Spelaeoecia styx Kornicker ex Kornicker, Yager & Williams 1990

Spelaeoecia styx Kornicker

ETYMOLOGY.—From the Latin and Greek Styx (river in the nether world).

MATERIAL.—El Dorado Cave, South Andros Island, Great Bahama Bank: USNM 193440, 1 adult male, appendages on 1 slide, carapace in alcohol (paratype); USNM 193441, 1 undissected adult male in alcohol (holotype); USNM 193442, 1 adult female, appendages on slide, carapace in alcohol (paratype).

DISTRIBUTION.—Known only from El Dorado Cave, South Andros Island.

DESCRIPTION OF ADULT MALE (Figures 2–6).—Carapace uncalcified, flexible, elongate, dorsal margin straight, ventral margin convex but fairly straight at midlength, anteroventral and posteroventral margins evenly rounded (Figure 2a,d,g); anterior incisur dorsal to midheight (Figure 2a,f; anterior of valve viewed from inside with edge of valve forming triangular rostrum (Figure 2g); edge of valve with slight sinuosity between anterior tip of rostrum and incisur (Figure 2g); anterior outer part of valve overreaching triangular rostrum to form broadly rounded extension of rostrum (Figure 2 g). Posterodorsal corner of right valve with gland-bearing protuberance (Figure 2d,e,g,i).

Ornamentation: Surface with few long single bristles; small bristle at tip of protuberance in posterdorsal corner of right valve (Figure 2e,g,i). Surface appearing smooth, but at high resolution (× 40 objective, × 15 ocular, transmitted light) indistinct oblique striations observed on parts of valves (Figure 2h) (striations probably more distinct on living specimens). Edge of valve with 1 or 2 (usually 1) minute bristles between each pair of glandular tubes (Figure 2j).

Infold (Figure 2g): Broad infold except along hinge, narrower just posterior to inner end of incisur (Figure 2g); narrow list at about midwidth of infold extending from just ventral to incisur to posteroventral corner of valve; a 2nd list forming sclerotized bar just posterior to posterior hinge juncture, then extending ventrally as narrow ridge intersecting valve edge at posteroventral corner (Figure 2g).

Glands: Posterodorsal corner of right valve with protuberance bearing along dorsal edge 5 or 6 minute glandular openings forming row just anterior to minute V-shaped indentation (detail in Figure 2g); small bristle at posterior corner of indentation. Left valve with minute glandular protuberance about halfway between posterior hinge juncture and posterodorsal corner of valve (Figure 2c). Outer edge of infold with minute tube-like glandular openings between point just ventral to incisur and posterior hinge juncture (USNM 193440 with about 18 tubes on each valve, mostly along anteroventral and ventral margins (Figure 2j), none observed along anterior edge of rostrum, and very few along posterior margin of valve).

Selvage (Figure 2g,j): Narrow bare lamellar prolongation extending from point just ventral to incisur to posterodorsal corner of each valve, not observed along either edge of rostrum or straight edge of valve between posterior hinge juncture and posterodorsal corner (on right valve prolongation not present dorsal to apex of glandular protuberance).

Hingement: Each valve with narrow sclerotized ridges forming bar between posterior juncture of hinge and posterodorsal corner of valve that may serve as locking structures when valves are closed (exact configuration of ridges difficult to resolve with light microscopes) (Figure 2g).

Central Adductor Muscle Attachments (Figure 2a,b,g): Difficult to determine exact number on specimens in collection, 3 of the scars on right valve of USNM 193440 shown in Figure 2g.

Carapace Size: USNM 193440, length 0.98 mm, height 0.55 mm; USNM 193441, length 0.98 mm, height 0.46 mm.

First Antenna (Figures 2b, 3a,b): With 8 joints but 3rd and 4th joints partly fused, boundary indicated by ventral and dorsal marginal indentations: 1st joint with terminal ventral lobe with numerous short spines; 2nd joint with short, indistinct, diaphanous, dorsal bristle, and distal medial spinules; 3rd joint bare, longer than 4th; 4th joint with short dorsal bristle with indistinct short marginal spines. 5th joint slightly shorter than 4th, with 1 or 2 long ventral unringed filaments. 6th joint shorter than 5th, bare. 7th joint about same length as combined 5th and 6th joints, with short spinous dorsal a-bristle, ventral b-bristle about length of c-bristle, and long ventral c-bristle. 8th joint small with 4 terminal bristles (anterior medial d-bristle about twice length of a-bristle; long lateral e-bristle about same length as c-bristle, stout with indistinct rings and few widely spaced minute marginal spines; medial, f-bristle narrow, about length of e-bristle and oriented obliquely ventrally; and long, stout, g-bristle lateral to f-bristle and ventral to e-bristle, about length of e-bristle). The b- to g-bristles with terminal papillae. (The diaphanous bristle on the dorsal margin of the 2nd joint could be easily overlooked; it appears spine-like with the ×10 and ×20 objectives, but is bristle-like with the ×40 and ×100 objectives using ordinary transmitted light).

