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Gordon's Mockorange

Philadelphus lewisii Pursh

Common Names

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Lewis' mock orange
mockorange
syringa
Gordon's mockorange
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Carey, Jennifer H. 1995. Philadelphus lewisii. In: Fire Effects Information System, [Online]. U.S. Department of Agriculture, Forest Service, Rocky Mountain Research Station, Fire Sciences Laboratory (Producer). Available: http://www.fs.fed.us/database/feis/

Cover Value

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More info for the terms: cover, cover type, shrub

Lewis' mock orange occurs in dense shrub habitats which probably provide
good cover for wildlife. In north-central Washington, Lewis' mock orange
occurs in a riparian cover type which is preferred in both summer and
winter by mule deer for thermal and security cover [6].
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Carey, Jennifer H. 1995. Philadelphus lewisii. In: Fire Effects Information System, [Online]. U.S. Department of Agriculture, Forest Service, Rocky Mountain Research Station, Fire Sciences Laboratory (Producer). Available: http://www.fs.fed.us/database/feis/

Description

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More info for the terms: capsule, fruit, shrub

Lewis' mock orange is a native, deciduous, erect to spreading shrub that
grows 3 to 10 feet (1-3 m) tall. The showy flowers occur in clusters of
three to fifteen. The fruit is a four-chambered capsule about 0.24 to
0.39 inch (0.6-1 cm) long [34]. Seeds are about 0.08 inch (0.2 cm)
long. Lewis' mock orange is extremely variable in both vegetative and
floral characteristics [39].
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bibliographic citation
Carey, Jennifer H. 1995. Philadelphus lewisii. In: Fire Effects Information System, [Online]. U.S. Department of Agriculture, Forest Service, Rocky Mountain Research Station, Fire Sciences Laboratory (Producer). Available: http://www.fs.fed.us/database/feis/

Distribution

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Lewis' mock orange occurs in the northwestern United States and southern
Canada. It occurs from extreme southern British Columbia south to
California, and east to north and central Idaho, western Montana, and
southwestern Alberta [14,15,34]. Philadelphus lewisii ssp. californicus
occurs from the southern Cascade Range of southwestern Oregon south
through the Sierra Nevada to Tulare County, California [14,32], and P.
l. var. gordonianus occurs in the Coast Ranges and the Cascade Range
from British Columbia south to northern California [43].



Distribution of Lewis' mock orange. Map courtesy of USDA, NRCS. 2018. The PLANTS Database.
National Plant Data Team, Greensboro, NC [44].

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Carey, Jennifer H. 1995. Philadelphus lewisii. In: Fire Effects Information System, [Online]. U.S. Department of Agriculture, Forest Service, Rocky Mountain Research Station, Fire Sciences Laboratory (Producer). Available: http://www.fs.fed.us/database/feis/

Fire Ecology

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More info for the terms: fire regime, top-kill

Lewis' mock orange occurs in both forested environments which frequently
experience fire and on rocky scree slopes which may not burn at all. In
western Montana, Lewis' mock orange occurs in Douglas-fir habitat types
which had historical fire frequencies of 5 to 45 years [10]. The
ability of Lewis' mock orange to sprout after top-kill by fire enables it
to persist in these forests.

FIRE REGIMES :
Find fire regime information for the plant communities in which this
species may occur by entering the species name in the FEIS home page under
"Find FIRE REGIMES".
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Carey, Jennifer H. 1995. Philadelphus lewisii. In: Fire Effects Information System, [Online]. U.S. Department of Agriculture, Forest Service, Rocky Mountain Research Station, Fire Sciences Laboratory (Producer). Available: http://www.fs.fed.us/database/feis/

Fire Management Considerations

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More info for the term: fuel

Lewis' mock orange palatability increases following fire. After a spring
fire in northern Idaho, utilization of Lewis' mock orange by Rocky
Mountain elk was significantly (p less than 0.05) greater on recently burned sites
than on adjacent unburned sites [21].

% of available twigs browsed

1st season 2nd season

Control 1.3 0.6
Burn 36.3 30.0

Allometric equations, which can be used to estimate fuel quantities,
have been developed for Lewis' mock orange. The equations use basal stem
diameter to estimate foliage biomass and total biomass [5].
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bibliographic citation
Carey, Jennifer H. 1995. Philadelphus lewisii. In: Fire Effects Information System, [Online]. U.S. Department of Agriculture, Forest Service, Rocky Mountain Research Station, Fire Sciences Laboratory (Producer). Available: http://www.fs.fed.us/database/feis/

Growth Form (according to Raunkiær Life-form classification)

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More info for the term: phanerophyte

Phanerophyte
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bibliographic citation
Carey, Jennifer H. 1995. Philadelphus lewisii. In: Fire Effects Information System, [Online]. U.S. Department of Agriculture, Forest Service, Rocky Mountain Research Station, Fire Sciences Laboratory (Producer). Available: http://www.fs.fed.us/database/feis/

Habitat characteristics

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Lewis' mock orange occurs on well-drained, moist sites [46]. It grows on
deep, rich alluvial loams to rocky or gravelly loams [43]. Lewis'
mock orange is commonly found on rocky sites, at the base of talus slopes
and cliffs, along streams, and in seasonally moist draws [6,15]. It
is found at talus margins in the Columbia River Basin [11]. It occurs
at seeps, springs, and rocky wet areas in the Crooked River National
Grasslands in central Oregon [16].

