelinka. "Wood Handbook, Chapter 04: Moisture Relations and Physical Properties of Wood". www.fpl.fs.fed.us, 2010. Forest Products Laboratory. Failitüüp: PDF. Vaadatud 18.11.2010. Inglise.
-
↑ David E. Kretschmann. "Wood Handbook, Chapter 05: Mechanical Properties of Wood". www.fpl.fs.fed.us, 2010. Forest Products Laboratory. Failitüüp: PDF. Vaadatud 18.11.2010. Inglise.
-
↑ Spruce Gum in Maine and Atlantic Canada (inglise keeles)
-
↑ The WoodExplorer (inglise keeles)
-
↑ Flora of North America (inglise keeles)
-
↑ Eino Laas. "Okaspuud", Tartu: Atlex, 2004.
Välislingid
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Punane kuusk: Brief Summary
(
Estonyaca
)
wikipedia ET tarafından sağlandı
Punane kuusk (Picea rubens) on igihaljas okaspuuliik kuuse perekonnast. Selle looduslik levila Põhja-Ameerika kirdeosas, kuid 18. sajandi keskel toodi seda ka Euroopasse ning vähesel määral kasvatatakse seda ka Soomes ja Eestis.
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Punakuusi
(
Fince
)
wikipedia FI tarafından sağlandı
Punakuusi (Picea rubens) on Pohjois-Amerikan itäosista kotoisin oleva ainavihanta, keskikokoinen havupuu, joka kuuluu kuusten sukuun ja mäntykasvien heimoon.[2][3] Se on valittu Nova Scotian provinssipuuksi.[4]
Ulkonäkö ja koko
Täysikasvuinen punakuusi.
Punakuusi on pitkäikäinen puu, joka alkaa tuottaa siementä 15–20 vuoden iässä ja elää usein yli 350 vuoden ikäiseksi.[2] Se kasvaa keskimäärin 18–23 metriä korkeaksi ja rinnankorkeusläpimitaltaan 30–61 senttimetriä paksuksi. Suurin elossa oleva punakuusi on 33,5 metriä korkea ja 133 senttimetriä paksu ja kasvaa Great Smoky Mountainsin kansallispuistossa Pohjois-Carolinassa.[5]
Punakuusella on lähelle maanpintaa levittäytyvä juuristo, kapean kartiomainen latvus ja siirottavat haarat, jotka ympäröivät runkoa säännöllisinä kiehkuroina.[2][3] Runkoa peittää karhea, hilseilevä, harmaan- tai punertavanruskea kaarna.[3][6] Kuluvan kesän kasvainranka on paksuhko, kellanruskea ja kalju tai karvainen. Kärkisilmut ovat punertavanruskeat, pistäväkärkiset ja läpimitaltaan 5–8 millimetriä.[6] Lehdet ovat kiiltäviä, kellan- tai tummanvihreitä, taipuisia neulasia.[6][7] Yksittäinen neulanen on 8–25 millimetriä pitkä, teräväkärkinen, läpileikkaukseltaan neliskulmainen ja kauttaaltaan ilmarakojen peittämä.[6]
Muiden kuusien tapaan punakuusi on yksikotinen eli samassa yksilössä kasvaa sekä hede- että emikukintoja.[3] Käpymäiset kukinnot syntyvät heinäkuussa edellisten vuosikasvaimien oksanhankoihin mutta kehittyvät lopullisesti vasta seuraavan vuoden toukokuussa. Hedekukinnot ovat riippuvat ja kirkkaanpunaiset. Emikukinnot ovat pystyt ja kirkkaanvihreät, hivenen purppuranpunaiseen vivahtavat.[5] Emikukinnot kypsyvät syys–lokakuuhun mennessä luumunmuotoisiksi kävyiksi, jotka ovat vaaleanpunaruskeita ja pituudeltaan 3–5 senttimetriä.[5][6][7][8] Ohuiden ja pyöreäkärkisten käpysuomujen alle kehittyy ohutkuorisia ja siivekkäitä siemeniä.[3][7] Kävyt eivät säily pitkään puussa ja niistä vapautuneet siemenet itävät yleensä jo seuraavana keväänä.[5][7]
Levinneisyys
Punakuusi on kotoisin Pohjois-Amerikan itäosista. Sen luontainen levinneisyysalue ulottuu Kanadassa Cape Bretonin saarelle, Nova Scotiaan, New Brunswickiin, Etelä-Québeciin ja Kaakkois-Ontarioon sekä Yhdysvalloissa Maineen, New Yorkiin, Koillis-Pennsylvaniaan, New Jerseyn pohjoisosiin ja Koillis-Massachusettsiin. Se on levinnyt Appalakkien vuoristoa pitkin myös Marylandin länsiosiin, Länsi-Virginian itäosiin, Virginian pohjois- ja itäosiin, Pohjois-Carolinan länsiosiin sekä Tennesseen itäosiin.[2]
Elinympäristö
Punakuusi kasvaa viileässä ja kosteassa ilmastossa aina 2 000 metrin korkeuteen asti.[2][6] Se kestää hyvin varjostusta ja viihtyy korvissa, humuspeitteisillä kallioilla sekä tuoreissa, niukkaravinteisissa metsissä.[2][7]
Käyttö
