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Taxonomic History

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Mystrium oberthueri Forel, 1897d PDF: 192 (w.) MADAGASCAR. Malagasy. Primary type information: Madagascar, Insel Ste. Marie (Island of Sainte Marie), coll. Perrot; CASENT0101993; MHNG AntCat AntWiki HOL

Taxonomic history

Lectotype designation: Yoshimura & Fisher, 2014 PDF: 85.[Mystrium oberthueri Wasmann, 1897a PDF: 250. Nomen nudum (attributed to Forel).].Emery, 1899e PDF: 267 (m.); Yoshimura & Fisher, 2014 PDF: 85 (w.eq.m.).Status as species: Emery, 1899e PDF: 267; Emery, 1911e PDF: 23; Wheeler, 1922: 1006; Menozzi, 1929e PDF: 530 (redescription); Brown, 1960a: 170; Bolton, 1995b: 287; Yoshimura & Fisher, 2014 PDF: 85 (redescription).
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California Academy of Sciences
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AntWeb. Version 8.45.1. California Academy of Science, online at https://www.antweb.org. Accessed 15 December 2022.
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Diagnostic Description

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Il Sig. Mocquerys ha raccolto una [[ worker ]] di questa specie e un [[ male ]] che credo dovere riferire alla medesima. Differisce dal [[ male ]] di M. mysticum per i punti seguenti: la grandezza minore, il torace piu largo, le antenne molto piu corte, col 2. ° articolo (1. ° del funicolo) brevissimo, piu corto che largo, lo scapo che non raggiunge il livello dell'ocello impari, le ali piu chiare e particolarmente la scultura. Le fossette piligere sono piu stipate che nel M. mysticum sul capo e sul torace; su tutti i segmenti dorsali e ventrali dell'addome, serbano l'aspetto di buchi tondi e profondi, opachi, mentre nel M. mysticum sono ridotti su questa parte del corpo a infossature superficiali, appena sensibili sui segmenti posteriori. Il clipeo, fittamente coperto di fossette, ha il margine distintamente dentellato. L'addome e strozzato dietro ciascun segmento. Sul torace, le linee parassidiali sono marcate.

L. 7 mm. Antenne circa 3 mm. (nel M. mysticum [[ male ]], le antenne misurano oltre 4 mm.).

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Emery, C., 1899, Formiche di Madagascar raccolte dal Sig. A. Mocquerys nei pressi della Baia di Antongil (1897-1898)., Bollettino della Societa Entomologica Italiana, pp. 263-290, vol. 31
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Emery, C.
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Diagnostic Description

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[[ worker ]] minor L. 6,8 bis 7 mm ohne, 8,3 bis 8,5 mm mit Oberkiefer. Oberkiefer kuerzer als bei voeltzkowi , ungefaehr wie bei mysticum , aber fast gerade, gleichmaessig schwach, nach aussen konvex und nach innen konkav, an der Spitze kaum breiter, stumpf abgestutzt, mit zwei Reihen von je ca. 10 kleinen Zaehnen, die etwa so stark sind wie bei mysticum . Der Oberkiefer an der Basis mindestens so dick (sogar etwas dicker) als an der Spitze, sehr fein genetzt und matt.

Kopf wie bei voeltzkowi , viel breiter als lang, sogar noch breiter im Verhaeltnis zur Laenge, hinten weniger verengt als bei genannter Art, vorn seitlich mit zwei sehr dicken und viel laengeren Zaehnen als bei voeltzkowi [[ worker ]] minor. Augen ziemlich konvex, etwas hinter der Mitte. Clypeusrand schaerfer gezaehnt als bei mysticum , aber kuerzer als bei voeltzkowi . Der Fuehlerschaft erreicht gerade den Hinterhauptrand. Geisselglieder etwas laenger als bei mysticum , cylindrischer und kuerzer als bei voeltzkowi , mit Ausnahme des langen zweiten Gliedes. Thorax, Stielchen und Hinterleib so geformt wie bei mysticum , aber etwas schmaeler und laenglicher, Thorax staerker eingeschnuert, Stielchen etwas laenger im Verhaeltnis zur Breite (etwas mehr als doppelt so breit als lang).