Lewis' mock orange occurs from sea level up to 7,000 feet (2,100 m) in
the Cascade Range [15,43]. Philadelphus lewisii ssp. californicus grows
from 1,000 to 5,000 feet (300-1,500 m) elevation on rocky slopes and in
canyons in the Sierra Nevada [36]. Lewis' mock orange grows best on
northern and eastern exposures [43].
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bibliographic citation
Carey, Jennifer H. 1995. Philadelphus lewisii. In: Fire Effects Information System, [Online]. U.S. Department of Agriculture, Forest Service, Rocky Mountain Research Station, Fire Sciences Laboratory (Producer). Available: http://www.fs.fed.us/database/feis/

Habitat: Cover Types

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This species is known to occur in association with the following cover types (as classified by the Society of American Foresters):

210 Interior Douglas-fir
212 Western larch
213 Grand fir
219 Limber pine
221 Red alder
222 Black cottonwood-willow
227 Western redcedar-western hemlock
229 Pacific Douglas-fir
230 Douglas-fir-western hemlock
233 Oregon white oak
234 Douglas-fir-tanoak-Pacific madrone
237 Interior ponderosa pine
244 Pacific ponderosa pine-Douglas-fir
245 Pacific ponderosa pine
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bibliographic citation
Carey, Jennifer H. 1995. Philadelphus lewisii. In: Fire Effects Information System, [Online]. U.S. Department of Agriculture, Forest Service, Rocky Mountain Research Station, Fire Sciences Laboratory (Producer). Available: http://www.fs.fed.us/database/feis/

Habitat: Ecosystem

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This species is known to occur in the following ecosystem types (as named by the U.S. Forest Service in their Forest and Range Ecosystem [FRES] Type classification):

More info for the term: shrub

FRES20 Douglas-fir
FRES21 Ponderosa pine
FRES22 Western white pine
FRES23 Fir-spruce
FRES25 Larch
FRES28 Western hardwoods
FRES29 Sagebrush
FRES34 Chaparral-mountain shrub
FRES36 Mountain grasslands
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Carey, Jennifer H. 1995. Philadelphus lewisii. In: Fire Effects Information System, [Online]. U.S. Department of Agriculture, Forest Service, Rocky Mountain Research Station, Fire Sciences Laboratory (Producer). Available: http://www.fs.fed.us/database/feis/

Habitat: Plant Associations

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This species is known to occur in association with the following plant community types (as classified by Küchler 1964):

More info for the terms: forest, shrub

K010 Ponderosa shrub forest
K011 Western ponderosa forest
K012 Douglas-fir forest
K013 Cedar-hemlock-pine forest
K014 Grand fir-Douglas-fir forest
K025 Alder-ash forest
K026 Oregon oakwoods
K028 Mosaic of K002 and K026
K034 Montane chaparral
K051 Wheatgrass-bluegrass
K055 Sagebrush steppe
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bibliographic citation
Carey, Jennifer H. 1995. Philadelphus lewisii. In: Fire Effects Information System, [Online]. U.S. Department of Agriculture, Forest Service, Rocky Mountain Research Station, Fire Sciences Laboratory (Producer). Available: http://www.fs.fed.us/database/feis/

Habitat: Rangeland Cover Types

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This species is known to occur in association with the following Rangeland Cover Types (as classified by the Society for Range Management, SRM):

More info for the terms: shrubland, woodland

109 Ponderosa pine shrubland
203 Riparian woodland
209 Montane shrubland
421 Chokecherry-serviceberry-rose
422 Riparian
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Carey, Jennifer H. 1995. Philadelphus lewisii. In: Fire Effects Information System, [Online]. U.S. Department of Agriculture, Forest Service, Rocky Mountain Research Station, Fire Sciences Laboratory (Producer). Available: http://www.fs.fed.us/database/feis/

Immediate Effect of Fire

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More info for the term: root crown

Lewis' mock orange is top-killed by fire, but the root crown usually
survives and produces sprouts [10,23,24].
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Carey, Jennifer H. 1995. Philadelphus lewisii. In: Fire Effects Information System, [Online]. U.S. Department of Agriculture, Forest Service, Rocky Mountain Research Station, Fire Sciences Laboratory (Producer). Available: http://www.fs.fed.us/database/feis/

Importance to Livestock and Wildlife

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Lewis' mock orange is of minor importance as a forage species for
livestock [43]. Sampson [36] considered it poor to useless browse for
cattle and horses. However, Shaw [37] reports that in riparian areas in
eastern Oregon, Lewis' mock orange is heavily browsed in areas accessible
to cattle.