Punakuusi on valkokuusen ohella Pohjois-Amerikan tärkeimpiä paperi- ja sahateollisuuden raaka-aineita. Sen puuaines on vaaleaa, kevyttä, suorasyistä ja sitkeää ja soveltuu erinomaisesti jousisoittimien rakentamiseen.[5] Pohjois-Amerikan intiaanit käyttivät punakuusen pitkiä juuria naruna ja tilkitsivät sen hartsilla koloja ja vuotavia saumoja.[8] 1800- ja 1900-lukujen vaihteessa sen pihkasta valmistettiin teollisesti purukumia Mainessa.[2][5]
Punakuusi on monille lintulajeille tärkeä pesimä- ja suojapaikka ja ravinnonlähde. Sen kaarnaa, siemeniä ja neulasia käyttävät ravinnokseen myös lumikenkäjänis, puupiikkisiat ja muut pienet nisäkkäät.[2]
Lähteet
- Hämet-Ahti, Leena & Palmén, Annikki & Alanko, Pentti & Tigerstedt, Peter M. A.: Suomen puu- ja pensaskasvio. Helsinki: Dendrologian Seura, 1992. ISBN 951-96557-0-0.
Viitteet
-
↑ a b Picea rubens IUCN Red List of Threatened Species. International Union for Conservation of Nature, IUCN, Iucnredlist.org. (englanniksi)
-
↑ a b c d e f g h Janet Sullivan: Picea rubens Fire Effects Information System. USDA Forest Service. Viitattu 8.4.2011. (englanniksi)
-
↑ a b c d e Alanko, 67.
-
↑ Red Spruce nslegislature.ca. Viitattu 21.4.2011. (englanniksi)
-
↑ a b c d e f Blum, Barton: Picea rubens Northeastern Area State & Private Forestry. Viitattu 18.8.2009.
-
↑ a b c d e f Hong Song: Picea rubens Flora of North America. Viitattu 18.8.2009.
-
↑ a b c d e Alanko, 72.
-
↑ a b Picea rubens The Gymnosperm Database. Viitattu 25.4.2011. (englanniksi)
Aiheesta muualla
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Punakuusi: Brief Summary
(
Fince
)
wikipedia FI tarafından sağlandı
Punakuusi (Picea rubens) on Pohjois-Amerikan itäosista kotoisin oleva ainavihanta, keskikokoinen havupuu, joka kuuluu kuusten sukuun ja mäntykasvien heimoon. Se on valittu Nova Scotian provinssipuuksi.
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Épinette rouge
(
Fransızca
)
wikipedia FR tarafından sağlandı
Picea rubens • épicéa rouge
L'épinette rouge ou épicéa rouge (Picea rubens) est un arbre (conifère) appartenant à la famille des Pinaceae et au genre Picea (connu sous le nom d'épinette en Amérique du Nord et d'épicéa en Europe) natif de l'Amérique du Nord. Son bois de qualité, léger mais résistant, est largement utilisé comme bois d'œuvre, pour la construction, et aussi pour la fabrication de pâte à papier et d'instruments de musique à cordes (guitare, violon).
Description
L'épinette rouge est une espèce de fin de succession, c'est-à-dire qu'elle occupe une place prépondérante dans les derniers stades de la mise en place d'une forêt naturelle. Elle est très tolérante à l'ombre et peut aisément reprendre une croissance vigoureuse après des décennies de domination par les arbres voisins. Cela se produit généralement à la faveur d'une éclaircie dans le couvert forestier via une perturbation tel un chablis partiel. L'Épinette rouge est un arbre de taille moyenne pouvant atteindre 26 m de hauteur et 60 cm de diamètre[1]. Cet arbre peut vivre jusqu'à 400 ans.
Son écorce mince et écailleuse est de couleur brune à rougeâtre pâle, et ses rameaux sont orangés. Ses aiguilles sont d'un vert jaunâtre, elles sont brillantes (lustrées), mesurent de 1,2 à 2,1 cm et leur section est quadrangulaire.[réf. nécessaire]
Répartition et habitat
Aire de répartition naturelle de l'épinette rouge (
Picea rubens)
Cette espèce est native de l'Est de l'Amérique du Nord (États-Unis, Canada). Préfère les sols secs à humides. Abondance au sud du St-Laurent, en forêt feuillue et mélangée (mixte).
Au Québec, elle est présente dans les domaines de l'érablière à tilleul, de l'érablière à bouleau jaune et surtout dans celui de la sapinière à bouleau jaune, principalement dans les Appalaches. Elle s'établit en général sur des stations fraîches. Un sol sableux profond bien ou modérément bien drainé lui offre des conditions idéales de croissance.
