dcsimg

Lifespan, longevity, and ageing

provided by AnAge articles
Maximum longevity: 46 years (captivity) Observations: It has been estimated that these animals live up to 46 years (Don Wilson and Sue Ruff 1999). One wild born specimen was about 36-37 years old when it died in captivity (Richard Weigl 2005).
license
cc-by-3.0
copyright
Joao Pedro de Magalhaes
editor
de Magalhaes, J. P.
partner site
AnAge articles

Behavior

provided by Animal Diversity Web

Perception Channels: tactile ; chemical

license
cc-by-nc-sa-3.0
copyright
The Regents of the University of Michigan and its licensors
bibliographic citation
Kiehl, K. 2001. "Lagenorhynchus obliquidens" (On-line), Animal Diversity Web. Accessed April 27, 2013 at http://animaldiversity.ummz.umich.edu/site/accounts/information/Lagenorhynchus_obliquidens.html
author
Katie Kiehl, Michigan State University
editor
Cynthia Sims Parr, University of Michigan-Ann Arbor
original
visit source
partner site
Animal Diversity Web

Conservation Status

provided by Animal Diversity Web

Pacific white-sided dolphins are not considered to be endangered. A recent estimate of the population of these mammals in the central North Pacific ranged between a minimum of about 500,000, to a maximum of 930,000. Therefore there is not any immediate danger for the extiction of these animals. They are hunted by Japanese coastal fishermen in the East China and Japan seas and taken accidentally in the North Pacific purse-seine fishery.

US Federal List: no special status

CITES: no special status

IUCN Red List of Threatened Species: least concern

license
cc-by-nc-sa-3.0
copyright
The Regents of the University of Michigan and its licensors
bibliographic citation
Kiehl, K. 2001. "Lagenorhynchus obliquidens" (On-line), Animal Diversity Web. Accessed April 27, 2013 at http://animaldiversity.ummz.umich.edu/site/accounts/information/Lagenorhynchus_obliquidens.html
author
Katie Kiehl, Michigan State University
editor
Cynthia Sims Parr, University of Michigan-Ann Arbor
original
visit source
partner site
Animal Diversity Web

Trophic Strategy

provided by Animal Diversity Web

Pacific white-sided dolphins eat fish that live in large schools, such as anchovies, herring, smelt, capelin, and mackerel. They feed in groups of 10-20 dolphins, each adult eating about 9 kilograms (20 lbs ) of food each day.

Animal Foods: fish

Primary Diet: carnivore (Piscivore )

license
cc-by-nc-sa-3.0
copyright
The Regents of the University of Michigan and its licensors
bibliographic citation
Kiehl, K. 2001. "Lagenorhynchus obliquidens" (On-line), Animal Diversity Web. Accessed April 27, 2013 at http://animaldiversity.ummz.umich.edu/site/accounts/information/Lagenorhynchus_obliquidens.html
author
Katie Kiehl, Michigan State University
editor
Cynthia Sims Parr, University of Michigan-Ann Arbor
original
visit source
partner site
Animal Diversity Web

Distribution

provided by Animal Diversity Web

Pacific white-sided dolphins have a primarily temperate distribution, remaining north of the tropics and south of the colder waters caused by arctic currents. Their range is from the Aleutian Islands through the Gulf of Alaska to the tip of Baja California in the eastern Pacific; and from Japan to the Kuril Islnads in the western Pacific.

Biogeographic Regions: pacific ocean (Native )

license
cc-by-nc-sa-3.0
copyright
The Regents of the University of Michigan and its licensors
bibliographic citation
Kiehl, K. 2001. "Lagenorhynchus obliquidens" (On-line), Animal Diversity Web. Accessed April 27, 2013 at http://animaldiversity.ummz.umich.edu/site/accounts/information/Lagenorhynchus_obliquidens.html
author
Katie Kiehl, Michigan State University
editor
Cynthia Sims Parr, University of Michigan-Ann Arbor
original
visit source
partner site
Animal Diversity Web

Habitat

provided by Animal Diversity Web

They are usually seen in deep waters up to 160 km (100 miles ) offshore. There seem to be local migrations inshore in the winter months.

