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Lifespan, longevity, and ageing

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Maximum longevity: 8.9 years (wild)
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Morphology

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The adult male rufous has a white breast with greenish back and crown. The back is sometimes glossed with metallic bronze-green. The pileum (the top of the head from the bill to the nape) is bronze-green, and the gorget (collar) is bright orange-red. The chin and throat is a shiny metallic scarlet color. The bill is long, straight, thin, black or dark brown in color. The feet are a dusky color. The adult female rufous has a metallic bronze-green back, and the pileum is a little duller than the male rufous. From the chin and the throat, down to the breast of the female, is a dull white color. Rufous Hummingbird has a body length of about 7.3 to 9.1 cm (2.87 to 3.58 in), and weighs around 2.8 g to 4.0 g (0.097 to 0.141 Oz). Unlike other birds that have large sound-producing muscles extending from the windpipe to the chest bone,the rufous has two sets of small vocal muscles in the trachea (Johnsgard 1983; Toops 1992; Sayre 1999).

Range mass: 2.8 to 4 g.

Average mass: 3.37 g.

Other Physical Features: endothermic ; bilateral symmetry

Average basal metabolic rate: 0.06853 W.

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Kong, B. 2001. "Selasphorus rufus" (On-line), Animal Diversity Web. Accessed April 27, 2013 at http://animaldiversity.ummz.umich.edu/site/accounts/information/Selasphorus_rufus.html
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Life Expectancy

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Average lifespan
Status: wild:
107 months.

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Habitat

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During breeding season, Rufous Hummingbirds are found in forests, on seed-tree harvest units, riparian shrub, and spruce-fir habitats. During the winter, it lives wherever flowers are present. It migrates to lowland stream bottoms, foothill brush land, seacoast and high mountain meadows (Johnsgard 1983; Paige et al. 1999).

Terrestrial Biomes: desert or dune ; savanna or grassland ; chaparral ; forest ; mountains

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Untitled

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The Rufous Hummingbird typically flaps its wings 50 times per second, and when it dives to mate, it can flap its wings 200 times per second (Toops 1992; Chloe 1999).

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Behavior

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Perception Channels: visual ; tactile ; acoustic ; chemical

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Kong, B. 2001. "Selasphorus rufus" (On-line), Animal Diversity Web. Accessed April 27, 2013 at http://animaldiversity.ummz.umich.edu/site/accounts/information/Selasphorus_rufus.html
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Conservation Status

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This widespread, abundant species is in no immediate danger of extinction. It is, like all hummingbirds, protected by the U.S. Migratory Bird Act and in CITES Appendix II.

US Migratory Bird Act: protected

US Federal List: no special status

CITES: appendix ii

State of Michigan List: no special status

IUCN Red List of Threatened Species: least concern

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Benefits

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Unknown

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Kong, B. 2001. "Selasphorus rufus" (On-line), Animal Diversity Web. Accessed April 27, 2013 at http://animaldiversity.ummz.umich.edu/site/accounts/information/Selasphorus_rufus.html
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Benefits

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Rufous Hummingbirds play an important role in pollinating at least 129 plant species (Paige et al. 1999).

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Trophic Strategy

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The Rufous Hummingbird consumes flies, ants, small beetles, tiny wasps and other small insects for a source of protein. Nectar is its most important food source for energy. It also drinks sap from the holes made by the Red-naped Sapsuckers for an extra food source. It feeds on nectar from several different flowering plants, such as honeysuckle, scarlet sage, horsemint, and black locust. This hummingbird is attracted to red and tubular flowers, preferring flowers that are spread farther apart giving it needed space for the beat of its wings. It eats about 1/2 to about 3 times its body weight. The Rufous Hummingbird feeds on nectar a minimum of sixty times a day. It consumes numerous small meals instead of a few big meals. It consumes nectar from flowering plants with its fork-like tongue at 13 licks per second ( Johnsgard 1983; Toops 1992; Chloe 1999; Sayre 1999; Gates and Gates 2000).

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Kong, B. 2001. "Selasphorus rufus" (On-line), Animal Diversity Web. Accessed April 27, 2013 at http://animaldiversity.ummz.umich.edu/site/accounts/information/Selasphorus_rufus.html
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Distribution

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Rufous Hummingbird, Selasphorus rufus, is found in western and coastal North America from March through August, and migrates to Mexico in the winter months of October through February (Johnsgard 1983).

