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Associations ( англиски )

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The predators of Crotalus viridis include some predatory birds such as red-tailed hawks. When Crotalus viridis senses a predator, it makes a rattling sound to warn it. Smaller snakes tend to wait until the predator is closer to begin rattling, as do pregnant females.

Anti-predator Adaptations: cryptic

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Morphology ( англиски )

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The various subspecies of Crotalus viridis may vary slightly in color. Most are greenish gray or greenish brown in color, although members of one subspecies are black with only slight markings. These rattlesnakes have from 33-55 dark blotches on their back, which flatten into rings on the tail, surrounded by lighter markings. Juviniles have similar markings to adults, but may have higher contrast in coloring. All Crotalus viridis have a rattle at the end of their tail, made up of segments of keratin, which knock together to make a rattling sound. The number of segments varies because each time the snake sheds its skin, it gains another segment. Crotalus viridis is generally about 91.5 cm in length, but can be from 89-114 cm. Males and females are dimorphic in that the number of rings on their tails differs. Males tend to have between 6-15 rings, and females tend to have between 4-11 rings.

There are nine subspecies of Crotalus viridis. Crotalus viridis viridis , or Western or Prairie rattlesnake, is the most widespread and is found in most regions of the species. There are eight others which are mostly named for the region they are found in. They are Grand Canyon rattlesnake (Crotalus viridis abyssus), Coronado Island rattlesnake (Crotalus viridis caliginis), Arizona Black rattlesnake (Crotalus viridis ceberus), Midget Faded rattlesnake (Crotalus viridis concolor), Southern Pacific rattlesnake (Crotalus viridis helleri), Great Basin rattlesnake (Crotalus viridis lutosus), and Hopi rattlesnake (Crotalus viridis oreganus). (LaDuc, 2000; Melli, 1999; Herp-edia, 1998; Reptiles and Amphibians of North Dakota, 1999)

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Life Expectancy ( англиски )

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Average lifespan
Status: captivity:
24.1 years.

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Без наслов ( англиски )

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The name Crotalus comes from the Greek word "crotalon", which means "little bell", and is used to describe the rattle. The name viridis comes from the latin word for the color green.

Crotalus viridis can live up to twenty-one years. 95% of deaths of Crotalus viridis occur during the active season, rather than the winter. (Greene, 1997; Melli, 1999)

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Conservation Status ( англиски )

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There are currently no conservation measures being taken for Crotalus viridis because their existence is not considered threatened. (Melli, 1999)

US Federal List: no special status

CITES: no special status

IUCN Red List of Threatened Species: least concern

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Benefits ( англиски )

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The only adverse affect that Crotalus viridis can have on humans is that they may bite an individual that provokes them. Their venom can cause injury and death. (Jacobs, 1999)

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Benefits ( англиски )

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Crotalus viridis help to control populations of rodents and other animals that they feed on.

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Trophic Strategy ( англиски )

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Crotalus viridis eat small mammals, ground nesting birds, amphibians, and reptiles, including sometimes other snakes. They locate their prey by using their tongue to sense in airborn chemicals given off by the prey. Then they rapidly strike out at them, biting them with their fangs, then letting them go quickly. Venom is released from their fangs when the snakes strike. It works to immobilize the prey, which the snake then tracks and eats. The venom also works to destroy tissue and help with the digestion of bulkier prey.

Their venom is very deadly. Crotalus viridis use from 20-55% of their stored venom when they bite a small mammal such as a mouse. This is approximately 300 times the amount of venom needed to kill that animal. Venom is stored in glands which are connected to the hollow fangs. Venom is the means used to kill the prey, as the bite of the fangs alone would not usually result in death. The fangs are covered by thin tissue and fold back against the roof of the mouth when the mouth is closed.

During early spring and late fall when the weather is warm, Crotalus viridis hunts during the day. When the weather gets hotter, Crotalus viridis tends to seek shelter during the day and wait until night when it is cooler to hunt. (Greene, 1997; Jacobs, 1999; Kardong et. al., 1998; Kardong, 1996; Melli, 1999; Reptiles and Amphibians of North Dakota, 1999)

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Distribution ( англиски )

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Crotalus viridis are found across most of the United States west of Texas and the Dakotas. They are also found in northern Mexico and southwest Canada. (Melli, 1999; LaDuc, 2000)

Biogeographic Regions: nearctic (Native )

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Habitat ( англиски )

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Crotalus viridis are found mostly in grasslands and prairies, and in brush. Various subspecies can be found in woods, forests, caves, rock ledges, and alongside streams. Crotalus viridis avoids desert. (LaDuc, 2000; Melli, 1999; Crotalus viridis viridis, 1999)

Habitat Regions: temperate ; terrestrial

Terrestrial Biomes: savanna or grassland ; chaparral ; forest

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Reproduction ( англиски )

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Crotalus viridis mate between March and May. They have internal fertilization; males have a hemipenis which they use for copulation. Females give birth to between 4-21 live young in late summer or early fall. The babies are able to take care of themselves immediately and require no parental care. They are from 22-28 cm long when born and are already venomous. They reach sexual maturity after three years. Males may compete for females during mating season; however, body size of males does not seem to be a trait that contributes to mating success. In areas where females are scarce, males spend more time searching for females, and are not generally observed fighting over females.