Aquatic Biomes: coastal

license
cc-by-nc-sa-3.0
copyright
The Regents of the University of Michigan and its licensors
bibliographic citation
Kiehl, K. 2001. "Lagenorhynchus obliquidens" (On-line), Animal Diversity Web. Accessed April 27, 2013 at http://animaldiversity.ummz.umich.edu/site/accounts/information/Lagenorhynchus_obliquidens.html
author
Katie Kiehl, Michigan State University
editor
Cynthia Sims Parr, University of Michigan-Ann Arbor
original
visit source
partner site
Animal Diversity Web

Life Expectancy

provided by Animal Diversity Web

Average lifespan
Status: captivity:
20.0 years.

license
cc-by-nc-sa-3.0
copyright
The Regents of the University of Michigan and its licensors
bibliographic citation
Kiehl, K. 2001. "Lagenorhynchus obliquidens" (On-line), Animal Diversity Web. Accessed April 27, 2013 at http://animaldiversity.ummz.umich.edu/site/accounts/information/Lagenorhynchus_obliquidens.html
author
Katie Kiehl, Michigan State University
editor
Cynthia Sims Parr, University of Michigan-Ann Arbor
original
visit source
partner site
Animal Diversity Web

Morphology

provided by Animal Diversity Web

Pacific white-sided dolphins have torpedo-shaped bodies which help them move quickly through water. Body length of Pacific white-sided dolphins ranges from 150 to 310 cm. Their coloration is one of their most distinguishing features, they are black or dark gray on the dorsal surface with a white underside, and have bicolored fins and flippers. This coloration is believed to act as a form of camouflage in their aquatic environment.

Range mass: 82 to 124 kg.

Other Physical Features: endothermic ; bilateral symmetry

license
cc-by-nc-sa-3.0
copyright
The Regents of the University of Michigan and its licensors
bibliographic citation
Kiehl, K. 2001. "Lagenorhynchus obliquidens" (On-line), Animal Diversity Web. Accessed April 27, 2013 at http://animaldiversity.ummz.umich.edu/site/accounts/information/Lagenorhynchus_obliquidens.html
author
Katie Kiehl, Michigan State University
editor
Cynthia Sims Parr, University of Michigan-Ann Arbor
original
visit source
partner site
Animal Diversity Web

Reproduction

provided by Animal Diversity Web

Pods are made up of one dominant male and a number of other males and females. The dominant male mates with reproductively available females.

Mating System: polygynous

Female Pacific white-sided dolphins reach sexual maturity around 5-6 years of age, males are sexually mature at 8-10 years. Generally breeding occurs in the summer or fall, and gestation lasts approximately 11-12 months. Females give brith to a single calf, which is almost 3 feet long and can weigh up to 14 pounds.

Breeding season: Generally breeding occurs in the summer or fall.

Average number of offspring: 1.

Range gestation period: 11 to 12 months.

Range age at sexual or reproductive maturity (female): 5 to 6 years.

Range age at sexual or reproductive maturity (male): 8 to 10 years.

Key Reproductive Features: iteroparous ; seasonal breeding ; gonochoric/gonochoristic/dioecious (sexes separate); sexual ; viviparous

Average birth mass: 14000 g.

Average number of offspring: 1.

license
cc-by-nc-sa-3.0
copyright
The Regents of the University of Michigan and its licensors
bibliographic citation
Kiehl, K. 2001. "Lagenorhynchus obliquidens" (On-line), Animal Diversity Web. Accessed April 27, 2013 at http://animaldiversity.ummz.umich.edu/site/accounts/information/Lagenorhynchus_obliquidens.html
author
Katie Kiehl, Michigan State University
editor
Cynthia Sims Parr, University of Michigan-Ann Arbor
original
visit source
partner site
Animal Diversity Web

Diagnostic Description

provided by FAO species catalogs
Pacific white-sided dolphins, like all members of the genus Lagenorhynchus, are stocky animals with very short, thick snouts. The large flippers have slightly rounded tips. The dorsal fin, the species' most diagnostic feature, is prominent, strongly recurved, and bi-coloured. The dark grey back and sides are distinctly set off from the white belly by a black border. Light grey streaks beginning on the sides of the melon sweep downwards behind the eye and expand into large grey thoracic patches. Grey "suspender stripes", which start above the eyes, widen to bands on the sides of the tail stock. The lips are black. Each tooth row contains 23 to 36 pairs of relatively fine, sharply pointed teeth. Can be confused with: Pacific white-sided dolphins are most likely to be confused with common dolphins, because both species are found in large schools and have large light coloured flank patches. Beak length and specifics of the colour pattern are the best keys to distinguishing them.
license
cc-by-nc-sa-3.0
bibliographic citation
Marine mammals of the world. Jefferson, T.A., S. Leatherwood & M.A. Webber - 1993. FAO species identification guide. Rome, FAO. 320 p. 587 figs. . 
author
Food and Agriculture Organization of the UN
original
visit source
partner site
FAO species catalogs