Lewis' mock orange is a moderately important winter forage species for
deer and elk in the northern Rocky Mountains. In southern British
Columbia, Lewis' mock orange is of moderate importance as a winter forage
species for white-tailed deer and Rocky Mountain elk, and of low
importance to other wild ungulates [4]. In Montana, a 1957 study based
on rumen samples showed that Lewis' mock orange constituted 2 percent of
mule deer diets in the winter and a trace in the summer [47]. In
northern Idaho, use by white-tailed deer was moderate, although a few
individual plants were browsed heavily [42]. Lewis' mock orange seeds
are eaten by quail and squirrels [46].
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bibliographic citation
Carey, Jennifer H. 1995. Philadelphus lewisii. In: Fire Effects Information System, [Online]. U.S. Department of Agriculture, Forest Service, Rocky Mountain Research Station, Fire Sciences Laboratory (Producer). Available: http://www.fs.fed.us/database/feis/

Key Plant Community Associations

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More info for the terms: forest, shrub, shrubs

Lewis' mock orange commonly occurs in open coniferous forests and at
forest edges [34], and is usually associated with other shrubs. In dry
Douglas-fir (Pseudotsuga menziesii) forests on the western slope of the
Cascade Range in Oregon, Lewis' mock orange is positively (significant at
P less than 0.05) associated with beaked hazelnut (Corylus cornuta), oceanspray
(Holodiscus discolor), poison-oak (Toxicodendron diversilobum), baldhip
rose (Rosa gymnocarpa), and hollyleaved barberry (Mahonia aquifolium).
It reaches its greatest importance in the Douglas-fir/hollyleaved
barberry/disporum (Disporum spp.) community type [26]. Lewis'
mock orange is positively (significant at P less than 0.05) associated with
ninebark (Physocarpus malvaceus) on talus slopes in western Montana [25].

Lewis' mock orange occurs in moist draws and riparian areas, especially
in drier regions of the Northwest. In eastern Oregon, Lewis' mock orange
is associated with willows (Salix spp.), alders (Alnus spp.), and
hawthorns (Crataegus spp.) [13]. It occurs with red-osier dogwood
(Cornus sericea) and Saskatoon serviceberry (Amelanchier alnifolia) in
north-central Washington [6]. It occurs with Saskatoon serviceberry and
common chokecherry (Prunus virginiana) on rocky outcrops of the National
Bison Range in western Montana [29]. In Oregon white oak (Quercus
garryana) woodlands in northern California, Lewis' mock orange occurs
along stream channels with oceanspray, cluster rose (R. pisocarpa), pale
serviceberry (A. pallida), and Indian plum (Oemleria cerasiformis). A
Lewis' mock orange/brittle bladderfern (Cystopteris fragilis) community
type has been described [41].

Lewis' mock orange occurs in seral shrubfields and chaparral communities.
In northern Idaho, Lewis' mock orange is a component of the tall shrub
union that follows logging and burning [48]. In southwestern Oregon,
Philadelphus lewisii ssp. californicus occurs in chaparral dominated by
wedgeleaf ceanothus (Ceanothus cuneatus). Other shrubs present include
skunkbush sumac (Rhus trilobata), yerba santa (Eriodictylon
californicum), chaparral honeysuckle (Lonicera interrupta), Klamath plum
(P. subcordata), hollyleaf redberry (Rhamnus crocea ssp. ilicifolia),
and pale serviceberry [7].
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Carey, Jennifer H. 1995. Philadelphus lewisii. In: Fire Effects Information System, [Online]. U.S. Department of Agriculture, Forest Service, Rocky Mountain Research Station, Fire Sciences Laboratory (Producer). Available: http://www.fs.fed.us/database/feis/

Life Form

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More info for the term: shrub

Shrub
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Carey, Jennifer H. 1995. Philadelphus lewisii. In: Fire Effects Information System, [Online]. U.S. Department of Agriculture, Forest Service, Rocky Mountain Research Station, Fire Sciences Laboratory (Producer). Available: http://www.fs.fed.us/database/feis/

Management considerations

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Lewis' mock orange is very sensitive to herbicide sprays [2,31]. Effects
of herbicides on Lewis' mock orange are described [27].
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bibliographic citation
Carey, Jennifer H. 1995. Philadelphus lewisii. In: Fire Effects Information System, [Online]. U.S. Department of Agriculture, Forest Service, Rocky Mountain Research Station, Fire Sciences Laboratory (Producer). Available: http://www.fs.fed.us/database/feis/

Occurrence in North America

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CA ID MT OR WA AB BC
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Carey, Jennifer H. 1995. Philadelphus lewisii. In: Fire Effects Information System, [Online]. U.S. Department of Agriculture, Forest Service, Rocky Mountain Research Station, Fire Sciences Laboratory (Producer). Available: http://www.fs.fed.us/database/feis/

Other uses and values

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Native Americans used the strong, hard branches of Lewis' mock orange for
bows, arrows, combs, tobacco pipes, cradles, and netting shuttles
[17,34]. Lewis' mock orange is cultivated as an ornamental, but
Philadelphus coronarius, a European species, is the most commonly grown
mock orange in the Northwest [39]. Lewis' mock orange is the Idaho state
flower [17]; it is illegal to collect Lewis' mock orange in Idaho for
export or sale [30].
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Carey, Jennifer H. 1995. Philadelphus lewisii. In: Fire Effects Information System, [Online]. U.S. Department of Agriculture, Forest Service, Rocky Mountain Research Station, Fire Sciences Laboratory (Producer). Available: http://www.fs.fed.us/database/feis/