De nos jours, la régression, voire la raréfaction de l'épinette rouge est attribuable à sa surexploitation et à ses difficultés de pouvoir se régénérer suffisamment après la coupe forestière.
Reproduction et développement
L'épinette rouge se reproduit à partir de graines. Les arbres semenciers sont productifs à partir de l’âge de 30 ans. Par la suite, l′intervalle de temps entre deux bonnes années semencières varie entre 3 et 8 ans. Entre-temps, le peu de semences produites est plus souvent qu'autrement consommé par les petits rongeurs (écureuils, souris). La viabilité des semences au sol est très faible (quelques mois, au maximum 1 an). La chute des graines se déroule d′octobre à mars et la germination a lieu au printemps qui suit. La dispersion des graines se fait sur une distance maximale de 100 m des semenciers. Très petites, les graines ont tout juste assez de réserves nutritives pour faire émerger les plantules, ces dernières devant rapidement par la suite puiser dans le substrat de sol pour se développer.
Épinette rouge mature en forêt mixte.
La stratégie de reproduction privilégiée de l'épinette rouge consiste à établir au fil du temps une banque de semis préétablis sous le couvert forestier. Pour s'établir, les semis requièrent un microenvironnement stable, ombragé, frais et suffisamment humide, comme des gros débris ligneux à un stade de décomposition avancé. Ce type de microsite surélevé favorise aussi les semis qui se retrouvent en meilleure position pour concurrencer les autres espèces. La croissance initiale des racines et de la tige des jeunes semis est très lente, ce qui les rend fragiles et vulnérables au recouvrement par la litière feuillue ou la neige. Une fois établie, la croissance de la régénération d'épinette rouge en forêt naturelle augmente avec l'ouverture progressive du couvert forestier (trouées engendrées par les épidémies d'insectes et le chablis partiel). Les semis et les gaules soudainement exposés à la pleine lumière et aux extrêmes de température, notamment à la suite d'opérations de coupe forestière, présentent des difficultés d'acclimatation physiologique pouvant causer leur dépérissement et leur mort à court ou moyen terme.
Étymologie
L'Épinette rouge est aussi connu sous le nom de prusqueur rouge[1]. « Épinette » provient de « pin » dont on a ajouté le suffixe « -ette ». Quant au « é- » il aurait été ajouté par analogie à « épine »[2].
Le genre Picea est dérivé du latin pix et signifie « poix » (ou résine). Il était à l'origine le nom latin du pin, mais a été plutôt retenu pour le genre de l'épinette (ou épicéa)[3]. Quant au spécifique rubens, il s'agit d'un mot latin signifiant « rougeâtre ». Il fait allusion au cônes qui sont de couleur brun-rougeâtre[4].
Notes et références
Annexes
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Épinette rouge: Brief Summary
(
Fransızca
)
wikipedia FR tarafından sağlandı
Picea rubens • épicéa rouge
L'épinette rouge ou épicéa rouge (Picea rubens) est un arbre (conifère) appartenant à la famille des Pinaceae et au genre Picea (connu sous le nom d'épinette en Amérique du Nord et d'épicéa en Europe) natif de l'Amérique du Nord. Son bois de qualité, léger mais résistant, est largement utilisé comme bois d'œuvre, pour la construction, et aussi pour la fabrication de pâte à papier et d'instruments de musique à cordes (guitare, violon).
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Brúngreni
(
İzlandaca
)
wikipedia IS tarafından sağlandı
Brúngreni, vísindaheiti Picea rubens, er tegund af greni ættuðu úr austurhluta Norður Ameríku, frá austur Quebec til Nova Scotia, og frá New England suður til Adirondack-fjöllum og Appalasíu-fjöllum til vestur Norður-Karólínu.[2][3][4]
Þessi tegund gengur undir fjölda nafna á ensku; red spruce, yellow spruce, West Virginia spruce, eastern spruce, og he-balsam (sjá Abies fraseri; "she-balsam").[5][6]
Lýsing
Brúngreni er sígrænt[7] shade-tolerant, late successional[8] barrtré sem verður við bestu aðstæður 18 til 40 metra hátt með stofnþvermál um 60 sm, þó geta einstaka tré náð 46 metra hæð og 1 meters þvermáli. Það er með mjókeilulaga krónu. Barrið er nálarlaga, gulgrænt, 12 til 15 mm langt, fjórhliða, sveigt, með hvössum oddi, og stendur út frá öllum hliðum greinarinnar. Börkurinn er grábrúnn að utan og rauðbrúnn að innan, þunnur og hreistraður. Viðurinn er léttur og mjúkur, með mjóum árhringjum, og er með lítilsháttar rauðum blæ.[9] Könglarnir eru sívalir, 3 til 5 sm langir, með gljáandi rauðbrúnum lit og stíft köngulhreistur.[2][3][4][10]
Búsvæði
Brúngreni vex meðalhratt til hægt, lifir í 250 til 450+ ár, og er mjög skuggþolið ungt.[11] Það er oft í hreinum skógum ("stands") eða í bland við Pinus strobus, balsamþin, eða svartgreni.[12] Búsvæði þess er rakur en vel drenaður sendinn moldarjarðvegur, yfirleitt hátt yfir sjávarmáli. Brúngreni skaðast auðveldlega af vindkasti og súru regni.
