Elpeddo kaikai Barnard & Drummond 1978

Elpeddo kaikai

DESCRIPTION OF FEMALE.—Head about 19 percent of total body length, greatest width about 63 percent of length; rostrum unconstricted, not reaching middle of article 2 on antenna 1. Eyes absent. Article 1 on peduncle of antenna 1 about 1.6 times as long as wide, about 1.8 times as wide as article 2, ventral margin with about 9 setules, weakly produced dorsal apex with 2 setules; article 2 about 0.5 times as long as article 1, with apicoventral row of 6 setae; primary flagellum with 10 articles, about as long as peduncle, lacking aesthetascs; accessory flagellum with 7 articles. Spine formula on article 4 of antenna 2 = 7 (or possibly 3–4), dorsal margin with notch bearing 3 setae and one spine, ventral margin with 2 groups of 2–3 long to short setae, one ventrodistal long thin setal-spine; article 5 about 0.7 times as long as article 4, facial spine formula = 0, dorsal margin naked, ventral margin with 2 sets of 2 medium to short setae, 3 ventrodistal short to medium spines, one subdistal facial spine; flagellum about 1.8 times as long as articles 4–5 of peduncle combined, with 10 articles. Mandibles with medium palpar hump; right incisor with 3 teeth and notch; left incisor with 4 humps in 2 branches; right lacinia mobilis bifid, distal branch shorter than proximal, flabellate, denticulate, proximal branch weakly flabellate but pointed, with marginal denticles; left lacinia mobilis with 4 teeth plus 2 accessory teeth; right rakers 6 plus 2 rudimentaries; left rakers 7; molar composed of bulbous hump demarcated mainly by spines, each molar with 3 long spines, none disjunct, each molar with plume; palp article 1 slightly elongate, article 2 with one short inner apical seta and one other shorter inner seta, article 3 about as long as article 2, oblique apex with 6 spine-setae, basofacial formula = 0. Inner plate of maxilla 1 large, bearing one long apical pluseta, one similar apicomedial seta, one apicolateral seta; palp article 2 with 6 apicalmedial-lateral marginal spines and setae. Inner plate of maxilla 2 shorter than outer, outer not broader than inner, outer with 3 apicolateral setae, inner with one medial seta. Inner plate of maxilliped with one thin apical spine, 3 apicofacial setae, one medial seta; outer plate with 6 medial and apical spines, no apicolateral setae; palp article 1 naked, article 2 with one apicolateral seta, medial margin of article 2 moderately setose, article 3 with 3 facial setae, no lateral setae, nail of article 4 medium, with one accessory setule. Coxa 1 scarcely expanded distally, anterior margin weakly concave; main ventral setae of coxae 1–4 = 5–5–5–0, posteriormost seta of coxae 1–2 slightly shortened; anterior and posterior margins of coxa 4 almost parallel, posterior margin convex, posterodorsal corner sharp–rounded, posterodorsal margin short, V-shaped, width–length ratio of coxa 4 = 38:45. Long posterior setae on article 2 of gnathopods 1–2 and pereopods 1–2 = (3–4)–4–4–(3–4), short posteriors = (1–2)–3–(3–5)–4, long anteriors = 1–3–0–0, short anteriors = 4–3–4–3. Gnathopods weakly enlarged; wrist of gnathopod 2 short; width ratios of articles 5–6 on gnathopods 1–2 = 27:40 and 28:44, length ratios = 62:74 and 45:72; palmar humps ordinary, palms strongly oblique; article 5 of gnathopod 1 ovate, posterior margin flat; article 5 of gnathopod 2 short, triangular, almost lobate. Pereopods 1–2 similar; facial setae formula on article 4 = 1 and 2, on article 5 = 2 and 2, main spine of article 5 extending to M. 75 on article 6, article 5 lacking proximoposterior spines; spine formula of article 6 = 2 and 3, no middistal seta, acclivity on inner margin of dactyls of pereopods 1–2 weak, emergent setule short, midfacial pluseta short. Coxae 5–7 posteroventral setule formula = 1–1–1. Articles 4–5 of pereopods 3–4 narrow, facial spine rows sparse; facial ridge formula on article 2 of pereopods 3–5 = 0–1–2 (posterior short); width ratios of articles 2, 4, 5, 6 of pereopod 3 = 44:27:25:11, of pereopod 4 = 68:31:20:9, of pereopod 5 = 90:19:18:8, length ratios of pereopod 3 = 70:22:30:34, of pereopod 4 = 76:58:37:48, of pereopod 5 = 110:25:23:27; article 2 of pereopod 5 almost reaching middle of article 5; medial apex of article 6 finely combed, lacking digital processes. Posteroventral corner of epimeron 1 rounded–quadrate, posterior margin almost straight, anteroventral margin with 1–3 short widely spread setae, posteroventral face with 0–1 long seta; posteroventral corner of epimeron 2 rounded–quadrate, posterior margin almost straight, facial setae = 3; posteroventral corner of epimeron 3 weakly protuberant, with setule sinus, posterior margin convex, with one setule notch, ventral margin naked; epimera 1–3 with setule on posterodorsal margin. Urosomite 1 with lateral setule at base of uropod 1, articulation line complete. Urosomite 3 unprotuberant dorsally. Rami of uropods 1–2 with immersed weak apical nails and accessory scale; outer ramus of uropod 1 with 2 dorsal spines, inner with one; outer ramus of uropod 2 with 1–2 dorsal spines, inner with one dorsomedial spine; peduncle of uropod 1 with one apicolateral spine and one basofacial seta, medially with 4 marginal spines plus apical inner enlarged spine; peduncle of uropod 2 with 3–4 dorsal spines, medially with one apical spine; apicolateral corners of peduncles on uropods 1–2 with comb. Peduncle of uropod 3 with 3 ventral spines, dorsally with one lateral spine, one medial spine; rami feminine, inner extending to M. 60 on article 1 of outer ramus, apex with one seta, medial and lateral margins naked, article 2 of outer ramus elongate, 0.50, bearing 2 short setae, medial apex of article 1 with one seta, lateral margin with 2 acclivities, spine formula = 2–2–2, setal formula = 0. Telson long, length–width ratio = 6:5, almost fully cleft, each apex narrow, truncate, lateral acclivity weak, bearing ordinary lateral setule, spine next medial of length equal to setule, next spine slightly shorter, midlateral setules subsequal. Cuticle with bulbar setules of varying sizes closely packed on coxae and article 2 of pereopod 5 but rare elsewhere, surface bearing fine striations in form of linear fingerprint pattern, emergent setules especially short.