(Duvall, 1997; Fitch, 1998; Melli, 1999; Reptiles and Amphibians of North Dakota, 1999; Crotalus viridis viridis, 1999)

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Amenazas ( шпански; кастиљски )

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Factores de riesgo

El principal riesgo que Lemos-Espinal (2001, personal) ha observado para las poblaciones de esta serpiente, es el hecho de que individuos de esta especie generalmente son atacados por el ser humano simplemente por ser considerados peligrosos.
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Lemos Espinal, J. A. 2003. Ficha técnica de Crotalus viridis. Fichas diagnósticas para 10 especies de anfibios y reptiles mexicanos. Facultad de Estudios Superiores Iztacala, Universidad Nacional Autónoma de México. Bases de datos SNIB-CONABIO. Proyecto No. W002. México, D.F.
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Biología de poblaciones ( шпански; кастиљски )

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Antecedentes del estado de la especie o de las poblaciones principales

No existe ningún estudio relacionado con el tamaño de las poblaciones de esta especie de serpiente. Hasta 1994 existían únicamente 6 registros de esta especie para México. Lemos-Espinal (2001, personal) ha observado que esta especie es relativamente común en la parte Oeste de los Médanos de Samalayuca, Chihuahua (Ranchos Luz Alba, El Gato y El Huérfano).
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Lemos Espinal, J. A. 2003. Ficha técnica de Crotalus viridis. Fichas diagnósticas para 10 especies de anfibios y reptiles mexicanos. Facultad de Estudios Superiores Iztacala, Universidad Nacional Autónoma de México. Bases de datos SNIB-CONABIO. Proyecto No. W002. México, D.F.
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Descripción ( шпански; кастиљски )

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Crotalus viridis tiene una coloración que varia de café claro a amarillo claro, puede llegar a medir 152.0 cm de longitud total. Tiene entre 35 y 55 manchas dorsales de color café oscuro sobre todo el cuerpo excluyendo la parte de la cola, estas manchas generalmente están limitadas por delgadas líneas blancas. La mayoría de estas manchas tienen una forma ovalada a la mitad del cuerpo, pero gradualmente se van haciendo más anchas y cortas formando bandas transversales en el tercio posterior del tronco. Hay dos líneas de pequeñas manchas a ambos lados, las manchas superiores son más grandes y difusas, las inferiores son más oscuras con colores claros en sus limites exteriores. Dos líneas oblicuas de color blanco son evidentes a los lados de la cabeza; la línea detrás del ojo pasa sobre la comisura de la boca. El vientre es blanco y sin manchas (Gloyd, 1940; Degenhardt et al. 1996).

Se reconocen nueve subespecies de C. viridis, tres de las cuales se distribuyen en la República Mexicana:
C. v. caliginis Klauber, 1949;
C. v. helleri Meek, 1905;
C. v. viridis (Rafinesque, 1818).
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Lemos Espinal, J. A. 2003. Ficha técnica de Crotalus viridis. Fichas diagnósticas para 10 especies de anfibios y reptiles mexicanos. Facultad de Estudios Superiores Iztacala, Universidad Nacional Autónoma de México. Bases de datos SNIB-CONABIO. Proyecto No. W002. México, D.F.
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Distribución ( шпански; кастиљски )

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Actual

Área prioritaria marina: Ensenadense

región prioritaria terrestre: Delta del Río Colorado

región prioritaria terrestre: El Vizcaino-El Barril

región prioritaria terrestre: Médanos de Samalayuca

En México esta serpiente se distribuye desde el Noreste del estado de Sonora hasta el Noreste del estado de Coahuila, incluyendo la parte Noroeste/Norte del estado de Chihuahua, Oeste de Baja California Norte, Norte de Baja California Sur e Isla Coronado del Sur (Campbell y Lamar, 1989; Lemos-Espinal et al., 1994).

región prioritaria terrestre: Punta Banda-Eréndida

región prioritaria terrestre: San Telmo-San Quintín

región prioritaria terrestre: Santa María-El Descanso

región prioritaria terrestre: Sierra de Juárez

región prioritaria terrestre: Sierra de San Luis-Janos

región prioritaria terrestre: Sierra de San Pedro Mártir

región prioritaria terrestre: Sierra Maderas del Carmen

región prioritaria terrestre: Sierras La Libertad-La Asamblea

región prioritaria terrestre: Valle de los Cirios
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Lemos Espinal, J. A. 2003. Ficha técnica de Crotalus viridis. Fichas diagnósticas para 10 especies de anfibios y reptiles mexicanos. Facultad de Estudios Superiores Iztacala, Universidad Nacional Autónoma de México. Bases de datos SNIB-CONABIO. Proyecto No. W002. México, D.F.
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Estado de conservación ( шпански; кастиљски )

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NOM-059-SEMARNAT-2001

Pr sujeta a protección especial

NOM-059-SEMARNAT-2010

Pr sujeta a protección especial
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Hábitat ( шпански; кастиљски )

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Médanos semiestablecidos, matorral xerófito.