Palatability

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Although generally considered of low palatability, Lewis' mock orange is
browsed heavily at times [21,37,42,43]. New sprouts of Lewis'
mock orange are very palatable [1,21,36]. See FIRE MANAGEMENT for
discussion of Lewis' mock orange palatability following fire.
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Carey, Jennifer H. 1995. Philadelphus lewisii. In: Fire Effects Information System, [Online]. U.S. Department of Agriculture, Forest Service, Rocky Mountain Research Station, Fire Sciences Laboratory (Producer). Available: http://www.fs.fed.us/database/feis/

Phenology

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More info for the term: fruit

Flowering of Lewis' mock orange occurs from May through July. Fruit
matures in late summer and seeds are dispersed in September or October
[39]. The following dates are general ranges drawn from several studies
in northern Idaho. Specific times for several years are reported [8,33].

Development Date of Occurrence

bud burst early April
leaf out late April to early May
leaf growth late April to mid-May
stem elongation early May to late May
flower bloom late June to July
fruit development begins in July
leaf fall late September to late November
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Carey, Jennifer H. 1995. Philadelphus lewisii. In: Fire Effects Information System, [Online]. U.S. Department of Agriculture, Forest Service, Rocky Mountain Research Station, Fire Sciences Laboratory (Producer). Available: http://www.fs.fed.us/database/feis/

Plant Response to Fire

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More info for the terms: habitat type, herbaceous, root crown, shrubs

Lewis' mock orange sprouts vigorously from the root crown following fire
[10,23,24]. After an April fire on a south-facing slope in
north-central Idaho, Lewis' mock orange increased to prefire densities by
the third postfire growing season [22].

The following two studies have investigated the sprouting response of
Lewis' mock orange to fire.

Seral brushfields within the grand fir (Abies grandis)/pachistima
(Pachistima myrsinites) habitat type in northern Idaho were burned in
either spring (late March - early April) or fall (October).
Temperatures during the fires ranged from 67 to 78 degrees Fahrenheit
(19-26 deg C), and relative humidities ranged from 25 to 48 percent.
Lewis' mock orange on sites burned in October did not sprout until the
following April. Lewis' mock orange on the spring-burned sites sprouted
4 to 8 weeks after the fire. Twenty completely top-killed Lewis'
mock orange (eleven on the fall burned sites and nine on the spring
burned sites) were measured at the end of the first postfire growing
season [24]:

Fall Fires Spring Fires

Average crown diameter (ft)
prefire 4.7 (143 cm) 4.7 (143 cm)
postfire 2.4 (73 cm) 2.2 (67 cm)

Average crown height (ft)
prefire 8.8 (268 cm) 9.6 (293 cm)
postfire 4.1 (125 cm) 3.7 (113 cm)

No. basal sprouts per plant
prefire 1.5 0.6
postfire 38.0 28.9

Average postfire sprout height (ft) 2.4 (73 cm) 2.1 (64 cm)

A multiple regression equation is presented which relates the number of
postfire basal sprouts to prefire crown height, crown diameter, and
crown volume [24].

In another study in north-central Idaho, a brushfield was burned three
times at 5-year intervals (31 March 1965, 3 May 1970, and 14 May 1975).
Maximum air temperatures during the fires were 77, 81, and 88 degrees
Fahrenheit (25, 27, and 31 deg C), and relative humidities at 4:00 pm
were 35, 16, and 27 percent, respectively. Leaves on shrubs and
succulent herbaceous growth depressed the fire in 1975. A single Lewis'
mock orange was followed during the study. The plant was dormant during
the first fire, but leaves were beginning to emerge at the time of the
second fire and were completely emerged at the time of the third fire.
With each successive fire, average sprout height decreased. Reduced
growth following the second and third fires may have resulted from the
advanced phenological stage at the time of those fires. Sprout height
and number were measured the first growing season following each fire.
Crown height and diameter were measured during the second postfire
growing season [23].

No. basal sprouts Average sprout height (ft)
1965 14 2.0 (61 cm)
1970 19 1.5 (46 cm)
1975 16 1.0 (30 cm)


Crown height (ft) Crown diameter (ft)
prefire 7.0 (213 cm) 1.5 (46 cm)

1966 4.0 (122 cm) 2.0 (61 cm)
1971 2.5 (76 cm) 2.0 (61 cm)
1976 2.5 (76 cm) 1.5 (46 cm)
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Carey, Jennifer H. 1995. Philadelphus lewisii. In: Fire Effects Information System, [Online]. U.S. Department of Agriculture, Forest Service, Rocky Mountain Research Station, Fire Sciences Laboratory (Producer). Available: http://www.fs.fed.us/database/feis/

Post-fire Regeneration

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More info for the terms: ground residual colonizer, root crown, shrub

Tall shrub, adventitious-bud root crown
Ground residual colonizer (on-site, initial community)
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Carey, Jennifer H. 1995. Philadelphus lewisii. In: Fire Effects Information System, [Online]. U.S. Department of Agriculture, Forest Service, Rocky Mountain Research Station, Fire Sciences Laboratory (Producer). Available: http://www.fs.fed.us/database/feis/

Regeneration Processes

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More info for the terms: fruit, root crown, seed, stratification

Lewis' mock orange reproduces vegetatively and by seed.