Biogeographic Regions: nearctic (Native )

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Reproduction

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The breeding season for the Rufous Hummingbird begins in April and ends in July. The peak of the season usually occurs in May. The male will mate with several females during the breeding season. The male arrives at the desired breeding territory 2 to 3 weeks before the females. He attracts the female by climbing high into the air (20 to 45 m (75 to 150 ft)) then diving toward the female, pulling out of the dive and arcing back up into the sky after bottoming out within 2 or 3 inches of the female. During this mating display his wings are flapping at a rate of 200 wing beats per second, which creates the unique metallical "buzzing" sound heard during these displays. The female signals acceptance of the male as a mate by displaying the white tips of her tail. Copulation lasts only 3 to 5 seconds (Johnsgard 1983; Toops 1992; Chloe 1999).

The female builds her nest in blackberry vines, huckleberry bushes and other well protected overgrowths. Nests are built of mosses, leaves, and lichens woven together with spider webs. Rufous Hummingbirds usually lay two eggs about 1.3 cm (1/2 in) in size. The eggs take anywhere from 12 to 14 days to hatch, and the young leave the nest about 1 week after hatching. The female fiercely defends her nest from predators, while the male plays no role in defending or raising the young. (Johnsgard 1983; Toops 1992; Chloe 1999).

Range time to hatching: 12 to 14 days.

Average fledging age: 7 days.

Key Reproductive Features: iteroparous ; gonochoric/gonochoristic/dioecious (sexes separate); sexual ; oviparous

Average eggs per season: 2.

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Brief Summary

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A mature rufous hummingbird is about 7-9 cm in length, has an 11 cm wingspan, and weighs between 2 and 5 g. Males have an iridescent red throat, a dull reddish back, and an orange tail with pointed black tips. Females have a white throat with a few red feathers, a green back and head, and an orange, green, and black tail. Rufous hummingbirds feed on flower nectar, small insects, and tree sap. Males can be identified by their aerial display during courtship in which they make a series of steep, J-shaped dives that end at the same point. Males are territorial year-round. This bird can be found in the winter from Southern California, through Mexico, and along the northern Gulf Coast. In the summer, it breeds from southern Alaska southward to northern California, south central Idaho, and western Montana. These birds frequently visit the eastern states and have the northernmost range of any hummingbird. Selasphorus rufus is declining over most of its range.
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Brief Summary

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The Rufous Hummingbird (Selasphorus rufus) is relatively small even for a hummingbird, but can be quite aggressive in defending a patch of flowers at which it feeds. Like many hummingbirds, Rufous Hummingbirds often preferentially visit red tubular flowers. This species breeds farther north than any other hummingbird, all the way up to south-central Alaska and southern Yukon. Of all the western hummingbirds, the Rufous Hummingbird shows up most often in eastern North America. Rufous Hummingbirds breed around forest edges, streamsides, and mountain meadows and winter mainly in pine-oak woods in Mexico (although some winter along the Gulf Coast in the southeastern United States and casually northward throughout much of the eastern United States). This species is closely related to Allen's Hummingbird (Selasphorus sasin), which breeds mainly in coastal California. Females and immatures of these two species are nearly impossible to distinguish in the field, but adult males can usually be distinguished by back color (typically solid green in Allen's and mostly rufous in Rufous). Although the male Rufous Hummingbird's courtship display has been described as tracing a steep U or vertical oval, climbing high and then diving steeply, with whining and popping sounds at the bottom of the dive, Hurly et al. (2001) report that this is not accurate for many populations and likely is true for none. In their observations, males made J-shaped dive displays that were concave upward during both the climb and the dive. Males produce a high-pitched whine during dives. Toward the bottom of the arc, they produce a distinctive pulsing sound (chu-chu-chu-chu), and then end with a buzzing or “rattle” sound during the waggle. The male also performs a shuttle-flight display a few centimeters above an intruder, usually a female perched in low vegetation, hovering in a horizontal orientation and throwing himself from side to side with his head facing the female and his tail describing an arc on the order of 130°, and approximately 40 to 50 cm in length, in the horizontal plane. A characteristic throbbing buzz can be heard as the male moves from side to side. A single male may mate with several females.(Hurly et al. 2001) The nest of a Rufous Hummingbird is typically well concealed in the lower part of a coniferous tree, deciduous shrub, or vine. It is generally less than 5 m above the ground, although it may be as high as 10 m. Built by the female alone, it is a compact cup of soft materials held together with pieces of spider web and lined with plant down, The outside is camouflaged with pieces of lichen. Old nests may be repaired and re-used. The clutch of two (sometimes one, possibly rarely three or even four) white eggs is incubated by the female alone for 15 to 17 days. The female is also solely responsible for feeding the young after hatching. Age at first flight is around 21 days. This species is widespread and abundant, although there is some evidence of recent declines. (Kaufman 1996; AOU 1998)
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Life Cycle