Macroclima

Habita en climas áridos templados, áridos semicálidos, muy áridos semicálidos y muy áridos templados: BSokw, BSohw, BWh(x'), BWk(x").

Altitud a la que se distribuye en la República Mexicana: desde el nivel del mar (Isla Coronado del Sur, Baja California) hasta 1,457 m sobre el nivel del mar (Rancho La Viuda, Chihuahua).

Situación actual del hábitat con respecto a las necesidades de la especie

La mayor cantidad de ejemplares recolectados de esta especie de serpiente provienen de la zona de los médanos de Samalayuca, Chihuahua. En los meses de Verano es frecuente observar a estas serpientes bajo las vías del ferrocarril, en el tramo Samalayuca. Al parecer esta región aún conserva las condiciones idóneas para el desarrollo de C. viridis. Igualmente, en la zona de los Ranchos El Espía y La Viuda, mpio. de Ascensión, Chihuahua, el matorral xerófito se encuentra en buenas condiciones, por lo que es fácil observar a individuos de esta especie en esta zona.
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Lemos Espinal, J. A. 2003. Ficha técnica de Crotalus viridis. Fichas diagnósticas para 10 especies de anfibios y reptiles mexicanos. Facultad de Estudios Superiores Iztacala, Universidad Nacional Autónoma de México. Bases de datos SNIB-CONABIO. Proyecto No. W002. México, D.F.
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Relevancia de la especie ( шпански; кастиљски )

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Relevancia de la especie

En el Norte de Chihuahua C. viridis coexiste con C. atrox y C. molossus. Desafortunadamente estos organismos por lo general son atacados por el ser humano tan pronto como son vistos. Estas acciones pueden estar mermando las poblaciones de las víboras de cascabel (incluyendo a C. viridis) de esta localidad.
La dieta de C. viridis incluye roedores, aves y sus huevos, lagartijas, ranas, sapos e insectos (Degenhardt et al. 1996), siendo esta una especie importante de la cadena alimentaría en el Desierto Chihuahuense.

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Lemos Espinal, J. A. 2003. Ficha técnica de Crotalus viridis. Fichas diagnósticas para 10 especies de anfibios y reptiles mexicanos. Facultad de Estudios Superiores Iztacala, Universidad Nacional Autónoma de México. Bases de datos SNIB-CONABIO. Proyecto No. W002. México, D.F.
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Distribution ( англиски )

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Continent: Middle-America North-America
Distribution: SW Canada (incl. Alberta, Saskatchewan), W USA (Washington, California, Oregon ?, Montana, South Dakota, New Mexico), Mexico (Baja California Norte, N Chihuahua, NW Coahuila) viridis: the Great Plains to the Rocky Mountains and from S Canada to Mexico (N Sonora, N Chihuahua, N Coahuila);
Type locality: œthe Upper Missouri. Restricted to œthe prairies between the Cannonball and Heart rivers, within 40 km of the Missouri River in North Dakota a location far distant from the previous restriction of the type locality to ˜˜vicinity of Kansas City, Missouri (Schmidt, 1953: 226). nuntius: NE Arizona
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Westliche Klapperschlange ( германски )

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Die Westliche Klapperschlange (Crotalus viridis), auch Prärieklapperschlange, ist eine Art der Klapperschlangen (Crotalus) innerhalb der Vipern (Viperidae), die in den westlichen USA bis nach Kanada im Norden und Mexiko im Süden verbreitet ist. Bis vor ein paar Jahren wurde sie gemeinsam mit der Pazifik-Klapperschlange (Crotalus oreganus) als eine Art geführt.[1]

Merkmale

Allgemeine Merkmale

Die Westliche Klapperschlange ist eine relativ große Art der Klapperschlangen mit einer Durchschnittslänge von etwa 1 Meter, die bisherige Maximallänge beträgt 1,51 Meter.[2] Die Unterart C. v. nuntius bleibt im Regelfall etwas kleiner als die Nominatform.

Schlangengift

Das Toxin der Westlichen Klapperschlange ist ein komplex aufgebautes Gemisch aus verschiedenen Proteinen (Eiweiße), darunter zu finden sind Enzyme wie Proteasen und Peptidasen. Neben hämotoxischer und gewebezerstörender Wirkung zeigt das Gift auch neurotoxische Eigenschaften. Für Bisse dieser Viper stehen spezifische Antivenine zur Verfügung.[3]

Verbreitung und Lebensraum

 src=
Verbreitungsgebiet
 src=
Prärieklapperschlange in Wyoming

Das Verbreitungsgebiet der Schlange umfasst in den Vereinigten Staaten von Amerika einen großen Bereich der Great Plains und reicht im Norden bis Kanada und im Süden bis Mexiko. In Kanada ist sie in den südlichen Gebieten von Saskatchewan und Alberta anzutreffen. In den USA umfasst das Verbreitungsgebiet Teile der Bundesstaaten Idaho, Montana, North Dakota, South Dakota, Wyoming, Nebraska, Colorado, Kansas, Oklahoma, Texas, New Mexico und den äußersten Süden Arizonas. In Mexiko findet sich die Schlange im nördlichen Coahuila und nordwestlichen Chihuahua.[2]

