^ abcDusbabek, Frantisek; Literak, Ivan; Capek, Miroslav; Havlicek, Martin (2007). "Ascid mites (Acari: Mesostigmata: Ascidae) from Costa Rican hummingbirds (Aves: Trochilidae), with description of three new species and a key to the Proctolaelaps belemensis species group". Zootaxa. 1484: 51–67.