Skyldar tegundir
Það er náskylt Svartgreni, og blendingar á milli þeirra eru algengir þar sem útbreiðslusvæði þeirra mætast.[2][3][4]
Nytjar
Brúngreni er notað sem jólatré og pappamassi og í timburframleiðslu.[13] Það er einnig frábært "tonewood", og er notað í marga vandaðri "acoustic" gítara og fiðlur. Hægt er að gera "spruce gum" úr safanum.[10] Greinar með barri eru notaðar til að gera grenibjór. Einnig er gerður úr því "grenibúðingur".
Brúngreni er héraðstré Nova Scotia.[3]
Skaðvaldar
Könglar af brúngreni í Pisgah National Forest
Eins og flest tré, eru ýmis skordýr sem leggjast á það. Aðal skaðvaldurinn er "spruce budworm" (nokkrar tegundir af ættkvíslinni Choristoneura, en sérstaklega Choristoneura fumiferana), þó er hann meira vandamál í hvítgreni og balsamþin.[14][15] Annað sem hefur hrjáð brúngreni er aukning súrs regns og núverandi loftslagsbreytingar.[16]
Tilvísanir
-
↑ Farjon, A. (2013). „Picea rubens“. IUCN Red List of Threatened Species. IUCN. 2013: e.T42335A2973542. doi:10.2305/IUCN.UK.2013-1.RLTS.T42335A2973542.en. Sótt 11. nóvember 2016.
-
↑ 2,0 2,1 2,2 Farjon, A. (1990). Pinaceae. Drawings and Descriptions of the Genera. Koeltz Scientific Books ISBN 3-87429-298-3.
-
↑ 3,0 3,1 3,2 3,3 "Picea rubens". Flora of North America (FNA). Missouri Botanical Garden – via eFloras.org.
-
↑ 4,0 4,1 4,2 Gymnosperm Database: Picea rubens
-
↑ Blum, Barton M. (1990). "Picea rubens". In Burns, Russell M.; Honkala, Barbara H. Conifers. Silvics of North America. Washington, D.C.: United States Forest Service (USFS), United States Department of Agriculture (USDA). 1 – via Northeastern Area State and Private Forestry [www.na.fs.fed.us]
-
↑ Peattie, Donald Culross (1. janúar 1948). A Natural History of Trees of Eastern and Central North America (enska). Houghton Mifflin Harcourt. bls. 51. ISBN 0-395-58174-5.
-
↑ „Red Spruce (Rubens)“. Garden Guides. Sótt 27. febrúar 2014.
-
↑ Dumais, D; Prevost, M (June 2007). „Management for red spruce conservation in Quebec: The importance of some physiological and ecological characteristics – A review“. Forestry Chronicle. 83 (3): 378–392. doi:10.5558/tfc83378-3. Sótt 27. febrúar 2014.
-
↑ „Red Spruce“ (PDF). USDA NRCS. Sótt 26. febrúar 2014.
-
↑ 10,0 10,1 Atlantic Forestry Centre: Red Spruce
-
↑ http://www.ldeo.columbia.edu/~adk/oldlisteast/#spp
-
↑ Peter White, "Boreal Forest," Encyclopedia of Appalachia (Knoxville, Tenn.: University of Tennessee Press, 2006), pp. 49-50.
-
↑ „Red Spruce“. The Wood Database. Sótt 27. febrúar 2014.
-
↑ Blum, Barton. „Red Spruce“. Encyclopedia of Life. Sótt 27. febrúar 2014.
-
↑ U.S. Department of Agriculture-Forest Servic: Spruce budworm
-
↑ Houle, Daniel (2012). „Compositional vegetation changes and increased red spruce abundance during the Little Ice Age in a sugar maple forest of north-eastern North America“. Plant Ecology. 213 (6): 1027–1035. doi:10.1007/s11258-012-0062-0.
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Brúngreni: Brief Summary
(
İzlandaca
)
wikipedia IS tarafından sağlandı
Brúngreni, vísindaheiti Picea rubens, er tegund af greni ættuðu úr austurhluta Norður Ameríku, frá austur Quebec til Nova Scotia, og frá New England suður til Adirondack-fjöllum og Appalasíu-fjöllum til vestur Norður-Karólínu.
Þessi tegund gengur undir fjölda nafna á ensku; red spruce, yellow spruce, West Virginia spruce, eastern spruce, og he-balsam (sjá Abies fraseri; "she-balsam").