Seeds accumulate in the seedbank. Sparsely distributed viable seeds
were collected from the top 2 inches (5 cm) of soil in forested sites in
central Idaho [19]

Stratification of Lewis' mock orange increases germination. Seeds
stratified for 8 weeks at 41 degrees Fahrenheit (5 deg C) then placed in
a sand medium at 72 to 79 degrees Fahrenheit (22-26 deg C) had 64
percent germination. The germination rate was less than 10 percent for
seeds stratified less than 8 weeks [39]. Germination was 52 percent
when stored at room temperature for 3 years and 39 percent when stored
at 40 degrees Fahrenheit (4 deg C) [28]. Germination was low when seeds
were fully exposed to light or kept in complete darkness [39].

Fruit development was adversely affected by drought in northern Idaho.
Fruit partially developed, turned brown, and opened, but no viable seed
was produced [8].

Lewis' mock orange sprouts from the root crown [10].
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Carey, Jennifer H. 1995. Philadelphus lewisii. In: Fire Effects Information System, [Online]. U.S. Department of Agriculture, Forest Service, Rocky Mountain Research Station, Fire Sciences Laboratory (Producer). Available: http://www.fs.fed.us/database/feis/

Regional Distribution in the Western United States

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This species can be found in the following regions of the western United States (according to the Bureau of Land Management classification of Physiographic Regions of the western United States):

1 Northern Pacific Border
2 Cascade Mountains
4 Sierra Mountains
5 Columbia Plateau
8 Northern Rocky Mountains
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Carey, Jennifer H. 1995. Philadelphus lewisii. In: Fire Effects Information System, [Online]. U.S. Department of Agriculture, Forest Service, Rocky Mountain Research Station, Fire Sciences Laboratory (Producer). Available: http://www.fs.fed.us/database/feis/

Successional Status

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More info for the term: shrub

Lewis' mock orange is tolerant of moderate shade [43]. It is an early to
mid-seral species [38] and is often present in seral shrub communities
following logging and burning [48]. Although normally scattered at low
densities [43], it sometimes occurs in dense, localized stands [17].
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Carey, Jennifer H. 1995. Philadelphus lewisii. In: Fire Effects Information System, [Online]. U.S. Department of Agriculture, Forest Service, Rocky Mountain Research Station, Fire Sciences Laboratory (Producer). Available: http://www.fs.fed.us/database/feis/

Synonyms

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Philadelphus californicus Benth. [18]
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Carey, Jennifer H. 1995. Philadelphus lewisii. In: Fire Effects Information System, [Online]. U.S. Department of Agriculture, Forest Service, Rocky Mountain Research Station, Fire Sciences Laboratory (Producer). Available: http://www.fs.fed.us/database/feis/

Taxonomy

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The currently accepted scientific name for Lewis' mock orange is
Philadelphus lewisii Pursh (Hydrangeaceae) [14,15,18]. Recognized
varieties [18] and subspecies [14,32] are as follows:

Philadelphus lewisii var. angustifolius (Rydb.) Hu
Philadelphus lewisii var. ellipticus Hu
Philadelphus lewisii var. gordonianus (Lindl.) Jepson
Philadelphus lewisii var. helleri (Rydb.) Hu
Philadelphus lewisii var. intermedius (A. Nels.) Hu
Philadelphus lewisii var. lewisii Pursh
Philadelphus lewisii var. oblongifolius Hu
Philadelphus lewisii var. parvifolius Hu
Philadelphus lewisii var. platyphyllus (Rydb.) Hu

Philadelphus lewisii subsp. californicus (Benth.) Munz.
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Carey, Jennifer H. 1995. Philadelphus lewisii. In: Fire Effects Information System, [Online]. U.S. Department of Agriculture, Forest Service, Rocky Mountain Research Station, Fire Sciences Laboratory (Producer). Available: http://www.fs.fed.us/database/feis/

Comprehensive Description

provided by North American Flora
Philadelphus gordonianus Lindl. Bot. Reg. 24 :
Misc. Notes 21. 1838.
Philadelphus cordatus^eu. &Kirchn. Arb. Muse. 203. 1864. Philadelphus Grahami Petz. & Kirchn. loc. ciL 1864. Philadelphus californicus Carr. Rev. Hortic. 37 : 339. 1866. Deutzia calif ornica Carr. loc. cit,, as a synonym. 1866. Philadelphus globosus K. Koch, Dendr. 1 . 344, as a synonym. 1869. Philadelphus columbarius Dippel, Handb. Laubh. 3 : 342. 1893.
A deciduous shrub, 2-4 m. high, with ascending branches; bark of young twigs red or chestnut, at first slightly pubescent, of the previous season's growth yellowish or gray, not exfoliating, and of the old stems gray and rarely peeling off in age ; buds enclosed in the bases of the petioles; petioles about 5 mm. long, hairy; leaf -blades ovate, more or less acuminate at the apex, acute or rounded, or in vigorous shoots subcordate at the base, usually coarsely dentate, but sometimes almost entire, more or less pubescent on both sides, densely so in the axils of the ribs beneath and all over on those of the young shoots, usually strongly 5-ribbed, all the ribs originating at the base ; flowering branches 1-1.5 dm. long with 2-4 pairs of leaves and 5-9 racemose flowers; hypanthium glabrous; sepals lance-ovate, abruptly acuminate into a slender point, glabrous or with a few hairs without, slightly tomentulose within; petals white, oblong to oval, rounded or often retuse at the apex, 1.5-2 cm. long; styles united half their length or nearly to the apex; stigmas clavate ; capsule ovoid, 8-10 mm. long, abruptly contracted below.
Type locality : Banks of the Columbia River.
Distribution : On hills and along streams, Idaho and British Columbia to northern California.
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bibliographic citation
John Kunkel SmaII, Per Axel Rydber, Nathaniel Lord Britton, Percy Wilson, Henry Hurd Rusby. 1905. ROSALES, PODOSTEMONACEAE, CRASSULACEAE, PENTHORACEAE and PARNASSIACEAE. North American flora. vol 22(2). New York Botanical Garden, New York, NY
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Comprehensive Description