Systematik

Die taxonomische Diskussion um die Westliche Klapperschlange ist derzeit im Fluss, bis vor wenigen Jahren wurde sie gemeinsam mit der Pazifik-Klapperschlange (C. oreganus) als eine Art geführt. Auf der Grundlage von Untersuchungen auf molekularbiologischer Basis durch Ashton und de Queiroz 2001 sowie Pook et al. 2000 wurden die beiden Taxa als Arten getrennt, wobei ein Großteil der Unterarten der Pazifik-Klapperschlange zugeordnet wurde.[4][5][2]

Bei der Westlichen Klapperschlange werden aktuell nur noch zwei Unterarten unterschieden:

  • C. v. nuntius Klauber (Hopi-Klapperschlange) 1935 im Nordosten von Arizona
  • C. v. viridis Rafinesque 1818 im gesamten Verbreitungsgebiet

Literatur

  1. Beispielsweise in Chris Mattison: Rattler! - A natural history of Rattlesnakes. Blandford, London 1996, ISBN 0-7137-2534-6.
  2. a b c J.A. Campbell, W.W. Lamar WW:The Venomous Reptiles of the Western Hemisphere. Comstock Publishing Associates, Ithaca and London 2004, ISBN 0-8014-4141-2.
  3. Franz-Xaver Reichl: Taschenatlas der Toxikologie, Kap. Tiergifte; Nikol Verlag; 2., aktualisierte Auflage. ISBN 978-3868200058.
  4. Viperidae - Crotalinae - 2001 Publications (Memento des Originals vom 26. Dezember 2007 im Internet Archive)  src= Info: Der Archivlink wurde automatisch eingesetzt und noch nicht geprüft. Bitte prüfe Original- und Archivlink gemäß Anleitung und entferne dann diesen Hinweis.@1@2Vorlage:Webachiv/IABot/biology.bangor.ac.uk at Wolfgang Wüster (Memento des Originals vom 25. September 2006 im Internet Archive)  src= Info: Der Archivlink wurde automatisch eingesetzt und noch nicht geprüft. Bitte prüfe Original- und Archivlink gemäß Anleitung und entferne dann diesen Hinweis.@1@2Vorlage:Webachiv/IABot/biology.bangor.ac.uk, School of Biological Sciences, Bangor University
  5. Pook CE, Wüster W, Thorpe RS. 2000. Historical biogeography of the western rattlesnake (Serpentes: Viperidae: Crotalus viridis), inferred from mitochondrial DNA sequence information. Molecular Phylogenetics and Evolution, 15: 269–282. PDF (Memento des Originals vom 21. Juli 2007 im Internet Archive)  src= Info: Der Archivlink wurde automatisch eingesetzt und noch nicht geprüft. Bitte prüfe Original- und Archivlink gemäß Anleitung und entferne dann diesen Hinweis.@1@2Vorlage:Webachiv/IABot/biology.bangor.ac.uk at Wolfgang Wüster (Memento des Originals vom 25. September 2006 im Internet Archive)  src= Info: Der Archivlink wurde automatisch eingesetzt und noch nicht geprüft. Bitte prüfe Original- und Archivlink gemäß Anleitung und entferne dann diesen Hinweis.@1@2Vorlage:Webachiv/IABot/biology.bangor.ac.uk, School of Biological Sciences, Bangor University

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Westliche Klapperschlange: Brief Summary ( германски )

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Die Westliche Klapperschlange (Crotalus viridis), auch Prärieklapperschlange, ist eine Art der Klapperschlangen (Crotalus) innerhalb der Vipern (Viperidae), die in den westlichen USA bis nach Kanada im Norden und Mexiko im Süden verbreitet ist. Bis vor ein paar Jahren wurde sie gemeinsam mit der Pazifik-Klapperschlange (Crotalus oreganus) als eine Art geführt.

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Crotalus viridis ( англиски )

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Crotalus viridis (Common names: prairie rattlesnake,[3][4] Great Plains rattlesnake,[5]) is a venomous pit viper species native to the western United States, southwestern Canada, and northern Mexico. Currently, two subspecies are recognized, including the prairie rattlesnake (Crotalus viridis viridis), the nominate subspecies, and the Hopi rattlesnake (Crotalus viridis nuntius).[3]

Taxonomy

The taxonomic history of this species is convoluted. Previously, seven other C. viridis subspecies were also recognized, including C. v. abyssus, C. v. caliginis, C. v. cerberus, C. v. concolor, C. v. helleri, C. v. lutosus and C. v. oreganus. However, in 2001 Ashton and de Queiroz described their analysis of the variation of mitochondrial DNA across the range of this species. Their results agreed broadly with those obtained by Pook et al. (2000). Two main clades were identified, east and west of the Rocky Mountains, which they argued were actually two different species: on the one hand C. viridis, including the conventional subspecies C. v. viridis and C. v. nuntius, and on the other C. oreganus, including all the other traditional subspecies of C. viridis. The authors retained the names of the traditional subspecies, but emphasized the need for more work to be done on the systematics of C. oreganus.[6][7][8]