Size

provided by FAO species catalogs
Adults of this species reach 2.5 m in length, with males slightly larger than females. Maximum weight is about 180 kg. Length at birth is unknown, but is thought to be about 1 m.
license
cc-by-nc-sa-3.0
bibliographic citation
Marine mammals of the world. Jefferson, T.A., S. Leatherwood & M.A. Webber - 1993. FAO species identification guide. Rome, FAO. 320 p. 587 figs. . 
author
Food and Agriculture Organization of the UN
original
visit source
partner site
FAO species catalogs

Brief Summary

provided by FAO species catalogs
Often seen in large herds of hundreds or even thousands, these highly gregarious dolphins are also commonly seen with other species, especially northern right whale dolphins and Risso's dolphins. They are highly acrobatic and playful, commonly bowriding, and often leaping, flipping, or somersaulting. Calving apparently occurs during a protracted summer breeding season, which extends into autumn. They feed mostly on small schooling fish and squid. There is evidence that these dolphins feed mostly on deep scattering layer (DSL) organisms, sometimes using cooperative foraging techniques.
license
cc-by-nc-sa-3.0
bibliographic citation
Marine mammals of the world. Jefferson, T.A., S. Leatherwood & M.A. Webber - 1993. FAO species identification guide. Rome, FAO. 320 p. 587 figs. . 
author
Food and Agriculture Organization of the UN
original
visit source
partner site
FAO species catalogs

Benefits

provided by FAO species catalogs
In the western Pacific, Japanese drive and harpoon fisheries take hundreds or thousands of Pacific white-sided dolphins in most years. Exploitation in the eastern Pacific has been limited to occasional incidental capture in fishing nets, and small numbers taken in a fishery for live animals. In the central Pacific, white-sided dolphins were recently killed in substantial numbers in the Asian pelagic driftnets fisheries for squid. The annual kill was on the order of 8 000 to 10 000 per year. IUCN: Insufficiently known.
license
cc-by-nc-sa-3.0
bibliographic citation
Marine mammals of the world. Jefferson, T.A., S. Leatherwood & M.A. Webber - 1993. FAO species identification guide. Rome, FAO. 320 p. 587 figs. . 
author
Food and Agriculture Organization of the UN
original
visit source
partner site
FAO species catalogs

Pacific white-sided dolphin

provided by wikipedia EN

The Pacific white-sided dolphin (Lagenorhynchus obliquidens), also known as the hookfin porpoise, is an active dolphin found in the cool or temperate waters of the North Pacific Ocean.[4][5]

Taxonomy

The Pacific white-sided dolphin was named by Smithsonian mammalogist Theodore Nicholas Gill in 1865. It is morphologically similar to the dusky dolphin, which is found in the South Pacific.[6] Genetic analysis by Frank Cipriano suggests the two species diverged around two million years ago.

Though traditionally placed in the genus Lagenorhynchus, molecular analyses indicate they are closer to dolphins of the genus Cephalorhynchus, in the Lissodelphininae subfamily, than to both the Atlantic white-sided dolphin and the White-beaked dolphin. It has therefore been proposed to move the Pacific white-sided dolphin to the resurrected genus Sagmatias together with other southern hemisphere Lagenorhyncus species (hourglass dolphin, Dusky dolphin and Peale's dolphin).[7] However, the detailed phylogenetic relationships within this group of dolphins is still not fully elucidated.

Description

Photo of dual-hemisphered brain
Pacific white-sided dolphin's brain at Okinawa Churaumi Aquarium

The Pacific white-sided dolphin has three colors. The chin, throat and belly are creamy white. The beak, flippers, back, and dorsal fin are a dark gray. Light gray patches are seen on the sides and a further light gray stripe runs from above the eye to below the dorsal fin, where it thickens along the tail stock. A dark gray ring surrounds the eyes.