OBSERVATIONS (female).—One spine on outer plate of maxilla 1 in holotype aberrantly trifid.

DESCRIPTION OF MALE (male “y,” 3.08 mm, only specimen known, possibly aberrant).—Antenna 1 with thickened articles 2–3; basal article of flagellum swollen and bearing rows of long aesthetascs (similar to Lysianassidae). Peduncle of antenna 2 somewhat thinner than in female; anterior triad of spines and setae on article 4 short, spine formula of article 4 = 1–7 or 1–6, one long blunt spine on ventrodistal margin in place of seta found on female; anterior margin of article 5 with 2 giant calceoli; right antenna 2 with female-like flagellum comprising 8 articles about 1.6 times as long as articles 4–5 of peduncle combined, calceoli absent, left flagellum masculine but broken after 4 articles, calceoli absent. Upper lip with small midhump ventrally, otherwise like female. Right mandible with 5 rakers; molar and incisor like female, palps broken and missing, [left mandible lost during dissection]. Maxillipedal palp article 4 with apical nail setule-like and completely immersed, no accessory setules (condition aberrant?). Coxae generally like those of female, setae identical, coxae 2–3 slightly longer than in female, posterodorsal margin of coxa 4 even more dorsad than in female. Long posterior setae on article 2 of gnathopods 1–2 and pereopods 1–2 = (2–3)–3–2–2, short posterior setae = 0–0–1–(2–3), long anterior setae = 1–2–0–0, short anterior setae = 3–4–4–4, short setae strawlike. Pereopods 1–2, setae and spine formulas of article 4 = 1–1, article 5 = 2–2, article 6 = 2 + 3 on both pairs, article 4 thinner than in female, of pereopod 1 with only 2 groups of 4 and 3 setae posteriorly, of pereopod 2 with groups of 4, 3, and 1. Article 2 of pereopod 3 slightly thinner than in female; article 5 of pereopod 5 narrower and more elongate than in female, medial apex of article 6 on pereopod 5 with comb composed of much enlarged elements. Shape of epimera generally similar to that of female, epimeron 1 with 2 anteroventral setae and one posteroventral seta, none facial [ventral setae probably spread posteriad more than in female, thus 3 of female spread into groups of 2 and 1]; epimeron 2 with 2 facial setae; epimeron 3 lacking dorsal and posterior setules except at corner sinus. Urosomite 1 with dorsal saddle and invagination similar to that of lysianassids and ampeliscids. Medial margin on peduncle of uropod 1 with spines crowded toward distal margin (see illustration), basofacial setae = 2 elongate; outer ramus of uropod 2 with one dorsal spine. Uropod 3 intermediate between masculine and feminine, inner ramus not elongate but bearing medial setae, outer ramus with medial setae on article 1. Telson slightly elongate, bearing irregular basomedial rows of denticles.