Second Antenna (Figures 2b, 3c–g): Protopodite with small medial spines forming rows near ventral margin (Figure 3c). Endopodite 3-jointed (Figure 3e–g): 1st joint elongate, with a slender spinous a- and b-bristle; 2nd joint with 2 terminal bare f- and g-bristles with parallel sides and minute papilla at tip (g-bristle medial, stouter and longer than f-bristle and weakly ringed proximally), 1 minute lateral peg near base of f-bristle, and 1 slender, dorsal, spinous c-bristle; 3rd joint with equilength h-, i-, and j-bristles, all about half length of g-bristle, bare with parallel sides; clasper elongate, straight, with small sclerotized process at terminal dorsal edge and 2 small terminal spines, clasper of right limb more than twice length that of left. Exopodite with 9 joints (Figure 3d): 1st joint divided into long proximal and short distal parts (well-developed separation of parts evident along sclerotized ventral edge of joint but not along dorsal edge) with few spines near midlength, and long bristle with natatory hairs and ventral spines reaching well past 9th joint; bristle of 2nd joint with ventral spines (stouter than those on bristle of 1st joint) and natatory hairs; bristle of joints 3–7 with only natatory hairs; bristle of 8th joint with dorsal spines and natatory hairs; 9th joint with 4 terminal bristles (dorsal bristle about same length as combined lengths of joints 6–9 and with few indistinct spines; adjacent bristle about 2 times longer and with dorsal spines; following bristle about 4 times longer and with dorsal spines; ventral bristle very long, with dorsal spines and natatory hairs); all long bristles of exopodite with 3 long proximal segments followed by closely spaced rings.

Mandible (Figures 2b, 4a–e): Coxale endite with proximal and distal sets of teeth separated by gap (Figure 4a–c): proximal set comprising 4 broad cusps plus small distal posterior triangular tooth; surface between cusps and just proximal to cusps with slender spines; 1 minute indistinct spinous bristle on corner just anterior to anterior cusp; 1 minute indistinct spinous bristle just posterior to posterior cusp; 3 spinous bristles adjacent to triangular tooth (exact number of bristles difficult to resolve, could be only 2); distal set of teeth comprising 2 flat teeth, each with 7 cusps; 1 stout curved spinous process and 1 minute bristle proximal to flat teeth. Basale (Figure 4d,e): distal edge with 5 terminal triangular cusps, 1 sharper triangular anterior cusp, and 1 small posterior cusp; lateral surface near distal edge with sharp tooth near midwidth; lateral surface distal to midlength with 2 minute and 4 longer bristles; anterior margin with 1 long bristle distal to midlength; posterior margin hirsute, with 2 distal bristles (proximal ringed in proximal and with unringed pointed tip, distal tubular); proximal medial surface with transparent plumose bristle on hirsute protuberance, and 1 short bristle near endopodite; 2 transparent plumose bristles on or close to dorsal margin; lateral surface near insertion of endopodite with 1 long bare bristle. Endopodite (Figure 4d,e): 1st joint with 3 bristles (1 dorsal, 1 long and 1 short on or near ventral margin); 2nd joint widening distally, with 2 or 3 terminal dorsal bristles (1 stout, unringed, claw-like, with marginal spines, 1 short, ringed, bare, medial, 0 or 1 short, ringed, bare, lateral), and 1 long, ringed, terminal, ventral bristle; 3rd joint with 2 long stout claw-like spinous terminal bristles, 4 short ringed bristles forming medial row along terminal edge, and 1 longer ringed bristle on terminal lateral edge; anterior margin and medial surface of 3rd joint hirsute.

Maxilla (Figures 2b, 4f): Endite I with 10 bristles (3 tubular); endite II with 2 proximal and about 7 terminal bristles; endite III with 1 proximal and about 4 terminal bristles; some bristles on each endite stout, pectinate. Coxale and basale fused; coxale with long stout dorsal plumose bristle; basale with 1 long ventral bristle. Endopodite: 1st joint with 5 anterior bristles (4 at midlength, 1 distal), 2 distal posterior bristles, 2 lateral and 2 medial bristles; end joint with 2 stout claws and 5 slender ringed bristles (rings not shown).