The species is an average-sized oceanic dolphin. Females weigh up to 150 kg (330 lb) and males 200 kg (440 lb) with males reaching 2.5 m (8.2 ft) and females 2.3 m (7.5 ft) in length. Pacific white-sided dolphins usually tend to be larger than dusky dolphins. Females reach maturity at seven years. From 1990 to 1991, a study conducted by Richard C. Ferrero and William A. Walker revealed the vast majority of Pacific white-sided dolphins that fell victim to the drift nets were between the ages of 8.3 to 11 when they sexually matured.[8] The gestation period usually last for one year. Individuals are believed to live up 40 years or more.[6]

The Pacific white-sided dolphin is extremely active and mixes with many of the other North Pacific cetacean species. It readily approaches boats and bow-rides. Large groups are common, averaging 90 individuals, with supergroups of more than 300. Prey includes mainly hake, anchovies, squid, herring, salmon, and cod.[9]

They have an average of 60 teeth.[10]

Range and habitat

The range of the Pacific white-sided dolphin arcs across the cool to temperate waters of the North Pacific.[6][11][12] Sightings go no further south than the South China Sea on the western side and the Baja California Peninsula on the eastern. Populations may also be found in the Sea of Japan and the Sea of Okhotsk. In the northern part of the range, some individuals may be found in the Bering Sea. The dolphins appear to follow some sort of migratory pattern – on the eastern side they are most abundant in the Southern California Bight in winter, but further north (Oregon, Washington) in summer. Their preference for off-shore deep waters appears to be year-round.[13][14] The only known predator of the Pacific white-sided dolphin is the killer whale,[15] but at least one case of predation by the Great White Shark has been recorded.[16]

The total population may be as many as 1 million.[6] However, the tendency of Pacific white-sided dolphins to approach boats complicates precise estimates via sampling.

Behavior

These dolphins keep close company.[17] White-sided dolphins swim in groups of 10 to 100, and can often be seen bow-riding and doing somersaults.[6][18] Members form a close-knit group and will often care for a sick or injured dolphin. Animals that live in such large social groups develop ways to keep in touch, with each dolphin identifying itself by a unique name-whistle. Young dolphins communicate with a touch of a flipper as they swim beside adults.

Studies conducted on Pacific white-sided dolphins, as well as Risso's dolphin have revealed a multitude of things about how they communicate as a species, which was revealed to be vastly different from bottlenose dolphins and common dolphins.[19] The studies have revealed that their notches and spectral peaks happen to be more low pitched when juxtaposed with the bottlenose dolphins and common dolphins as mentioned earlier.[20] Other studies have revealed similar behaviors. Two studies conducted back in 2010 and 2011 revealed that the vocalizations of Pacific white-sided dolphins can range differently only from their behavioral states, indicating strong similarities between the acoustic and surface behavior for various foraging behaviors, including the possibility of an undescribed subspecies. The oceanographical data in the area can also effect the behavioral patterns of the dolphins. The studies also revealed that the different types of echolocations do vary based on the geographical locations; the first population of Pacific white-sided dolphins that were observed, inhabiting the waters near the Pacific United States seemed to more activity during the night while the second population of Pacific white-sided dolphins, that were also observed, inhabiting areas near Baja California, were observed to be more active during the day, possibly due the seasons and the dolphins' search for prey.[21][22]

The first sighting of the species on Commander Islands involved a single dolphin to travel along with a pod of killer whales in 2013.[23]

Pacific white-sided dolphins are known to sleep on average seven hours a night.[24]

Relation to humans

Left: High-jump during Pacific white-sided dolphin show at the Vancouver Aquarium
Right: Pacific white-sided dolphin named Spinnaker at Vancouver Aquarium

Protection

Until the United Nations banned certain types of large fishing nets in 1933, many Pacific white-sided dolphins were killed in drift nets. Some animals are still killed each year by Japanese hunting drives.