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Picea rubens
(
İtalyanca
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Il peccio rosso (Picea rubens Sarg., 1898) — conosciuto anche come abetina rossa, picea rossa del Canada e altri nomi simili[2] — è una specie di peccio nativo dell'America nord-orientale, diffuso dal Québec orientale alla Nuova Scozia, e dal sud della Nuova Inghilterra nei Monti Adirondack e Appalachi alla parte occidentale della Carolina del Nord.[3][4][5]
Etimologia
Il nome generico Picea, utilizzato già dai latini, potrebbe, secondo un'interpretazione etimologica, derivare da Pix picis = pece, in riferimento all'abbondante produzione di resina.[6] Il nome specifico rubens è riferito al colore rosso dei coni femminili.[7]
Descrizione
Il peccio rosso è un albero delle Conifere che in condizioni ottimali cresce fino a 18-40 m di altezza con un tronco del diametro di circa 60 cm, sebbene esemplari eccezionali possano raggiungere i 46 m di altezza e i 30 cm di diametro. Ha una corona conica ristretta. Le foglie sono aghiformi, giallo-verdi, lunghe 12-15 mm, quadrangolari, curve con una punta acuminata, e si estendono da tutti i lati del ramoscello. La corteccia è grigio-bruna sulla superficie e rosso-bruna all'interno, sottile e squamosa. Gli strobili sono cilindrici, lunghi 3-5 cm, di colore vitreo rosso-bruno e con scaglie dure. Gli strobili pendono verso il basso dai rami.[3][4][5][8]
Distribuzione e habitat
Il peccio rosso cresce a una velocità da lenta a moderata, vive da 250 a 450 anni e oltre, e da giovane tollera bene l'ombra.[9] Si trova spesso in semplici boschi o foreste mescolato con il pino bianco americano, l'abete balsamico o il peccio nero. Insieme all'abete di Fraser, il peccio rosso è uno dei due tipi principali di alberi della foresta di abeti rossi degli Appalachi Meridionali, un ecosistema distinto che si trova solo nelle altezze più elevate dei Monti Appalachi Meridionali.[10] Il suo habitat è il limo sabbioso, umido ma ben drenato, spesso ad alta quota. Il peccio rosso può essere facilmente danneggiato dalle raffiche di vento e dalle piogge acide.
Foreste notevoli di pecci rossi si possono vedere nell'Area panoramica di Gaudineer (Gaudineer Scenic Area) una foresta vergine di pecci rossi localizzata nella Virginia Occidentale, nella Valle di Canaan, nella Riserva naturale occidentale di Roaring Plains, Riserva naturale di Dolly Sods, nel Monte Spruce e nello Spruce Knob, anch'essi tutti in Virginia Occidentale, e in tutti i siti in cui un tempo sorgevano estese foreste di pecci rossi. Alcune aree di queste foreste, in particolare nella Riserva naturale occidentale di Roaring Plains, nella Riserva naturale di Dolly Sods come pure in certe zone del Monte Spruce, stanno avendo un recupero piuttosto sostanziale.
Tassonomia
È strettamente imparentato con il peccio nero e gli ibridi tra i due sono frequenti dove i loro ambienti naturali coincidono.[3][4][5]
Sinonimi
Si annoverano i seguenti sinonimi:[11]
-
Abies rubra (Du Roi) Poir.
-
Picea acutissima (Münchh.) J.G.Jack
-
Picea americana Suringar
-
Picea australis Small
-
Picea rubens f. virgata Fernald & Weath.
-
Picea rubra var. pusilla Peck
-
Picea rubra var. virgata (Rehder) Rehder
-
Picea rubra f. virgata Rehder
-
Pinus americana Gaertn.
-
Pinus rubra var. violacea Endl.
Usi
Il peccio rosso si usa per gli alberi di Natale ed è un legno importante per ricavare la polpa della carta. È anche un eccellente legno per gli strumenti musicali e si usa in molte chitarre e violini di pregio. La linfa può essere utilizzata per fare la "gomma di peccio" (spruce gum, un tipo di gomma da masticare).[8] I ramoscelli fronzuti a loro volta vengono bolliti come parte della ricetta per la preparazione della "birra di peccio" (spruce beer, una birra aromatizzata con le essenze del peccio rosso).
Il peccio rosso, infine, è l'albero che compare nello stemma della provincia canadese della Nuova Scozia.[4]
Conservazione
L'Iniziativa per il ripristino dei pecci degli Appalachi Centrali (Central Appalachian Spruce Restoration Initiative, CASRI)[12] cerca di unire diversi soci con l'obiettivo di ripristinare gli ecosistemi storici degli abeti rossi in tutti i luoghi panoramici ad alta quota degli Appalachi Centrali.
Note
-
^ (EN) Farjon, A. 2013., Picea rubens, su IUCN Red List of Threatened Species, Versione 2020.2, IUCN, 2020.