OBSERVATIONS (male).—Possibly aberrant, only one specimen known.

ILLUSTRATIONS.—Inner plate of maxilliped skewed laterally in drawing; coxa of pereopod 4 reconstructed by reference to male “y” and female “c”; illustration of urosome and uropods 1–3 of male “y” partially reconstructed, very slightly shriveled on specimen; head of male dorsally similar to female, not illustrated.

HOLOTYPE—AM, female “a,” 3.40 mm.

TYPE-LOCALITY.—SBS C6S3, 22 May 1972, off Malabar, New South Wales, Australia, 75 m, sandy gravel.

VOUCHER MATERIAL.—SBS CxS4, male “y,” 3.08 mm (illus); SBS C6S2, female “c,” 3.05 mm (illus.).

MATERIAL.—SBS, 3 samples (3).

DISTRIBUTION.—New South Wales, off Malabar, 75–92 m, sandy gravel.

Ganba, new genus

DIAGNOSIS.—Eyes present. Flagella of antenna 1–2 unreduced in female. Article 2 of antenna 1 especially shortened, ventral setae confined apically. Article 1 of antenna 2 not ensiform, article 3 with 2 setules, facial spines on article 4 in one main row, article 5 short. Right mandibular incisor with 3 teeth; molar not triturative, small, bearing 3 long clumped spines with common base, not bearing fuzz; palpar hump medium. Palp of maxilla 1 biarticulate; inner plate with 2 setae. Setation of maxilla 2 weak. Inner plates of maxillipeds mostly fused together; apex of palp article 3 not protuberant, dactyl elongate, apical nail distinct, medium. Gnathopods dissimilar; gnathopod 2 weakly to moderately enlarged; article 5 of gnathopod 2 very short, cryptic, ordinary on gnathopod 1, without eusirid attachment; palms oblique, hands of gnathopods 1–2 elongate, basally broadened, poorly setose anteriorly. Article 5 of pereopods 1–2 setose posteroproximally. Article 2 of pereopod 3 of broad form, articles 4–5 of pereopods 3–4 narrow to medium, article 2 of pereopods 3–4 not setose posteriorly; pereopod 5 ordinary, article 2 naked ventrally, article 3 ordinary, dactyl normal. Epimera 1–2 lacking long posterior setae, without midfacial setae above ventral facial ridge; epimeron 3 of rounded classification and lacking long setae. Urosomite 1 generally naked except for sparse apicoventral setae or spines near base of uropod 1; urosomite 3 without dorsal hook or special process. Peduncle of uropod 1 normally elongate, without apicoventral spike, without special enlarged apciolateral-medial spine; peduncular apices of uropods 1–2 not combed; inner ramus of uropod 1 naked, no rami continuously spinose to apex, inner ramus of uropod 2 ordinary. Uropod 3 ordinary, article 2 of outer ramus carrying one long apical seta. Telson ordinary, with only one apical spine on each lobe plus setules.

DESCRIPTION.—Rostrum fully developed. [Fuzz on article 1 of antenna 1 in male unknown, calceoli on male antennae 1–2 unknown.] Prebuccal parts extended forward, poorly separated from each other, upper lip dominant. Right lacinia mobilis bifid, thin; mandibular palp thin, article 1 slightly elongate, article 2 without outer setae, apex of article 3 transverse. Lower lip bearing cones. Outer plate of maxilla 1 with 9 spines, no spine especially thickened. Inner plates of maxilliped poorly armed, thick. Coxae 2–4 with special anterodorsal humps. Some posterior spines on article 6 of pereopods 1–2 thin and seta-like, most thick and stiff, midapical spine or seta absent. Article 2 of pereopod 5 without facial setae. Peduncle of uropod 1 with dorsolateral spines confined apically, medial spines partially spread; peduncle of uropod 2 with only one medial spine confined apically, Peduncle of uropod 3 lacking extra subapical setae or spines. Telson with ordinary pair of midlateral or dorsal setules on each side.

TYPE-SPECIES.—Ganba pellati, new species.

COMPOSITION.—Unique.

RELATIONSHIP.—This genus differs from Kuritus and Wildus in the presence of only one seta on the outer ramus of uropod 3 and from Brolgus in the presence of mostly short and stout spines on article 6 of pereopods 1–2, in the much thinner gnathopod 1 and the partially fused inner plates of the maxilliped. Unlike other members of the Brolgus group of genera, Ganba lacks apical combs on the peduncles of uropods 1–2.