Fifth Limb (Figures 2b, 5a–d): Epipodite with plumose bristles in 3 groups of 4, 6, and 5 (Figure 5d). Protopodite with lateral glandular process (Figure 5b,c), lateral spines, and 2 ventral endites: endite I with 3 bristles with long barbs; endite II with 1 proximal medial bristle and 3 ventral bristles (1 with long spines, 2 bare or with short spines). Basale with medial hairs and spinules, 1 long lateral anterior bristle with long spines, 1 proximal medial bristle, and 6 ventral bristles (2 tending to be claw-like, 1 with long proximal spines, 3 ringed, either bare or with short marginal spines). Endopodite with 1 proximal medial bristle (bare or with few short spines) and 9 additional bristles (1 short tooth-like medial, subventral bristle, 1 short lateral subventral bristle, 2 claw-like unringed ventral bristles, 3 ringed ventral bristles (either bare or with short spines, and 2 anterior long bristles with long spines); few indistinct minute spines near base of tooth-like bristle (not shown on illustration). Exopodite (Figure 5a,b): 1st joint: dorsal margin with 1 long subterminal bristle, and 1 plumose bristle with base on or near dorsal margin and proximal to subterminal bristle; ventral margin with weak suture at about midlength dividing joint into 2 parts: proximal part with 3 slender ventral bristles (bare or with short spines), 1 long plumose lateral bristle near ventral margin, and 1 short medial distal bristle near ventral margin (with short marginal spines); distal part with 1 or 2 subterminal ventral bristles and 2 distal plumose lateral bristles. 2nd joint: dorsal margin with 1 distal bristle; ventral margin with 3 slender bristles near midlength. 3rd joint with 2 stout, claw-like, unringed bristles, and 1 slender ringed ventral bristle; all bristles of 3rd joint bare. (Poulsen's (1969:12) interpretation of the protopodite, basale, and endopodite is followed in the above description of the 5th limb.)

Sixth Limb (Figures 2b, 5e,f): Epipodite with plumose bristles in 3 groups of 6, 6, and 5 (Figure 5e). Protopodite separated from basale by indistinct suture and divided ventrally by incision into 2 shallow lobes interpreted to be precoxale and coxale (Figure 5f): precoxale with 3 or 4 plumose bristles; coxale with 4 bristles (3 plumose, 1 bare). Basale with 7 plumose bristles (6 on or near ventral margin, 1 near distodorsal corner). Endopodite well developed, with 5 long bristles (3 plumose, 2 bare). Exopodite 3-jointed: 1st joint with 2 or 3 long bare ventral bristles; 2nd joint with 3 bare bristles (2 ventral, 1 dorsal); 3rd joint with 3 bristles (dorsal and middle bristles claw-like (middle claw may have minute spines distally on ventral edge, but their presence could not be resolved with certainty); dorsal and ventral bristles bare). Protopodite and 1st exopodial joint with medial hairs. (Terminology of segments is that suggested by Angel and Iliffe, 1987:551.)

Seventh Limb (Figures 2b, 4g): Elongate with 3 long terminal bristles.

Furca (Figures 2b, 5g): Each lamella with 7 claws, all with marginal spines, and basal sutures; claw 1 with 4 weakly developed sutures in proximal half; posterior end of furca with 2 closely spaced bristles in place of “unpaired” bristle. Apron anterior to furca short, just reaching proximal anterior edge of furca.

Bellonci Organ (Figure 3a): Elongate, bifurcating just distal to midlength, with rounded tips.

Lips: Upper lip projecting posteriorly, spinous, with inward pointing spine on each side of medial line (Figure 4h,i). Lower lip with triangular process on each side of mouth (Figure 4j).

Copulatory Organ (Figure 5h–j): Consisting of 2 parts on left side of body. Posterior rod-shaped organ with very long styliform process with hirsute tip (Figure 5j). Anterior part broad with 2 or 3 toothed prongs at tip (Figure 5i).

Sperm Clusters (Figures 4k, 6): USNM 193440 with about 17 pear-shaped sperm clusters near copulatory limbs, each cluster containing abundant thread-like filaments. (Müller (1894, pl. 38: fig. 32) illustrated similar clusters in a testis of Conchoecia agassizii)

Gut Content: Gut of USNM 193440 with abundant thread-like filaments.