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These hummingbirds have the longest known avian migration proportional to their body size, migrating over 1,500 km from Alaska to Mexico. Migration occurs between March and May. Nests are open cups placed in a shrub or on a small twig or branch of trees like conifers. Clutch size is two eggs. Upon hatching, the birds are helpless and naked.
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Pollinator

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Rufous hummingbirds (Selasphorus rufus) have been called the most important hummingbird pollinator in western North America. This is because rufous hummingbirds migrate from Alaska to Mexico, pollinating plants the entire journey. As the birds stop along nectar corridors to eat they inadvertently collect pollen and transfer it to another plant, often times moving genetic material great distances. In the 1960s these birds were seen pollinating the giant trumpets (Macromeria viridiflora) plant in Arizona, the first documented case of a hummingbird pollinating a member of the Boraginaceae family in western North America. M. viridiflora is a tall perennial herb with yellow tubular flowers. The hummingbird probes the floral tubes by fitting its bill into the upper part of the floral tube and extending its tongue to reach the nectar in the narrow, lower part of the tube. Pollen then becomes dusted on the hummingbird's bill and is inadvertently transferred when the bird moves from plant to plant eating nectar. The hummingbird's role as a pollinator during migration is believed to have influenced the speciation of California flowers. This species of hummingbird is declining throughout its range, and the loss of nectar corridors across its migration route may continue to threaten this bird.
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Pollinator

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Rufous hummingbirds (Selasphorus rufus) have been called the most important hummingbird pollinator in western North America. This is because rufous hummingbirds migrate from Alaska to Mexico, pollinating plants the entire journey. As the birds stop along nectar corridors to eat they inadvertently collect pollen and transfer it to another plant, often times moving genetic material great distances. In the 1960s these birds were seen pollinating the giant trumpets (Macromeria viridiflora) plant in Arizona, the first documented case of a hummingbird pollinating a member of the Boraginaceae family in western North America. M. viridiflora is a tall perennial herb with yellow tubular flowers. The hummingbird probes the floral tubes by fitting its bill into the upper part of the floral tube and extending its tongue to reach the nectar in the narrow, lower part of the tube. Pollen then becomes dusted on the hummingbird's bill and is inadvertently transferred when the bird moves from plant to plant eating nectar. The hummingbird's role as a pollinator during migration is believed to have influenced the speciation of California flowers. This species of hummingbird is declining throughout its range, and the loss of nectar corridors across its migration route may continue to threaten this bird.
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Rufous hummingbird

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The rufous hummingbird (Selasphorus rufus) is a small hummingbird, about 8 cm (3.1 in) long with a long, straight and slender bill. These birds are known for their extraordinary flight skills, flying 2,000 mi (3,200 km) during their migratory transits. It is one of nine species in the genus Selasphorus.

Taxonomy

The rufous hummingbird was formally described in 1788 by the German naturalist Johann Friedrich Gmelin in his revised and expanded edition of Carl Linnaeus's Systema Naturae. He placed it with all the other hummingbirds in the genus Trochilus and coined the binomial name Trochilus rufus.[3] Gmelin based his description on the Ruff-necked humming-bird that had been described by John Latham in 1782 and the Ruffed honeysucker that had been described by Thomas Pennant in 1785.[4][5] The type locality is Nootka Sound on the west coast of Vancouver Island in western Canada.[6] The rufous hummingbird is now placed with eight other species in the genus Selasphorus that was introduced in 1832 by the English naturalist William John Swainson.[7][8] The genus name combines the Ancient Greek selas meaning "light" or "flame" with -phoros meaning "-carrying". The specific epithet rufus is the Latin word for "red".[9] The species is considered as monotypic: no subspecies are recognised.[8]

Description

A perched male rufous hummingbird

The adult male has a white breast, rufous face, flanks and tail and an iridescent orange-red throat patch or gorget. Some males have some green on their back and/or crown. The female has green, white, and some iridescent orange feathers in the center of the throat, and a dark tail with white tips and rufous base.