provided by North American Flora
Philadelphus lewisii Pursh, Fl. Am. Sept. 329. 1814
Philadelphus confusus Piper, Bull. Torrey Club 29 : 225, in part. 1902.
A deciduous shrub, 1-2 m. high, with ascending branches; bark of the young twigs reddish and glabrous, of the previous season's growth red or rarely yellowish-gray, with conspicuous cross-cracks and sooner or later peeling off, and of the old stems light-gray and furrowed ;'buds enclosed in the bases of the petioles ; petioles 5-8 mm. long ; leaf-blades ovate-lanceolate or ovate, thick, acute at the apex, acute or rounded at the base, glabrous or sparingly hairy on the veins or more densely so in the axils of the veins beneath, entire or denticulate, 3or 5-ribbed, the stronger laterals arising some distance from the base of the leaf-blade; hypanthium glabrous; sepals lance-ovate, abruptly acuminate, glabrous without, finely pubescent within ; petals white, elliptic to oval, rounded or retuse at the apex, 10-15 mm. long; stamens numerous; styles united half-way up or to near the top; stigmas clavate, smaller than the anthers ; capsule short-obovoid with an almost conical base, about 1 cm. long.
Type locality : Waters of Clark's River (Clark's Fork of Columbia River in Montana). DiSTRiBUTio?^ : Hills and copses, from central Montana to Washington and Oregon. 1^^ NORTH AMERICAN FI,ORA [V01.UMK 22
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John Kunkel SmaII, Per Axel Rydber, Nathaniel Lord Britton, Percy Wilson, Henry Hurd Rusby. 1905. ROSALES, PODOSTEMONACEAE, CRASSULACEAE, PENTHORACEAE and PARNASSIACEAE. North American flora. vol 22(2). New York Botanical Garden, New York, NY
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Comprehensive Description