DESCRIPTION OF ADULT FEMALE (Figures 7, 8).—Shell decalcified and flexible, similar to that of adult male (Figure 7a).

Size: USNM 193442, length 0.82 mm, height 0.53 mm (because of film-like condition of decalcified shell, height dimension probably greater than when when animal was alive).

First Antenna (Figure 7b): Both limbs with only 1 ventral filament on 5th joint; limb otherwise similar to that of adult male.

Second Antenna (Figure 7c–g): Protopodite bare. Endopodite 3-jointed but 2nd and 3rd joints fused (Figure 7c–e); 1st joint with a slender a- and b-bristle; 2nd joint with stout f-bristle and shorter and more slender g-bristle (each with minute terminal papilla), and minute lateral peg near base of f-bristle (Figure 7e); 3rd joint with h-, i-, and j-bristles, each with terminal papilla; h-bristle with indistinct, minute, widely separated spines. Exopodite (Figure 7f,g); no spines observed on 1st joint, otherwise similar to that of adult male (as on adult male bristles of joints 1 and 2 with ventral spines in middle part, and bristle of 8th joint and 4 bristles of 9th joint with dorsal spines, and 1st joint divided into long proximal and short distal segments).

Mandible (Figure 7h,i): Similar to that of adult male.

Manxilla (Figure 7j,k): Not examined in detail but appearing similar to that of adult male.

Fifth Limb: Epipodite with bristles forming 3 groups, each with 4, 6, and 4 plumose bristles (Figure 7l). Protopodite without glandular process. Bristles of exopodial joints 2 and 3 similar to those of adult male. Bristles of protopodite and 1st exopodial joint not examined in detail but appearing similar to those of adult male.

Sixth Limb (Figure 7m,p): Epipodite with bristles forming 3 groups with 7 (6 long, 1 short and proximal), 5, and 4 or 5 plumose bristles (Figure 7m). Protopodite with 4 plumose bristles on precoxale and 5 on coxale. Remaining parts of limb similar to those of adult male.

Seventh Limb (Figure 7n,p): Similar to that of adult male.

Furca (Figure 7o): Similar to that of adult male except “unpaired” bristle bifurcate (not 2 separate bristles as on adult male).

Bellonci Organ (Figure 7b): Similar to that of adult male: tapering towards midlength then broadening and bifurcating, with broadly rounded tips and bare surface. Under oil immersion lens (×100), suggestion of suture at midlength where organ narrowest.

Lips (Figure 8): Similar to those of adult male.

Genitalia: Small process bearing terminal spine present on left side of body between epipodite of 6th limb and proximal end of 7th limb in vicinity of receptaculum seminis, which was not visible (Figure 7p).

Gut Content: Gut gorged with many crustacean-like claws, but not otherwise identified. (The contents of the gut suggest that the species is either a carnivore or scavenger.)

Maturity and Sex of Specimen: Not known with certainty. Two oval masses containing globules, both inside the posterior part of the body, could be eggs. The furca of the specimen bears 7 claws on each lamella, the same number as on the adult male, which indicates the specimen is either the adult or A-1 stage. The absence of a copulatory limb, as well as the lack of a convex dorsal margin on the endopodite of the 2nd antenna, both present on the A-1 male S. bermudensis described by Kornicker and Iliffe (1989b), indicate that the specimen is not male. The carapace of the specimen is shorter than those of the 2 adult males in the collection (0.82 mm for specimen, 0.98 mm for 2 adult males). The specimen bears a small spined process on the left side of the body between the epipodial appendage of the 6th limb and the 7th limb (Figure 7p); a similar process was found on the adult female S. bermudensis described by Kornicker and Iliffe (1989b), as well as on the adult female of S. sagax, new species, described herein. Because of the spined process, it is tentatively concluded that the specimen is an adult female.

COMPARISONS.—The carapace of S. styx differs from that of S. bermudensis in being smaller and in having the gland in the posterodorsal corner of the right valve on a small projecting process. The 1st antenna differs in not having ventral bristles on the 3rd and 4th joints; also, the dorsal bristle on the 2nd joint of styx differs from that of bermudensis in being diaphanous and shorter. The furca of styx differs from that of bermudensis in not having claw 2 on a stout protuberance. Known specimens of bermudensis have claw 2 of the furca broken off near its base, whereas all 3 specimens of S. styx in the present collection have claw 2 unbroken. Additional differences are listed in Table 3.