The female is slightly larger than the male. Females and the rare green-backed males are extremely difficult to differentiate from Allen's hummingbird. The typical "notched" shape of the second rectrix (R2) is considered an important field mark to distinguish the adult male rufous hummingbird from the adult male Allen's hummingbird.[10] This is a typical-sized hummingbird, being a very small bird. It weighs 2–5 g (0.071–0.176 oz), measures 7–9 cm (2.8–3.5 in) long and spans 11 cm (4.3 in) across the wings.[11]

Distribution and habitat

A perched female rufous hummingbird

Western rufous hummingbirds migrate through the Rocky Mountains and nearby lowlands from May to September to take advantage of the wildflower season. They may stay in one local region for the entire summer, in which case the migrants, like breeding birds, often aggressively take over and defend feeding locations. Most winter in wooded areas in the Mexican state of Guerrero, traveling over 2,000 mi (3,200 km) by an overland route from its nearest summer home – a prodigious journey for a bird weighing only 3 to 4 g.[12]

Adult male rufous hummingbirds tend to migrate slightly earlier than females or young.[13] Since juveniles or females are essentially indistinguishable from Allen's hummingbirds unless confirmed by close inspection, eastern rufous migrants may be classified as "rufous/Allen's hummingbirds".[11]

Behavior and ecology

Food and feeding

They feed on nectar from flowers using a long extendable tongue or catch insects on the wing. These birds require frequent feeding while active during the day and become torpid at night to conserve energy. Because of their small size, they are vulnerable to insect-eating birds and animals.

Hovering and sexual dimorphism

Hovering male rufous hummingbird

A study that used digital imaging velocimetry to look at wing movements found that the rufous hummingbird supports its body weight during hovering primarily by wing downstrokes (75% of lift) rather than by upstrokes (25% of lift).[14][15] When hovering during fasting, rufous hummingbirds oxidize fatty acids to support metabolism and food energy requirements, but can rapidly switch to carbohydrate metabolism (within 40 minutes) after feeding on flower nectar.[16]

A juvenile male rufous hummingbird nectaring on Rocky Mountain Beeplant in Wyoming, USA

Both males and females are territorial; however, they defend different types of territories.[17] The more aggressive males fight to defend areas with dense flowers, pushing females into areas with more sparsely populated flowers.[17] Males generally have shorter wings than females, therefore their metabolic cost for hovering is higher. This allows males to beat their wings at high frequencies, giving them the ability to chase and attack other birds to defend their territory.[17] The metabolic cost of short wings is compensated for by the fact that these males do not need to waste energy foraging for food, because their defended territory provides plenty of sustenance.[18] Females on the other hand are not given access to the high concentration food sources, because the males fight them off.[17] Therefore, females generally defend larger territories, where flowers are more sparsely populated, forcing them to fly farther between food sources.[17] The metabolic cost of flying farther is compensated for with longer wings providing more efficient flight for females.[17] The differences in wing length demonstrate a distinct sexual dimorphism, allowing each sex to best exploit resources in an area.

Breeding

Their primary breeding habitats are open areas, mountainsides and forest edges in western North America from southern Alaska through British Columbia and the Pacific Northwest to California,[11] nesting further north (Alaska) than any other hummingbird. The female builds a nest in a protected location in a shrub or conifer. Males are promiscuous, mating with several females.[11]

Conservation status

In 2018, the rufous hummingbird was uplisted from least concern to near threatened on the IUCN Red List, on the basis that due to its reliance on insect prey during the wintering season, it will be heavily affected by the global decline in insect populations due to pesticides and intensified agriculture.[19] Due to climate change, many flowers that the rufous hummingbird feeds on during the breeding season have started blooming two weeks prior to the birds' arrival to their breeding locations, which may lead to rufous hummingbirds arriving too late to feed on them.[19]