Captivity

Although overshadowed in popularity by bottlenose dolphins, Pacific white-sided dolphins are also a part of some marine theme park shows. Roughly 100 reside in dolphinaria in North America and Japan. In captivity, they tend to consume less amounts of food when compared to their wild counterparts, this could be the case due to the fact of temperatures changing in the water based on the seasons. However, the condition in which the dolphins lives, most likely in an aquarium tank, will impact how much energy is required for a captive dolphin to thrive in captivity. Studies have also shown that the highest amount of food intake that a captive Pacific white-sided dolphin displays in autumn when the dolphin increase their food intake as well as their body mass.[25][26]

See also

References

  1. ^ Mead, J. G.; Brownell, R. L. Jr. (2005). "Order Cetacea". In Wilson, D. E.; Reeder, D. M. (eds.). Mammal Species of the World: A Taxonomic and Geographic Reference (3rd ed.). Johns Hopkins University Press. pp. 723–743. ISBN 978-0-8018-8221-0. OCLC 62265494.
  2. ^ Ashe, E.; Braulik, G. (2018). "Lagenorhynchus obliquidens". IUCN Red List of Threatened Species. 2018: e.T11145A50361866. doi:10.2305/IUCN.UK.2018-2.RLTS.T11145A50361866.en. Retrieved 19 November 2021.
  3. ^ "Appendices | CITES". cites.org. Archived from the original on 2021-01-19. Retrieved 2022-01-14.
  4. ^ "Pacific White-Sided Dolphin". NOAA Fisheries. National Oceanic and Atmospheric Administration. 19 October 2021. Archived from the original on 15 November 2021. Retrieved 14 November 2021.
  5. ^ "Kenai Fjords National Park". National Park Service. Archived from the original on 15 November 2021. Retrieved 17 November 2021.
  6. ^ a b c d e Wursig, Bernd; Perrin, William F.; Thewissen, J. 'Hans' (2009-02-26). Encyclopedia of Marine Mammals. Academic Press. pp. 817–818. ISBN 978-0-08-091993-5. Archived from the original on 2021-12-27. Retrieved 2021-12-27.
  7. ^ Vollmer, Nicole L.; et al. (January 31, 2019). "Taxonomic revision of the dolphin genus Lagenorhynchus". Marine Mammal Science. 35 (3): 957–1057. doi:10.1111/mms.12573. S2CID 92421374. Archived from the original on 2021-11-11. Retrieved 2021-09-17.
  8. ^ Ferrero, R.C.; Walker, W.A. (1996). "Age, growth, and reproductive patterns of the Pacific white-sided dolphin (Lagenorhynchus obliquidens) taken in high seas drift nets in the central North Pacific Ocean". Canadian Journal of Zoology. 74 (9): 1673–1687. doi:10.1139/z96-185.
  9. ^ Morton, A. (January 2000). "Occurrence, Photo‐identification and Prey of Pacific White‐sided Dolphins (Lagenorhyncus obliquidens) in the Broughton Archipelago, Canada 1984–1998". Marine Mammal Science. 16 (1): 80–93. doi:10.1111/j.1748-7692.2000.tb00905.x.
  10. ^ Black, Nancy A. (2009). Perrin, William F.; Würsig, Bernd; Thewissen, J. G. M. (eds.). Encyclopedia of Marine Mammals (2nd ed.). Burlington Ma.: Academic Press. ISBN 978-0-12-373553-9. Archived from the original on 2009-11-09.
  11. ^ Salvadeo, CJ; Lluch-Belda, D; Gómez-Gallardo, A; Urbán-Ramírez, J; MacLeod, CD (2010-03-10). "Climate change and a poleward shift in the distribution of the Pacific white-sided dolphin in the northeastern Pacific". Endangered Species Research. 11: 13–19. doi:10.3354/esr00252. ISSN 1863-5407.
  12. ^ "PACIFIC WHITE-SIDED DOLPHIN (Lagenorhynchus obliquidens): North Pacific Stock" (PDF). NOAA Fisheries: 127–130. 2012. Archived (PDF) from the original on 2022-03-03. Retrieved 2022-12-31 – via ResearchGate.
  13. ^ Black, N.A. (1994). Behavior and ecology of Pacific white-sided dolphins (Lagenorhynchus obliquidens) in Monterey Bay, California. San Francisco State University.