-
^ Center for Wood Anatomy Research - USDA Forest Service - Forest Products Laboratory, Technology Transfer Fact Sheet - Picea rubens Sarg. (PDF), su fpl.fs.fed.us. URL consultato il 04-06-2013.
-
^ a b c Farjon, A. (1990). Pinaceae. Drawings and Descriptions of the Genera. Koeltz Scientific Books ISBN 3-87429-298-3.
-
^ a b c d Flora of North America: Picea rubens
-
^ a b c Gymnosperm Database: Picea rubens Archiviato il 2 maggio 2006 in Internet Archive.
-
^ Picea rubens, in Treccani.it – Vocabolario Treccani on line, Istituto dell'Enciclopedia Italiana. URL consultato il 22 maggio 2020.
-
^ (EN) Aljos Farjon, A Handbook of the World's Conifers (2 vols.): Revised and Updated edition, Brill, 2017, p. 614. URL consultato il 22 maggio 2020.
-
^ a b Atlantic Forestry Centre: Red Spruce Archiviato il 18 agosto 2008 in Internet Archive.
-
^ Eastern OLDLIST: A database of maximum tree ages for Eastern North America
-
^ Peter White, "Boreal Forest", Encyclopedia of Appalachia, Knoxville, Tenn., University of Tennessee Press, 2006, pp. 49-50.
-
^ Picea rubens Sarg., su Plants of the world. URL consultato il 22 maggio 2020.
-
^ Burks, Evan (2010), "Return of the Red Spruce", Wonderful West Virginia; vol. 74, n. 12 (dicembre), pp. 6-11.
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Picea rubens: Brief Summary
(
İtalyanca
)
wikipedia IT tarafından sağlandı
Il peccio rosso (Picea rubens Sarg., 1898) — conosciuto anche come abetina rossa, picea rossa del Canada e altri nomi simili — è una specie di peccio nativo dell'America nord-orientale, diffuso dal Québec orientale alla Nuova Scozia, e dal sud della Nuova Inghilterra nei Monti Adirondack e Appalachi alla parte occidentale della Carolina del Nord.
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Raudonoji eglė
(
Litvanca
)
wikipedia LT tarafından sağlandı
Raudonoji eglė (lot. Picea rubens, angl. Red Spruce) – pušinių (Pinaceae) šeimos, eglių (Picea) genties visažalis spygliuotis medis. Paplitusi Šiaurės Amerikos rytuose (Kvebeke, Adirondako kalnuose, Naujojoje Anglijoje, Apalačuose iki Šiaurės Karolinos).
Aukštis apie 18-40 m. Kamieno skersmuo 0,6 m, nors pasitaiko medžių, kurių kamieno skersmuo 1,3 m. Spygliai 12-15 mm ilgio, gelsvai žali. Kankorėžiai cilindriški, 3-5 cm ilgio, nuo šakos nusvirę žemyn.
Raudonoji eglė artima juodajai eglei, ir tarpusavyje kryžminasi.
Raudonoji eglė yra Naujosios Škotijos provincijos (Kanada) simbolis. Medis naudojamas medienos, popieriaus pramonėje, muzikos instrumentų gamybai.
Vikiteka
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Rødgran
(
Norveççe
)
wikipedia NO tarafından sağlandı
Rødgran (Picea rubens) er et bartre i furufamilien som vokser i østlige Nord-Amerika.
Den blir opptil 40 m høy med en stammediameter på 100 cm. Krona er smalt kjegleformet. Barken er gråbrun eller rødbrun. Nålene er 0,8–2,5 cm lange og firkantet i tverrsnitt. Konglene er 2,3–4,5 cm lange.[1][2][3]
Rødgran vokser i rene bestander som danner store skoger. Den kan også vokse sammen med andre trær som balsamgran, svartgran, papirbjørk, weymouthfuru, canadahemlokk, sukkerlønn, amerikabøk, gulbjørk, østamerikansk tuja, kvitgran og rødlønn. Vanlige busker er blant annet bærlyng, korsved, bringebær, krypberglyng, virginiatrollhassel, tresøtmispel, nebbhassel og canadabarlind. Trevirket er likt andre granarter og brukes til trelast og papirmasse.[4]
Arten er utbredt fra De kanadiske kystprovinser og Saint-Pierre og Miquelon sørvestover til Maine, sørlige Québec, sørøstlige Ontario, sentrale New York, østlige Pennsylvania, nordlige New Jersey og Massachusetts. Spredte forekomster finnes videre sørover i Appalachene: helt vest i Maryland, øst i Vest-Virginia, nord og øst i Virginia, vest i Nord-Carolina og øst i Tennessee.[4][3]
Referanser
Eksterne lenker
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Rødgran: Brief Summary
(
Norveççe
)
wikipedia NO tarafından sağlandı
Rødgran (Picea rubens) er et bartre i furufamilien som vokser i østlige Nord-Amerika.