Description

Crotalus viridis, closeup of head

This species commonly grows to more than 100 cm (3.3 ft) in length. The maximum recorded size is 151.5 cm (4.97 ft). In Montana, specimens occasionally exceed 120 centimetres (3.9 ft) in length; the species reaches its maximum size in this region. One of the most characteristic features is the presence of three or more, usually four, internasal scales.[6]

Identification characteristics will vary depending on which subspecies is encountered. Generally, prairie rattlesnakes are usually lightly colored in hues of brown. Patches of dark brown are often distributed in a dorsal pattern. A color band may be seen at the back of the eye. The prairie rattlesnake group carries the distinctive triangle-shaped head and pit sensory organs on either side of the head. A key characteristic that can help differentiate a prairie rattlesnake from other rattlesnakes is the presence of two internasals contacting the rostral.[9]

Distribution and habitat

Prairie rattlesnake (Crotalus v. viridis), Hidalgo County, New Mexico (5 Sep 2014)

Prairie rattlesnakes are found in North America over much of the Great Plains, the eastern foothills and some intermontane valleys of the Rocky Mountains, from southwestern Canada through the United States to northern Mexico. In Canada, they occur in Alberta and Saskatchewan; in the US in eastern Idaho, most of Montana (where it is one of 10 snake species and the only venomous one), North Dakota, South Dakota, Wyoming, Nebraska, Colorado, Kansas, Oklahoma, Texas, New Mexico, northeastern Arizona, extreme western Iowa, and in Mexico in northern Coahuila and northwestern Chihuahua. Its vertical range is from 100 m (330 ft) near the Rio Grande to over 2,775 m (9,104 ft) in elevation in Wyoming.[6]

Wright and Wright (1957) and Klauber (1997) both mention Utah as within the range of this species, including maps showing it confined to the extreme southeastern part of the state.[5][10] The type locality is described as "the Upper Missouri [Valley, USA]". An emendation was proposed by H.M. Smith and Taylor (1950) to "Gross, Boyd County, Nebraska."[2]

Habitat characteristics can vary depending on subspecies and range. Generally, western rattlesnakes occupy areas with an abundant prey base. Many subspecies occupy somewhat rocky areas with outcrops serving as den sites. Prairie rattlesnakes have also been known to occupy burrows of other animals.[11] They seem to prefer dry areas with moderate vegetation coverage. Vegetation cover will vary depending on region and subspecies.[9]

Behavior

A Hopi rattlesnake (Crotalus viridis nuntius) in a defensive posture

Prairie rattlesnakes are primarily terrestrial, but occasionally climb in trees or bushes. Some even rest in crevices or caves. They are typically active diurnally in cooler weather and nocturnally during hot weather C. viridis. This species complex is equipped with powerful venom, using about 20-55 percent of venom in one bite, and will defend themselves if threatened or injured. As with other rattlesnake species, prairie rattlesnakes will rapidly vibrate their tails, which produces a unique rasping sound to warn intruders.[12] Prairie rattlesnakes prefer sinuosity while migrating even in human-dominated environments because it leads to a higher chance of survival and better body condition throughout and after migration.

[13]

The venom of the Prairie rattlesnake is a complexly structured mixture of different proteins with enzymes such as proteases and peptidases found among them.[14] Besides the hemotoxin and its tissue destructive effect, the venom also has neurotoxic properties.[15]

Diet

Prairie rattlesnakes, because of their expansive distribution, have a wide array of prey. Generally, this species prefers small mammals, such as ground squirrels, ground nesting birds, mice, rats, small rabbits and prairie dogs. They will occasionally feed on amphibians and reptiles, and sometimes even other snakes. This is more commonly seen in juvenile snakes.[9]

Reproduction

A juvenile Hopi rattlesnake (Crotalus viridis nuntius) in captivity

Prairie rattlesnakes are viviparous and can produce from one to 25 young per reproduction event. The average number of young ranges from four to 12, but can vary greatly due to availability of food and environmental conditions. Males may compete for females during the breeding season, but western rattlesnake females may not necessarily breed every year.[16] They give birth in late summer or early fall, being their breed 22–28 cm long, without the need for parental care. In addition, the young are toxic as soon as they are born. They reach sexual maturity at three years of age. It is also common for females to give birth at communal den sites with the young born between August and October.[9][12]

Conservation status

This species is classified as Least Concern on the IUCN Red List of Threatened Species (v3.1, 2001). Species are listed as such due to their wide distribution, presumed large population, or because they are unlikely to be declining fast enough to qualify for listing in a more threatened category. The population trend was stable when assessed in 2006.[1]