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Philadelphus ellipticus Rydb. sp. no v
A shrub about 1 m. high, with spreading branches ; bark of the young twigs brown and more or less shortvillous, of the previous season's growth brown, exfoliating, and of the old stems gray ; buds hidden in the bases of the short petioles ; leaf-blades elliptic, oval or rarely ovate, acute at the base, acutish at the apex, entire-margined, more or less leathery, 1-2.5 cm. long, glabrous or nearly so and shining above, pale and strigose beneath, 3-ribbed ; flowering branches about 5 cm. long or less, with 2-4 pairs of leaves and usually solitary terminal flowers ; the very short pedicel and hypanthium densely white-villous ; sepals ovate or lance-ovate, acuminate, about 6 mm. long, densely white-villous without and along the margin within ; petals white, broadly obovate, 15-18 mm. long, usually retuse at the apex ; stamens many ; styles wholly and the oblong stigmas partly united ; capsule broadly obovoid, about 5 mm. long.
Type collected near Mesilla Park, New Mexico, in 1896, by J. D. Tinsley. Distribution : Southern New Mexico.
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John Kunkel SmaII, Per Axel Rydber, Nathaniel Lord Britton, Percy Wilson, Henry Hurd Rusby. 1905. ROSALES, PODOSTEMONACEAE, CRASSULACEAE, PENTHORACEAE and PARNASSIACEAE. North American flora. vol 22(2). New York Botanical Garden, New York, NY
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Philadelphus angustifolius Rydb. sp. no v
A slender shrub with spreading branches; barkof the young twigs red or chestnut, sparingly hairy, and of the previous season's growth chestnut, with prominent cross-cracks and peeling off; buds hidden in the bases of the petioles; petioles 5 mm. longer less; leafblades lanceolate, acute at the base and acute or somewhat acuminate at the apex, 3-5 cm. long, usually denticulate with a few distant teeth, sparingly hairy along the veins on both sides; flowering branches 5-10 cm. long, with 1-3 pairs of leaves and a raceme of about 5 flowers ; hypanthium glabrous; sepals about 5 mm. long, lanceolate, acute ; petals oblong, rounded at the apex, 10-12 mjn. long; styles united to near the top; stigmas clavate, narrower than the anthers.
Type collected at Palace Camp, Washington, in 1883, by Mrs. Bailey Willis. Distribution : Type locality and near Tacoma.
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John Kunkel SmaII, Per Axel Rydber, Nathaniel Lord Britton, Percy Wilson, Henry Hurd Rusby. 1905. ROSALES, PODOSTEMONACEAE, CRASSULACEAE, PENTHORACEAE and PARNASSIACEAE. North American flora. vol 22(2). New York Botanical Garden, New York, NY
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Philadelphus platyphyllus Rydb. sp. nov
A tall, deciduous shrub with ascending branches ; bark of the young twigs light-brown, glabrous, of the previous season's growth and older stems gray, not exfoliating ; buds enclosed in the bases of the petioles ; petioles 8-10 mm. long; leaf-blades rounded, ovate or broadly oval, short-acuminate at the apex, mostly rounded at the base, coarsely dentate, thin, 4-7 cm. long, 3-5 cm. wide, sparingly hairy on both sides, densely so in the axils of the ribs beneath ; ribs 5, originating near the base ; flowering branches 1-2 dm. long, with 2- pairs of leaves and 5-11 racemose, rather crowded flowers ; hypanthium glabrous or nearly so ; sepals glabrous, lanceolate, acute, 6-7 mm. long ; petals white, elliptic or oval, retuse or emarginate or rounded at the apex ; stamens numerous ; styles united half their length; stigmas clavate, narrower than the anthers; capsule elongated-obovoid, fully 1 cm. long.
Type collected on the south side of Mt. Shasta, California, in 1897, by H. BBrown, no. 561. Distribution : Northern California and apparently also western Montana.
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John Kunkel SmaII, Per Axel Rydber, Nathaniel Lord Britton, Percy Wilson, Henry Hurd Rusby. 1905. ROSALES, PODOSTEMONACEAE, CRASSULACEAE, PENTHORACEAE and PARNASSIACEAE. North American flora. vol 22(2). New York Botanical Garden, New York, NY
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Philadelphus helleri Rydb. sp. nov
A shrub 2-3 m. high, with ascending branches ; bark of young twigs brownish, glabrous, and of the previous season's growth yellowish or grayish or partly brown, not exfoliating; buds enclosed in the bases of the petioles ; petioles about 5 mm. long; leaf-blades ovate or lance-ovate, acute at both ends or rounded at the base, denticulate or entire, 4-7 cm. long, sparingly hairy along the veins, or glabrous except the axils of the veins beneath; flowering branches 7-15 cm. long, with 2 or 3 pairs of leaves and 5-11 racemose flowers; hypanthium glabrous; sepals 7-8 mm. long, lance-ovate, shortacuminate, glabrous without, tomentose within ; petals white, lance-ovate, 15-18 mm. long, obtuse or acute at the apex ; stamens numerous ; styles united to near the apex, or two thirds their length; stigmas clavate, narrower than the anthers.
Type collected about Lake Waba, Nez Perces County, Idaho, i*n 1896, by A. A. and EGertrude Heller, no. 3374.
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John Kunkel SmaII, Per Axel Rydber, Nathaniel Lord Britton, Percy Wilson, Henry Hurd Rusby. 1905. ROSALES, PODOSTEMONACEAE, CRASSULACEAE, PENTHORACEAE and PARNASSIACEAE. North American flora. vol 22(2). New York Botanical Garden, New York, NY
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Philadelphus lewisii

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Philadelphus lewisii, the Lewis' mock-orange, mock-orange, Gordon's mockorange, wild mockorange,[1] Indian arrowwood, or syringa,[2] is a deciduous shrub native to western North America, and is the state flower of Idaho.[2]

It was first collected for science by scientist and explorer Meriwether Lewis in 1806 during the Lewis and Clark Expedition, and so was named after him.[3][4]

Description

The perennial shrub[5] is rounded and grows to 1.5–3 meters (4 ft 11 in – 9 ft 10 in) in height. It sends out long stems which are red when new and fade to gray with age, the bark shredding in small flakes.[6]

The oppositely arranged leaves vary in size across individual plants but they are usually oval, 3–5 centimeters (1+14–2 in) long, smooth or serrated along the edges, and light green in color with a rough texture.[6]

The flowers are produced in clusters at the ends of long stems, with four white petals up to 4 cm (1+12 in) long and numerous yellow stamens. At the height of flowering, the plant is covered in a mass of blossoms. The flowers have a heavy, sweet scent similar to orange blossoms with a hint of pineapple.[6]

The fruit is a small hard capsule about 1 cm long with woody, pointed wings, containing many brown seeds.[7] Drought will stunt fruit development and prevent the production of viable seeds.[8]

The plant is somewhat similar in appearance to serviceberry.[9]

Fruits

Distribution and habitat

Lewis' mock-orange occurs from northwestern California in the Sierra Nevada, north through the Pacific Northwest to southern British Columbia, and east to Idaho and Montana.[10] In the Cascades it occurs from sea level up to 7,000 feet (2,100 m), while in the Sierra Nevada it grows from 1,000–5,000 feet (300–1,520 m).[8]

Though it is tolerant of moderate shade, it prefers full sun. It is common in open coniferous forests and on forest edges, and in drier regions of the Northwest it occurs mostly in wetter and riparian areas. It is also found in chaparral and seral communities.[8]

It occurs on well-drained, moist sites, and is tolerant of rocky soil, and it is hardy from USDA zones 3 to 9.[7][8]

Ecology

Leaves and stems.