Den blir opptil 40 m høy med en stammediameter på 100 cm. Krona er smalt kjegleformet. Barken er gråbrun eller rødbrun. Nålene er 0,8–2,5 cm lange og firkantet i tverrsnitt. Konglene er 2,3–4,5 cm lange.
Rødgran vokser i rene bestander som danner store skoger. Den kan også vokse sammen med andre trær som balsamgran, svartgran, papirbjørk, weymouthfuru, canadahemlokk, sukkerlønn, amerikabøk, gulbjørk, østamerikansk tuja, kvitgran og rødlønn. Vanlige busker er blant annet bærlyng, korsved, bringebær, krypberglyng, virginiatrollhassel, tresøtmispel, nebbhassel og canadabarlind. Trevirket er likt andre granarter og brukes til trelast og papirmasse.
Arten er utbredt fra De kanadiske kystprovinser og Saint-Pierre og Miquelon sørvestover til Maine, sørlige Québec, sørøstlige Ontario, sentrale New York, østlige Pennsylvania, nordlige New Jersey og Massachusetts. Spredte forekomster finnes videre sørover i Appalachene: helt vest i Maryland, øst i Vest-Virginia, nord og øst i Virginia, vest i Nord-Carolina og øst i Tennessee.
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Świerk czerwony
(
Lehçe
)
wikipedia POL tarafından sağlandı
Świerk czerwony (Picea rubens Sarg.) – gatunek drzewa z rodziny sosnowatych (Pinaceae). Pochodzi ze wschodniej Ameryki Północnej. Obszar rodzimego występowania rozciąga się od wschodniego Quebecu do Nowej Szkocji oraz od Nowej Anglii na południe przez Appalachy do zachodniej Północnej Karoliny. Jest uprawiany w wielu krajach świata[3]
Morfologia
- Pokrój
- Drzewo o wąskiej, stożkowatej koronie.
- Pień
- Osiąga wysokość 18-30 m, średnica pnia ok. 60 cm, wyjątkowo może osiągnąć 46 m wysokości i 130 cm średnicy pnia. Kora szarobrązowa z wierzchu, czerwonobrązowa od wewnątrz, cienka i łuszcząca się. Gałęzie rosną poziomo, nie zwisają.
- Liście
- Igły jasnozielone do ciemnozielonych, o długości 12-15 mm i kwadratowym przekroju poprzecznym ułożone są równomiernie dookoła na gałęziach. Pozostają na drzewie przez wiele lat.
- Kwiaty
- Gatunek jednopienny. Kwiaty męskie cylindryczne, czerwonawe, z czasem żółto-brązowe. Kwiaty żeńskie fioletowo-zielone.
- Szyszki
- Cylindryczne, długości 3-5 cm, czerwonobrązowe. Łuski zaokrąglone, czasem lekko ząbkowane. Nasiona uwalniane jesienią, rozsiewane przez wiatr lub deszcz. Szyszki opadają przeważnie już pierwszej zimy.
- Korzeń
- Płytki system korzeniowy, większość korzeni znajduje się tuż pod powierzchnią ziemi, do głęgokości 40-50 cm.
Biologia i ekologia
Drzewo długowieczne, żyje średnio 250-350 lat. Toleruje zacienienie. Tworzy lasy jednogatunkowe lub mieszane, razem z sosną wejmutką (Pinus strobus), jodłą balsamiczną (Abies balsamea), jodłą Frasera (Abies fraseri). Występuje w klimacie o chłodnych, wilgotnych latach i mroźnych zimach. Rośnie na wysokościach od 0 do 1450 m n.p.m., w Appalachach od 980 do 1890 m, powyżej zastępuje go jodła Frasera.
Zastosowanie
- Świerk czerwony chętnie używany jest jako świąteczna choinka
- Wykorzystywany do wyrobu papieru.
Zagrożenia
- Świerkowi czerwonemu zagrażają wiatrołomy i kwaśne deszcze.
- Ze względu na cienką korę, płytkie korzenie i zapalne igły jest wrażliwy na ogień, przez co drzewa nie przeżywają pożaru.