  14. ^ Walker, W.A.; Leatherwood, S.; Goodrich, K.R.; Perrin, W.F.; Stroud, R.K. (1986). "Geographical variation and biology of the Pacific white-sided dolphin, Lagenorhynchus obliquidens, in the north-eastern Pacific". Research on Dolphins. 441: 465.
  15. ^ Dahlheim, M.E.; Towell, R.G. (1994). "Occurrence and Distribution of Pacific White-Sided Dolphins (Lagenorhynchus obliquidens) in Southeastern Alaska, With Notes On an Attack by Killer Whales (Orcinus orca)". Marine Mammal Science. 10 (4): 458–464. doi:10.1111/j.1748-7692.1994.tb00501.x. Archived from the original on 2020-09-25. Retrieved 2019-06-26.
  16. ^ Peter Klimley, A.; Ainley, David G. (3 April 1998). Great White Sharks: The Biology of Carcharodon carcharias. ISBN 9780080532608. Archived from the original on 14 March 2023. Retrieved 25 March 2022.
  17. ^ "Archived copy". academic.oup.com. Archived from the original on 2022-12-31. Retrieved 2022-12-31.{{cite web}}: CS1 maint: archived copy as title (link)
  18. ^ "Archived copy". academic.oup.com. Archived from the original on 2022-12-31. Retrieved 2022-12-31.{{cite web}}: CS1 maint: archived copy as title (link)
  19. ^ Henderson, E. Elizabeth; Hildebrand, John A.; Smith, Michael H. (July 2011). "Classification of behavior using vocalizations of Pacific white-sided dolphins (Lagenorhynchus obliquidens)". The Journal of the Acoustical Society of America. 130 (1): 557–567. doi:10.1121/1.3592213. ISSN 0001-4966. PMID 21786921. Archived from the original on 2023-03-14. Retrieved 2022-12-31.
  20. ^ Soldevilla, M.S.; Henderson, E.E.; Campbell, G.S.; Wiggins, S.M.; Hildebrand, J.A.; Roch, M.A. (2008). "Classification of Risso's and pacific white-sided dolphins using spectral properties of echolocation clicks". The Journal of the Acoustical Society of America. 124 (1): 609–24. Bibcode:2008ASAJ..124..609S. doi:10.1121/1.2932059. PMID 18647003. S2CID 14685920. Archived from the original on 2020-07-27. Retrieved 2019-06-26.
  21. ^ Henderson, E.E.; Hildebrand, J.A.; Smith, M.H. (2011). "Classification of behavior using vocalizations of pacific white-sided dolphins (Lagenorhynchus obliquidens)". The Journal of the Acoustical Society of America. 130 (1): 557–67. Bibcode:2011ASAJ..130..557H. doi:10.1121/1.3592213. PMID 21786921.
  22. ^ Soldevilla, M.S.; Wiggins, S.M.; Hildebrand, J.A. (2010). "Spatio-temporal comparison of pacific white-sided dolphin echolocation click types". Aquatic Biology. 9: 49–62. doi:10.3354/ab00224.
  23. ^ Commander Islands Nature and Biosphere Reserve. Тихоокеанский белобокий дельфин Sagmatias obliquidens Gill, 1865 Archived 2017-08-23 at the Wayback Machine. Retrieved on August 24, 2017
  24. ^ Goley, Patricia Dawn (October 1999). "Behavioral Aspects of Sleep in Pacific White-Sided Dolphins (Lagenorhynchus Obliquidens, Gill 1865)1". Marine Mammal Science. 15 (4): 1054–1064. doi:10.1111/j.1748-7692.1999.tb00877.x. ISSN 0824-0469.
  25. ^ Piercey, R. (2013). "Seasonal changes in the food intake of captive Pacific white-sided dolphins (Lagenorhynchus obliquidens)". Aquatic Mammals. 39 (3): 211–220. doi:10.1578/am.39.3.2013.211.
  26. ^ "Seasonal Resting Metabolic Rate and Food Intake of Captive Pacific White-Sided Dolphins (Lagenorhynchus obliquidens)". Archived from the original on 2017-08-08. Retrieved 2017-07-12.

license
cc-by-sa-3.0
copyright
Wikipedia authors and editors
original
visit source
partner site
wikipedia EN

Pacific white-sided dolphin: Brief Summary

provided by wikipedia EN

The Pacific white-sided dolphin (Lagenorhynchus obliquidens), also known as the hookfin porpoise, is an active dolphin found in the cool or temperate waters of the North Pacific Ocean.

license
cc-by-sa-3.0
copyright
Wikipedia authors and editors
original
visit source
partner site
wikipedia EN