Spelaeoecia styx Kornicker, 1990

Spelaeoecia styx Kornicker in Kornicker et al., 1990:6, figs. 2–8.—Kornicker and Iliffe, 1998:26, figs. 17–29.

HOLOTYPE—USNM 193441, undissected adult male in alcohol.

TYPE LOCALITY.—El Dorado Cave, South Andros Island, Great Bahama Bank.

DISTRIBUTION.—South Andros Island: El Dorado Cave, Stargate Blue Hole. Exuma Cays: Norman's Pond Cave, Oven Rock Cave.

MATERIAL.—Stargate Blue Hole, South Andros Island: Sta 97-026: USNM 194544, adult male on slide and in alcohol; USNM 194546A,B, 2 adult females in alcohol; USNM 194546C,D, 2 adult males in alcohol; USNM 194546E, 1 A–1 ?female in alcohol; USNM 194546F, 1 A–2 instar in alcohol. Sta 97–029: USNM 194545A, 1 adult male in alcohol; USNM 194545B,C, 2 adult females in alcohol.

SUPPLEMENTARY DESCRIPTION OF ADULT MALE (Figure 14).—Carapace similar to that of holotype (Figure 14a).

Carapace Size (length, height in mm): USNM 194544, 1.03, 0.56; USNM 194545A, 1.04, 0.58; USNM 194546C, 1.04, 0.60; USNM 194546D, 1.03, 0.58.

First Antenna (Figure 14b,c), Second Antenna (Figure 14d,e), Mandible, and Maxilla (Figure 14f): Similar to those of holotype.

Fifth Limb (Figure 14f): Protopodite with elongate lateral glandular process projecting outward between maxilla and 6th limb.

Sixth Limb (Figure 14f), Seventh Limb: Similar to those of holotype.

Furca (Figure 14g): Each lamella with 7 claws.

Copulatory Organ (Figure 14h,i): Anterior branch complex, with proximal flat process with teeth along distal tip and distal toothed process; processes between flat process and distal toothed process difficult to interpret (Figure 14i).

SUPPLEMENTAL DESCRIPTION OF ADULT FEMALE.—Carapace Size (length, height in mm): Sta 97-026: USNM 194546A, 1.13, 0.57; USNM 194546B, 1.05, 0.56. Sta 97-029: USNM 194545B, 1.03, 0.56; USNM 194545C, 1.04, 0.52.

SUPPLEMENTAL DESCRIPTION OF A–1 INSTAR.—Carapace Size (length, height in mm): Sta 97-026, USNM 194546E (?female), 0.84, 0.50.

Furca: Each lamella with 7 claws.

SUPPLEMENTAL DESCRIPTION OF A–2 INSTAR.—Carapace Size (length, height in mm): Sta 97-026, 0.73, USNM 194546F, 0.41.

Furca: Each lamella with 6 claws followed by small triangular process (incipient claw).

REMARKS CONCERNING CARAPACE SURFACE.—The carapace of an adult male of S. styx that had been collected on Exuma Cay (Kornicker and Iliffe, 1998) and had distinct epicuticle reticulations prior to being immersed in glycerine was examined with a Scanning Electron Microscope; most reticulations were no longer clearly visible, but a few remained. The carapace of a specimen of S. styx that had been collected in Andros Island and then immersed in glycerine, which had resulted in the reticulations being no longer visible, also was examined with a Scanning Electron Microscope, and a few reticulations were found to be present. The reticulations from both localities seem sufficiently similar to indicate a close relationship between the two populations. The specimens were treated by freezing point evaporation prior to micrography.

An adult female (USNM 194298) and an A–1 male (USNM 194414) from Exuma Cays, whose reticulations had disappeared after immersion in glycerin, were placed in dyes (Chlorozol Black and Hematoxylin, respectively), but these failed to bring out surface ridges. Fresh material from Andros Island should be examined to be certain that the carapace reticulations are similar to the carapace reticulations of the population from Exuma Cays.

COMPARISONS.—As discussed in Kornicker and Iliffe (1998:37), the 2nd joint of the 1st antennae of Andros Island specimens have a shorter dorsal bristle (Figure 14b,c) than the specimens from Exuma Cays, suggesting that the Andros Island and Exuma Cays populations may not be conspecific. Although the specimens are too few for statistical certainty, the carapace lengths of specimens from the four localities appear to vary (Table 6).