Matt. Kopf sehr grob genetzt, mit Tendenz zur Laengsrunzelung, besonders auf der Stirne, Clypeus mit feiner Skulptur, sowie die abschuessige Flaeche des Metanotum, die Vorderflaeche des Stielchens und die drei letzten Hinterleibssegmente. Thorax, Stielchen und erstes Hinterleibssegment sehr grob und regelmaessig laengsgerunzelt, (Metanotum, Stielchen und erstes Hinterleibssegment fast gestreift). Zweites Hinterleibssegment fein laengsgerunzelt. Ueberall ist die feine Grundskulptur unregelmaessig, sehr fein genetzt-gerunzelt (am Hinterleib punktiert-genetzt).

Behaarung zerstreut, aeusserst kurz, dick, ueberall ziemlich anliegend, aber nirgends deutlich kolbig oder loeffeifoermig, wodurch diese Art von allen anderen abweicht. An den Beinen ist die Behaarung reichlicher und dicker.

Schwarz. Fuehlerkeule, erstes Geisselglied, Spitze der Mandibeln, Stirnleisten, Tarsen, Gelenke und Segmentraender roetlichgelb oder rotbraun, Fuehler, Beine und Basis der Mandibeln im uebrigen braun.

[[ worker ]] major: L. ohne Mandibeln 9,7 mm, mit den Mandibeln 12,7 mm. Mandibeln wie bei M. voeltzkowi , [[ worker ]] major, viel laenger als beim [[ worker ]] minor und mit staerker nach innen konkaver Endhaelfte, mit viel staerkeren Zaehnen (die Zaehne aber stumpfer und kuerzer als beim voeltzkowi [[ worker ]] major). Die Vorderecken mit je einem langen, spitzen und gekruemmten Dorn, der dreimal so lang ist als die Breite seiner Basis und doppelt so lang wie bei voeltzkowi [[ worker ]] major. Das 3. Abdominalsegment ist noch gestreift; Kopf querrechteckig, sonst wie der [[ worker ]] minor.

Durch die laugen Kopfdornen des [[ worker ]] major, durch die Skulptur und durch die Behaarung von allen andern Arten verschieden, dem voeltzkowi am naechsten stehend, aber groesser und weniger schlank.

Insel Ste. Marie de Madagascar. Aus der Sammlung des Herrn R. Oberthuer, von Herrn Perrot gesammelt, durch Herrn Prof. Wasmann erhalten.

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Forel, A., 1897, Ameisen aus Nossi-Be, Majunga, Juan de Nova (Madagaskar), den Aldabra-Inseln und Sansibar. Gesammelt von Herrn Dr. A. Voeltzkow aus Berlin., Abhandlungen herausgegeben von der Senckenbergischen Naturforschenden Gesellschaft, pp. 185-208, vol. 21
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Forel, A.
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Description