The foliage of Lewis' mock orange is of moderate importance as winter forage for elk and deer in British Columbia, Idaho, and Montana. In Montana, a 1957 study found that it comprised 2% of mule deer diets in the winter and a trace in the summer.[8]

The seeds are eaten by quail and squirrels.[8]

It occurs in dense shrub habitats which provide good thermal and security cover for wildlife.[8]

Philadelphus lewisii is able to spread vegetatively and from seed. It forms seedbanks in the top 2 inches (5.1 cm) of soil.[8]

Fire ecology

The shrub is native to relatively arid regions of the American West which experience frequent wildfires, and is therefore quite well adapted to fire. Although mock-orange is typically completely top-killed by fires, it will enthusiastically resprout from rhizomes and root crowns afterward.[8] A 1971 study found that in the next growing season after a fire, mock-orange had already regrown to 50% of its previous diameter and height, and that those plants had an average of 28.9 to 38.0 sprouts per plant postfire compared to just 0.6 to 1.5 before.[11]

Lewis' mock orange palatability for Rocky Mountain elk is much greater after fire, with 36.3% of twigs browsed compared to only 1.3% on adjacent unburned sites.[8]

Human use

Ethnobotany

Native American tribes used P. lewisii for numerous purposes. The hard wood was useful for making hunting and fishing tools, snowshoes, pipes, combs, cradles, netting shuttles, and furniture. The leaves and bark, which contain saponins, were mixed in water for use as a mild soap.[12]: 787  [1] The flowers were used in preparing perfumes and teas.[12]: 787 

Cultivation

Lewis' mock-orange prefers full sun to partial sun. It is drought-tolerant, will grow in poor soils, and is suitable for xeriscaping.[7] It provides a landscape with flashy flowers and a fruity scent.

Waterton Mockorange Philadelphus lewisii 'Waterton'

The Waterton Mockorange Philadelphus lewisii 'Waterton' was hybridized by the Alberta Horticultural Research Station in Brooks, Alberta, Canada. It grows to 4–6 ft. (1.2-1.8 m) in height. It is named for Augustus Griffin, who in 1933 noted that this plant was growing in what is now Waterton Lakes National Park in Canada.[7]

Symbolism

Philadelphus lewisii is the state flower of Idaho. The plant is protected by Idaho state law along with other native wildflowers and shrubs, and it is illegal to collect wild specimens on public property for export, sale, or transport without approval.[13]

References

  1. ^ a b Lewis' Mockorange Philadelphus lewisii Pursh (PDF), Plant guide, 21 November 2003, retrieved 23 March 2016
  2. ^ a b "State Symbols USA: Idaho State Flower". 2 October 2014.
  3. ^ "Wildflowers - Syringa Philadelphus lewisii Hydrangea family". USDA. Retrieved 23 March 2016.
  4. ^ Shaw, N. L. Philadelphus lewisii. In: Francis, J. K. (ed). Wildland Shrubs of the United States and its Territories: Thamnic Descriptions. USDA Forest Service. GTR IITF-WB-1.
  5. ^ "Lady Bird Johnson Wildflower Center - The University of Texas at Austin". www.wildflower.org. Retrieved 2022-01-01.
  6. ^ a b c "Mock Orange, Philadelphus lewisii". calscape.org.{{cite web}}: CS1 maint: url-status (link)
  7. ^ a b c d "Waterton Mockorange Philadelphus lewisii Waterton", Dave's Garden, retrieved 23 March 2016
  8. ^ a b c d e f g h i j "Philadelphus lewisii". www.fs.fed.us. Retrieved 2021-05-07.
  9. ^ Taylor, Ronald J. (1994) [1992]. Sagebrush Country: A Wildflower Sanctuary (rev. ed.). Missoula, MT: Mountain Press Pub. Co. p. 166. ISBN 0-87842-280-3. OCLC 25708726.
  10. ^ "Philadelphus lewisii". www.wnps.org.{{cite web}}: CS1 maint: url-status (link)
  11. ^ Leege, Thomas A.; Hickey, William O. (July 1971). "Sprouting of Northern Idaho Shrubs after Prescribed Burning". The Journal of Wildlife Management. 35 (3): 508. doi:10.2307/3799705. ISSN 0022-541X. JSTOR 3799705.
  12. ^ a b Bonner, Franklin T. (July 2008). The Woody Plant Seed Manual. USDA. Washington, DC. ISBN 978-0-16-081131-9. Retrieved 23 March 2016.
  13. ^ "Section 18-3911 – Idaho State Legislature". Retrieved 2021-05-07.

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Philadelphus lewisii: Brief Summary

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Philadelphus lewisii, the Lewis' mock-orange, mock-orange, Gordon's mockorange, wild mockorange, Indian arrowwood, or syringa, is a deciduous shrub native to western North America, and is the state flower of Idaho.

It was first collected for science by scientist and explorer Meriwether Lewis in 1806 during the Lewis and Clark Expedition, and so was named after him.

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