Spelaeoecia styx Kornicker, 1990

Spelaeoecia styx Kornicker in Kornicker et al., 1990:6, figs. 2–8.

HOLOTYPE.—USNM 194270, undissected adult male in alcohol.

TYPE LOCALITY.—El Dorado Cave, South Andros Island, Great Bahama Bank.

MATERIAL.—Norman's Pond Cave. Norman's Pond Cay, Exuma Cays: Sta 93-001: USNM 194266, undissected adult female in alcohol; USNM 194295A,B, 2 undissected A–3 instars in alcohol; USNM 194295C, undissected A–4 instar in alcohol. Sta 93-002: USNM 194270, undissected adult male in alcohol; USNM 194276, undissected adult male (lost); USNM 194273, partly dissected A–1 male in alcohol; USNM 194275, undissected A–2 instar (sex unknown) in alcohol; USNM 194274, partly dissected A–3 instar (sex unknown) in alcohol; USNM 194272, undissected A–4 instar (sex unknown) in alcohol; USNM 194271, partly dissected A–5 instar (sex unknown) in alcohol; USNM 194296A–C, 3 undissected adult females in alcohol; USNM 194296D, undissected adult male in alcohol; USNM 194296E–G, 3 undissected A–1 females in alcohol; USNM 194296H, undissected A–2 instar (sex unknown) in alcohol; USNM 194296J,K, 2 undissected A–3 instars (sex unknown) in alcohol; USNM 194300, partly dissected adult male on slide and in alcohol. Sta 93–003: USNM 194260, adult male on slide and in alcohol; USNM 194261, adult female on slide and in alcohol; USNM 194325A, B, 2 undissected adult males in alcohol; USNM 194325C, undissected adult female in alcohol; USNM 194325D, undissected A–2 instar (sex unknown) in alcohol; USNM 194326, undissected A–1 female in alcohol; USNM 194327, 2 undissected A–4 instars (sex unknown) in alcohol. Sta 93-004: USNM 194285, A–3 instar (sex unknown) on slide and in alcohol. Sta 94–016: USNM 194434A, 2 undissected A–4 instars (sex unknown) and 2 undissected A–5 instars (sex unknown) in alcohol; USNM 194434B, 2 undissected A–3 instars (sex unknown) and 3 undissected A–2 instars (sex unknown) in alcohol; USNM 194434C, 3 undissected A–2 instars (sex unknown) and 1 undissected A–1 female in alcohol; USNM 194434D, 4 undissected A–1 females in alcohol; USNM 194434E, 4 undissected adult males and 4 undissected adult females in alcohol; USNM 194434F, 3 undissected adult males, 3 undissected adult females, and an empty carapace, all in alcohol. Oven Rock Cave, Great Guana Cay, Exuma Cays: Sta 93-006: USNM 194278, undissected A–3 instar (sex unknown) in alcohol; USNM 194297A, adult male on slide and in alcohol; USNM 194297B, undissected A–1 male in alcohol; USNM 194297C,D, 2 undissected A–1 females in alcohol; USNM 194297E, undissected A–2 instar (sex unknown) in alcohol. Sta 93–008: USNM 194298, undissected adult female in alcohol. Sta 93-009: USNM 194277, partly dissected A–5 instar in alcohol; USNM 194299A, undissected adult female in alcohol; USNM 194299B, undissected A–1 female in alcohol. Sta 94-014: USNM 194414, undissected A–1 male in alcohol; USNM 194415, undissected A–5 instar in alcohol. Sta 95-012: USNM 194450A, undissected adult male in alcohol; USNM 194450B,C, 2 undissected adult females in alcohol; USNM 194450D,E, 2 undissected A–1 females in alcohol; USNM 194450F, undissected A–2 instar (sex unknown) in alcohol; USNM 194450G, undissected A–1 male in alcohol. El Dorado Cave, South Andros Island, paratype, USNM 193440, adult male.

DISTRIBUTION.—Exuma Cays, Great Bahama Bank: Norman's Pond Cave, Norman's Pond Cay (Sta 93–001, 93-002, 93-003, 94-016) from water column at depths of 6–35 m and from fine silt on ledge at depth of 6–18 m, salinity 35–36 ppt. Oven Rock Cave, Great Guana Cay (Sta 93-006, 93-008, 93-009, 95-012) from water column at depths of 0–22 m and from dark brown silt on floor at 8 m, salinity 35–36 ppt. South Andros Island, Great Bahama Bank, El Dorado Cave (Kornicker et al., 1990:2).