Gallery

References

  1. ^ BirdLife International (2020). "Selasphorus rufus". IUCN Red List of Threatened Species. 2020: e.T22688296A178595564. doi:10.2305/IUCN.UK.2020-3.RLTS.T22688296A178595564.en. Retrieved 18 November 2021.
  2. ^ "Appendices | CITES". cites.org. Retrieved 2022-01-14.
  3. ^ Gmelin, Johann Friedrich (1788). Systema naturae per regna tria naturae : secundum classes, ordines, genera, species, cum characteribus, differentiis, synonymis, locis (in Latin). Vol. 1, Part 1 (13th ed.). Lipsiae [Leipzig]: Georg. Emanuel. Beer. p. 497.
  4. ^ Latham, John (1782). A General Synopsis of Birds. Vol. 1, Part 2. London: Printed for Benj. White. p. 785, Plate 35.
  5. ^ Pennant, Thomas (1785). Arctic Zoology. Vol. 2. London, United Kingdom: Printed by Henry Hughs. p. 290.
  6. ^ Peters, James Lee, ed. (1945). Check-List of Birds of the World. Vol. 5. Cambridge, Massachusetts: Harvard University Press. p. 141.
  7. ^ Swainson, William John; Richardson, J. (1831). Fauna boreali-americana, or, The zoology of the northern parts of British America. Vol. Part 2. The Birds. London: J. Murray. p. 324. The title page bears the year 1831 but the volume did not appear until 1832.
  8. ^ a b Gill, Frank; Donsker, David; Rasmussen, Pamela, eds. (January 2022). "Hummingbirds". IOC World Bird List Version 12.1. International Ornithologists' Union. Retrieved 12 July 2022.
  9. ^ Jobling, James A. (2010). The Helm Dictionary of Scientific Bird Names. London: Christopher Helm. pp. 352, 341. ISBN 978-1-4081-2501-4.
  10. ^ Pyle, Peter (1997). Identification Guide to North American Birds, Part I: Columbidae to Ploceidae. Steve N. G. Howell, Siobhan Ruck, Institute for Bird Populations, Point Reyes Bird Observatory. Bolinas, Calif.: Slate Creek Press. ISBN 0-9618940-2-4. OCLC 38593534.
  11. ^ a b c d "Rufous Hummingbird". All About Birds. Cornell Lab of Ornithology.
  12. ^ "Rufous Hummingbird Migration: Temperature, Patterns and Timing". Journey North. University of Wisconsin-Madison Arboretum. Retrieved 2021-03-18.{{cite web}}: CS1 maint: url-status (link)
  13. ^ "Rufous Hummingbird". Audubon Guide to North American Birds. 2014-11-13. Retrieved 2021-03-18.{{cite web}}: CS1 maint: url-status (link)
  14. ^ "Master fliers of the bird kingdom". BBC. 27 June 2005. Retrieved 30 October 2009.
  15. ^ Warrick DR, Tobalske BW, Powers DR (2005). "Aerodynamics of the hovering hummingbird". Nature. 435 (7045): 1094–7. Bibcode:2005Natur.435.1094W. doi:10.1038/nature03647. PMID 15973407. S2CID 4427424.
  16. ^ Welch KC Jr, Suarez RK (2007). "Oxidation rate and turnover of ingested sugar in hovering Anna's (Calypte anna) and rufous (Selasphorus rufus) hummingbirds". J Exp Biol. 210 (12): 2154–62. doi:10.1242/jeb.005363. PMID 17562889.
  17. ^ a b c d e f Kodric-Brown, A.; Brown, J.H. (1978). "Influence of economics, interspecific competition, and sexual dimorphism on territoriality of migrant Rufous hummingbirds". Ecology. 59 (2): 285–296. doi:10.2307/1936374. JSTOR 1936374.
  18. ^ Feinsinger; Chaplin (1975). "On the relationship between wing disc loading and foraging strategy in hummingbirds". The American Naturalist. 109 (966): 217–224. doi:10.1086/282988. S2CID 84567304.
  19. ^ a b BirdLife International (22 November 2018). "Red List update for birds" (PDF). BirdLife. Retrieved 23 November 2018.

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Rufous hummingbird: Brief Summary

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The rufous hummingbird (Selasphorus rufus) is a small hummingbird, about 8 cm (3.1 in) long with a long, straight and slender bill. These birds are known for their extraordinary flight skills, flying 2,000 mi (3,200 km) during their migratory transits. It is one of nine species in the genus Selasphorus.

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