Subspecies

Crotalus viridis nuntius Klauber, 1935,[3] the Hopi rattlesnake, inhabits the United States from northeastern and north-central Arizona, from the New Mexico border to Cataract Creek, including the Little Colorado River basin, the southern section of the Apache Indian Reservation, the Hopi Reservation, and the Coconino Plateau from the southern rim of the Grand Canyon to US Highway 66 in the south.[10]

Crotalus viridis viridis (Rafinesque, 1818),[3] the prairie rattlesnake, inhabits the North American Great Plains from the Rocky Mountains to 96° W and from southern Canada to extreme northern Mexico, including southwestern Saskatchewan, southeastern Alberta, eastern Washington, Idaho in the Lemhi Valley, Montana east of the higher Rockies, southwestern North Dakota, west, central and extreme southeastern South Dakota, western Iowa, central and western Nebraska, Wyoming except for the Rockies, Colorado, central and western Kansas, Oklahoma, extreme southeastern Utah, northeastern Arizona, New Mexico, western and southwestern Texas, northeastern Sonora, northern Chihuahua, northern Coahuila.[10]

See also

References

  1. ^ a b Frost, D.R.; Hammerson, G.A.; Santos-Barrera, G. (2007). "Crotalus viridis". IUCN Red List of Threatened Species. 2007: e.T64339A12771847. doi:10.2305/IUCN.UK.2007.RLTS.T64339A12771847.en. Retrieved 19 November 2021.
  2. ^ a b McDiarmid RW, Campbell JA, Touré TA. 1999. Snake Species of the World: A Taxonomic and Geographic Reference, Volume 1. Washington, District of Columbia: Herpetologists' League. 511 pp. ISBN 1-893777-00-6 (series). ISBN 1-893777-01-4 (volume).
  3. ^ a b c d "Crotalus viridis ". Integrated Taxonomic Information System. Retrieved 28 November 2006.
  4. ^ "Crotalus ". Integrated Taxonomic Information System. Retrieved 28 November 2006.
  5. ^ a b Wright AH, Wright AA. 1957. Handbook of Snakes of the United States and Canada. Ithaca and London: Comstock Publishing Associates. (7th printing, 1985). 1,105 pp. (2 volumes). ISBN 0-8014-0463-0.
  6. ^ a b c Campbell JA, Lamar WW. 2004. The Venomous Reptiles of the Western Hemisphere. Ithaca and London: Comstock Publishing Associates. 870 pp. 1500 plates. ISBN 0-8014-4141-2.
  7. ^ Viperidae - Crotalinae - 2001 Publications. The Arizona subspecies was further split into its own species, Crotalus cerberus. Archived 26 December 2007 at the Wayback Machine at Wolfgang Wüster Archived 25 September 2006 at the Wayback Machine, School of Biological Sciences, Bangor University. Accessed 7 April 2008.
  8. ^ Pook CE, Wüster W, Thorpe RS. 2000. "Historical biogeography of the western rattlesnake (Serpentes: Viperidae: Crotalus viridis), inferred from mitochondrial DNA sequence information". Molecular Phylogenetics and Evolution 15: 269-282. PDF Archived 15 August 2016 at the Wayback Machine at Wolfgang Wüster, School of Biological Sciences, Bangor University. Accessed 11 July 2016.
  9. ^ a b c d Stebbins RC. 2003. A Field Guide to Western Reptiles and Amphibians, Third edition. The Peterson Field Guide Series ®. New York: Houghton Mifflin Company. xiii + 533 pp. ISBN 978-0-395-98272-3.
  10. ^ a b c Klauber LM. 1997 (Reprint). Rattlesnakes: Their Habitats, Life Histories, and Influence on Mankind. Second Edition. Berkeley: University of California Press. 1,580 pp. (2 volumes). (First published in 1956, 1972). ISBN 0-520-21056-5.
  11. ^ Shipley, B.K., D. Chiszar, K.T. Fitzgerald, and A.J. Saviola. "Spatial ecology of Prairie Rattlesnakes (Crotalus viridis) associated with Black-tailed Prairie Dog (Cynomys ludovicianus) colonies in Colorado." Herpetological Conservation and Biology 8, no. 1 (2013): 240-250. http://www.herpconbio.org/Volume_8/Issue_1/Shipley_etal_2013.pdf
  12. ^ a b C. virdis. Accessed 7 October 2009.
  13. ^ Martin, A.E.; Jørgensen, D.; Gates, C.C. (December 2017). "Costs and benefits of straight versus tortuous migration paths for Prairie Rattlesnakes ( Crotalus viridis viridis ) in seminatural and human-dominated landscapes". Canadian Journal of Zoology. 95 (12): 921–928. doi:10.1139/cjz-2017-0031. hdl:1807/80397. ISSN 0008-4301.
  14. ^ Saviola, A.J., Pla, D., Sanz, L., Castoe, T.A., Calvete, J.J. and Mackessy, S.P., 2015. Comparative venomics of the Prairie Rattlesnake (Crotalus viridis viridis) from Colorado: Identification of a novel pattern of ontogenetic changes in venom composition and assessment of the immunoreactivity of the commercial antivenom CroFab®. Journal of proteomics, 121, pp.28-43. https://dx.doi.org/10.1016/j.jprot.2015.03.015
  15. ^ Reichl, Franz-Xaver. 2008. Taschenatlas der Toxikologie: Substanzen, Wirkungen, Umwelt [=Pocket Atlas of Toxicology: Substances, Effects, Environment ]. (Chapter: "Venoms"). Hamburg: Nikol Verlag. ISBN 978-3868200058. (in German).
  16. ^ C. viridis
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Crotalus viridis: Brief Summary ( англиски )