Przypisy
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Świerk czerwony: Brief Summary
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Lehçe
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wikipedia POL tarafından sağlandı
Świerk czerwony (Picea rubens Sarg.) – gatunek drzewa z rodziny sosnowatych (Pinaceae). Pochodzi ze wschodniej Ameryki Północnej. Obszar rodzimego występowania rozciąga się od wschodniego Quebecu do Nowej Szkocji oraz od Nowej Anglii na południe przez Appalachy do zachodniej Północnej Karoliny. Jest uprawiany w wielu krajach świata
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Picea rubens
(
Ukraynaca
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wikipedia UK tarafından sağlandı
Поширення, екологія
Країни проживання: Канада (Нью-Брансвік, Ньюфаундленд, Нова Шотландія, Онтаріо, Острови Принца Едуарда, Квебек); США (Коннектикут, Мен, Массачусетс, Нью-Гемпшир, Нью-Джерсі, Нью-Йорк, Північна Кароліна, Пенсільванія, Теннессі, Вермонт, Вірджинія, Західна Вірджинія). Проживає від близько рівнем моря, на узбережжях морських провінцій Канади, до схил гір Аппалачі (1100 м до 1500 м над рівнем моря). У районах пн.-сх. рівнин вид росте в основному на кислих ґрунтах (рН 4-5,5) алювіального походження, в горах — на кислих, торф'яних або кам'янистих ґрунтах. Живе в районах з прохолодним, вологим океанічним кліматом, з річною кількістю опадів між 875 мм і 2000 мм (збільшення з висотою). Зазвичай змішується з Picea glauca чи Abies balsamea рідше з Picea mariana. Широколистяні дерева може бути поширеними або домінуючими, особливо на кращих землях.
Опис
Дерева в 40 м у висоту і 100 см діаметром. Крона вузько-конічна. Кора від сіро-коричневого до червонувато-коричневого кольору. Гілки горизонтального розповсюдження. Бруньки червонувато-коричневі, 5-8 мм, вершина гостра. Листя довжиною 0.8-2.5 (3) см, від жовто-зеленого до темно-зеленого кольору. Насіннєві шишки довжиною 2.3-4.5 (5) см. Шишки яйцевидні, блискучі, оранжево-коричневі. 2n = 24.
Найвище відоме дерево 46.33 м заввишки. У 2005 році було знайдено дерево віком 445 років в провінції Нью-Брансвік, Канада.
Використання
Є важливим джерелом деревини серед дерев середнього розміру. Деревина має малу вагу, кремового кольору, сильна і прямошарувата. Використовується для паперової маси, будівництва, будівництво човнів, флагштоків, особливо на музичних інструментах.
Загрози та охорона
Вид широко експлуатувався для деревини у 19 і початку 20 століть, особливо в південній частині ареалу в Аппалачах. У цей період також значно зріс вплив пожеж і розчищення лісів для ведення сільського господарства. Зовсім недавно (останні 100 років), цей вид став залежати від атмосферних забруднюючих речовин і кислотних осаджень. Проте, недавні перевірки цього виду в США виявили, що не було ніякого істотного зниження площ зростання і що в більшості районів було розширення цих площ. Цей вид відомий з ряду національних парків та інших охоронних територій.
Посилання
Галерея
Це незавершена стаття про родину Соснові.
Ви можете допомогти проекту, виправивши або дописавши її.
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Vân sam đỏ
(
Vietnamca
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wikipedia VI tarafından sağlandı
Picea rubens là một loài thực vật hạt trần trong họ Thông. Loài này được Sarg. miêu tả khoa học đầu tiên năm 1898.[1]
Chú thích
Liên kết ngoài
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Vân sam đỏ: Brief Summary
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Vietnamca
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Picea rubens là một loài thực vật hạt trần trong họ Thông. Loài này được Sarg. miêu tả khoa học đầu tiên năm 1898.
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Ель красная
(
Rusça
)
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Вид: Ель красная
Международное научное название
Picea rubens Sarg., 1898
Синонимы Pinus abies var. acutissima Münchh.
Ареал
Систематика
на ВикивидахИзображения
на Викискладе ITIS 18034NCBI 3333EOL 1033620GRIN t:316777IPNI 195909-2TPL kew-2563588 Ель красная (лат. Picea rubens) — один из видов ели, классифицирован Ч. Сарджентом.
Ареал
Ареал вида — район от реки Святого Лаврентия до Атлантического побережья Новой Шотландии и Новой Англии. На юге ель красная может встречаться на высотах свыше 1000 м от Адирондака до гор Аллегейни в Аппалачах, изолированные ареалы до Северной Каролины. Наиболее вид распространён в Новой Англии и атлантическом побережье Канады. Ель красная — один из символов Новой Шотландии.[1]
Ель красная выращивается и как декоративное растения (формы «Virgata» и «Nana»).[2]
Описание вида
Ель красная представляет собой вечнозелёное хвойное дерево. В зависимости от условий высота дерева достигает 20—40 м, но в сложных условиях высота может составлять 4—5 м. Диаметр ствола в среднем 60 см, но может достигать 1,5 м.[3] Хвоя около 10—15 мм длиной и около миллиметра шириной. Семена 2—3 мм длиной, с большим крылом. Шишки яйцевидные, 3—5 см длиной и 1,5—3,5 см шириной, зрелом виде — красновато-коричневые. Живёт ель красная 250—400 лет.
Наиболее близкий вид — ель чёрная, в общем ареале в естественных условиях могут образовываться гибриды.[4]
Галерея
Примечания
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Ель красная: Brief Summary
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Rusça
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Ель красная (лат. Picea rubens) — один из видов ели, классифицирован Ч. Сарджентом.
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