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Worker. Description. Measurements: lectotype. HL 2.40, HW 2.61, SL 1.80, ML 2.92, HD 1.67, WL 2.79, PnW 1.39, PpW 1.21, PtW 1.18, PtL 0.81, CI 108.7, SI 69.0, MI 111.8, PpI 87.4, PtI 144.9. HL 2.00–2.58, HW 2.12–2.76, SL 1.55–1.96, ML 2.39–3.17, HD 1.33–1.69, WL 2.43–2.96, PnW 1.17–1.38, PpW 1.02–1.29, PtW 1.00–1.32, PtL 0.67–0.81, CI 104.4–108.5, SI 68.8–74.9, MI 106.9–122.9, PpI 87.2–93.8, PtI 139.5–163.2 (10 specimens measured). Posterolateral corner of head moderately expanding posteriorly. Posterior face of vertex forming almost right angle with dorsal face on median line of head, so that declivity of vertex on lateral part as steep as on median part. Whole region of vertex finely striated. Eye moderately small. Anterior margin of clypeus straight or weakly convex with long conical setae. Genal tooth of head relatively long, as long as lateral lobe of clypeus. Masticatory surface of mandible almost invisible in full-face view, and width of dorsal surface of mandible almost identical from mandibular shaft to distal portion. Second maxillary palpomere longer than third. First flagellomere (third antennal segment) about 1.2–1.5× length of pedicel (second antennal segment). Strong, deep longitudinal striae regularly impressed on whole central part of pronotal dorsum. Strong, deep longitudinal striae impressed on lateral surface of pronotum. Mesonotum differentiated from propodeum in dorsal view, length slightly shorter than that of propodeum. Metanotal groove shallowly and gently impressed in lateral view. Metapleural gland bulla moderately developed, and propodeal declivity in lateral view almost straight. Petiole widened on posterior 1/3 and gently narrowing anteriorly in dorsal view, anterior margin straight to gently rounded and often edged by thick striae. Body color black. Ergatoid queen. Description. Measurements: HL 1.40–1.93, HW 1.46–1.96, SL 1.07–1.47, ML 1.49–2.00, HD 1.00–1.35, WL 1.92–2.56, PnW 0.83–1.11, PpW 0.85–1.13, PtW 0.81–1.07, PtL 0.53–0.70, CI 101.6–108.6, SI 71.6–76.7, MI 96.3–108.8, PpI 96.0–104.4, PtI 149.1–177.1 (10 specimens measured). Wings vestigial and reduced to small but distinct appendages. Wing sclerites undeveloped. Posterolateral corner of head moderately expanding posteriorly, expansion not differentiated from that in workers. Posterior face of vertex forming almost a right angle with dorsal face on median line of head, so that declivity of vertex on lateral part as steep as on median part. Ventral half of vertex sculptured. Eye small but distinct. Ocelli absent. Anterior margin of clypeus almost straight with long to moderate conical setae. Anterolateral portion of head with short spine. Masticatory margin of mandible almost invisible in full-face view, and dorsal surface on distal portion as wide as that on mandibular shaft. Spatulate seta present on basal side of each basal denticle on masticatory margin of mandible. First flagellomere (third antennal segment) long, about 1.2-1.5× length of pedicel (second antennal segment). Setae on pronotum distinctly spatulate, widened distally with sharp or blunt apex. Metapleural gland bulla moderately developed, not expanding dorsally to propodeal spiracle, so that propodeal declivity in lateral view weakly convex and rounded posteriorly on its ventral 1/3. Petiole relatively long in dorsal view, about 0.8× length of abdominal segment III. Body color blackish to reddish brown. Male. Description. Measurements: HL 1.44, HW 2.09, SL 0.62, EL 0.95, WL 3.55, MnW 1.94, CI 145.1, SI 29.6, EI 66.2, MnI 92.9 (1 specimen measured). Eye moderately large, occupying 0.6× of head length. Ocelli relatively distant from dorsal margin of head in full-face view. Dorsal margin of head in full-face view straight. Both anterior and lateral ocelli small. Lateral ocellus small and distant from eye: distance between these more than 1.5× maximum diameter of lateral ocellus. Posterior face of vertex not clearly differentiated from dorsal face, so that vertex almost continuously rounded. Palpal formula 4,3. First maxillary palpomere flattened and distinctly wider than second segment. Second maxillary palpomere longer than third. Notauli clearly impressed on mesoscutum. Petiole in dorsal view thin, 0.8× as long as abdominal tergite III. Petiolar dorsum covered with shallow, irregular punctures. Abdominal tergum VIII without deep punctures, almost smooth. Distal portion of abdominal sternum IX smooth and not punctured. Basal ring moderately long, expanding basoventrally. Telomere distinctly extending distally farther than digitus. Basoventral expansion of aedeagus moderately developed basoventrally, longer than dorsal extension. Ventral margin of aedeagus gently convex in lateral view. Aedeagus moderately narrowing distally and distal portion rounded. On forewing, cu-a located far basal from junction of Media (M) and Cubitus (Cu). Body color reddish brown to black.
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Masashi Yoshimura, Brian L. Fisher
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Yoshimura M, Fisher B (2014) A revision of the ant genus Mystrium in the Malagasy region with description of six new species and remarks on Amblyopone and Stigmatomma (Hymenoptera, Formicidae, Amblyoponinae) ZooKeys 394: 1–99
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Masashi Yoshimura
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Brian L. Fisher
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Distribution

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MADAGASCAR: as in Figure 56D.
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Masashi Yoshimura, Brian L. Fisher
bibliographic citation
Yoshimura M, Fisher B (2014) A revision of the ant genus Mystrium in the Malagasy region with description of six new species and remarks on Amblyopone and Stigmatomma (Hymenoptera, Formicidae, Amblyoponinae) ZooKeys 394: 1–99
author
Masashi Yoshimura
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Brian L. Fisher
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