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Crotalus viridis (Common names: prairie rattlesnake, Great Plains rattlesnake,) is a venomous pit viper species native to the western United States, southwestern Canada, and northern Mexico. Currently, two subspecies are recognized, including the prairie rattlesnake (Crotalus viridis viridis), the nominate subspecies, and the Hopi rattlesnake (Crotalus viridis nuntius).

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Crotalus viridis ( шпански; кастиљски )

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Crotalus viridis es una especie de serpiente venenosa nativa del oeste de Estados Unidos, suroeste de Canadá, y norte de México. En la actualidad, dos subespecies son reconocidas (véase más abajo).[3]

Descripción

Esta especie crece hasta 1 m de longitud, siendo su máxima longitud registrada de 1,52 m (Klauber, 1937). En Montana, los especímenes ocasionalmente superan los 12 dm de longitud. Klauber (1972) menciona que la especie alcanza su máximo tamaño en esta región. Una de sus características típicas es la de la presencia de tres o más (normalmente cuatro) escamas internasales.[4]

Identificación

Las características que permiten la identificación varían dependiendo de cual subespecie se encuentre. Generalmente, las serpientes de cascabel del oeste (u occidentales) tienen colores claros con varias tonalidades de café. Manchas de color café más obscuro son comúnmente visibles en un patrón dorsal. Una banda de color puede estar presente detrás del ojo. El grupo de los cascabeles del oeste tiene una cabeza de característica forma triangular y los órganos sensoriales a cada lado de la cabeza. Una característica clave, que permite distinguir las cascabeles del oeste de otras cascabel, es la presencia de dos escamas internasales en contacto con la rostral.[5]

Hábitat

Las características del hábitat varían dependiendo de la subespecie y su área. Generalmente, las cascabeles del oeste ocupan áreas con presa abundante. Muchas subespecies ocupan zonas rocosas con salientes que sirven como guaridas. También es común que ocupen las habitaciones de otros animales. Parecen preferir zonas secas con poca vegetación. Las cascabeles de las praderas también pueden ser encontradas cerca de pantanos, estanques o cerca del agua.[5]

Dieta

Las cascabeles del oeste, debido a su amplia distribución, tienen una gama variada de presas. Generalmente, esta especie prefiere pequeños mamíferos tales como ardilla de tierra, ratón, conejo, perrillo de las praderas. Ocasionalmente se alimenta de anfibios o reptiles, observándose más comúnmente en serpientes jóvenes.[5]

Reproducción

La cascabel del oeste es vivípara y puede producir de 1 a 25 crías por evento reproductivo. El promedio de crías va de 4 a 12 pero varía mucho por la disponibilidad de alimento y otras condiciones ambientales. Hembras de la cascabel no necesariamente criarán cada año.[6]​ Es común que las hembras den a luz en guaridas comunes. La cría nace entre agosto y octubre.[5][7]

Comportamiento

Las cascabeles del oeste permanecen activas normalmente durante el día durante tiempo frío y de noche en tiempo caliente. C. viridis. La especie posee un veneno poderoso y se defenderá si se le amenaza o hiere. Al igual que otras especies de cascabel, vibrara su cola que producirá un sonido característico para advertir a los intrusos, a menos que esté conservando el calor y se enrosque sobre su cascabel.[7]

Rango geográfico

Se le encuentra en América del Norte en gran parte de las planicies, desde el sur de Canadá siguiendo hacia el sur a través de los Estados Unidos hasta el norte de México. En Canadá aparece en Columbia Británica, Alberta y Saskatchewan; en EE. UU. en el este de Idaho, Montana, Dakota del Norte, Dakota del Sur, Wyoming, Nebraska, Colorado, Kansas, Oklahoma, Texas, Nuevo México, el extremo este de Arizona; en México en el norte de Coahuila y norte de Chihuahua. Su rango vertical va desde los 100 m cerca del Río Bravo hasta más de 2,775 m de altura en Wyoming.[4]

Wright y Wright (1957) y Klauber (1997) mencionan Utah dentro del rango de la especie, incluyendo mapas mostrándola confinada al extremo sureste del estado.[8][9]

La localidad tipo se describe como "valle del alto Misuri" (EE. UU.). Una enmienda se propuso por Smith y Taylor (1950) ("Gross, Condado de Boyd (Nebraska).")[2]

Nombres comunes

  • cascabel de las praderas,[3]​ cascabel del oeste,[10]​ cascabel de las planicies,[8], cascabel negro, cascabel común, gran cascabel de la pradera, cascabel de la gran meseta, cascabel manchada, cascabel del pacífico occidental.

Estado de Conservación

La especia está clasificada como del menor riesgo (LC) por la IUCN Red List of Threatened Species (v3.1, 2001).[1]​ Las especies se enlistan como tales debido a su amplia distribución, cuando se presume una elevada población, o debido a que es poco probable que su población disminuya rápidamente como para calificar en una categoría más amenazada. La tendencia de la población es estable.[1]

Subespecies

Subespecies[3]​ Autor del Taxon[3]​ Nombre Común[3]​ Rango Geográfico[9]C. v. nuntius Klauber, 1935 Cascabel Hopi En Estados Unidos desde el noreste y norte-centro de Arizona, desde la frontera con Nuevo México hasta Cateract Creek, incluyendo la cuenca del Río Pequeño Colorado, el sur de la reservación Indígena Apache, la Reserva Hopi, y la Meseta Coconino desde el borde sur del Gran Cañón hasta la U.S. Highway 66 al sur. C. v. viridis (Rafinesque, 1818) Cascabel de la pradera En las grandes planicies de América del Norte desde las Montanas Rocallosas hasta la longitud 96° Oeste. y desde el sur de Canadá al extremo norte de México, incluyendo el suroeste de Saskatchewan, sureste de Alberta, Idaho en el valle de Lemhi, Montana al este de las altas Montañas Rocallosas, suroeste de Dakota del Norte, oeste, centro y extremo sureste de Dakota del Sur, Iowa occidental, centro y oeste de Nebraska, Wyoming excepto por las Rocosas, Colorado, centro y oeste de Kansas, Oklahoma, extremo sureste de Utah, noreste de Arizona, Nuevo México, este y suroeste de Texas, noreste de Sonora, norte de Chihuahua, y norte de Coahuila.

Taxonomía

La historia taxonómica de la especie es compleja. Anteriormente, se reconocían otras siete especies de C. viridis, incluyendo abyssus, caliginis, cerberus, concolor, helleri, lutosus y oreganus. Sin embargo, en el 2001 Ashton y de Queiroz publicaron un estudio detallando su análisis de la variación del DNA mitocondrial entre las diversas subespecies. Sus resultados coincidieron ampliamente con aquellos de Pook et al. (2000). Dos clados principales fueron identificados, al este y oeste de las Montañas Rocallosas, los cuales, ellos alegaron, pertenecían a dos especies diferentes: por un lado C. viridis, incluyendo la subespecie convencional viridis y nuntius, y por otra parte C. oreganus, incluyendo todas las otras subespecies tradicionales de C. viridis. Los autores retuvieron los nombres de las subespecies tradicionales, pero enfatizaron que se requería más trabajo en la sistematización de C. oreganus.[4][11][12]

Referencias

  1. a b c Frost, D.R., Hammerson, G.A. & Santos-Barrera, G. (2007). «Crotalus viridis». Lista Roja de especies amenazadas de la UICN 2015.2 (en inglés). ISSN 2307-8235. Consultado el 26 de julio de 2015.
  2. a b McDiarmid RW, Campbell JA, Touré T. 1999. Snake Species of the World: A Taxonomic and Geographic Reference, vol. 1. Herpetologists' League. 511 pp. ISBN 1-893777-00-6 (series). ISBN 1-893777-01-4 (volume).
  3. a b c d e Sistema Integrado de Información Taxonómica. «Crotalus viridis (TSN 174319)» (en inglés).
  4. a b c Campbell, J.A.; Lamar W.W. (2004). The Venomous Reptiles of the Western Hemisphere. Ithaca & Londres: Comstock Publishing Associates. p. 870. ISBN 0-8014-4141-2. La referencia utiliza el parámetro obsoleto |coautores= (ayuda)
  5. a b c d Stebbins, R.C. 2003. A field guide to western reptiles and amphibians. Third edition. Houghton Mifflin Company, New York, New York, USA.
  6. http://imnh.isu.edu/digitalatlas/bio/reptile/serp/crvi/crvifram.htm
  7. a b «C. viridis». Consultado el 7 de octubre de 2009.
  8. a b Wright AH, Wright AA. 1957. Handbook of Snakes. Comstock Publishing Associates. (7ª impresión, 1985). 1105 p. ISBN 0-8014-0463-0.
  9. a b Klauber LM. 1997. Rattlesnakes: Their Habitats, Life Histories, and Influence on Mankind. Second Edition. 2 volumes. Reprint, University of California Press, Berkeley. ISBN 0-520-21056-5.
  10. Sistema Integrado de Información Taxonómica. «Crotalus (TSN 174305)» (en inglés).
  11. Viperidae - Crotalinae - 2001 Publications at Wolfgang Wüster, School of Biological Sciences, Bangor University. Accesada el 7 de abril de 2008.
  12. Pook CE, Wüster W, Thorpe RS. 2000. Historical biogeography of the western rattlesnake (Serpentes: Viperidae: Crotalus viridis), inferred from mitochondrial DNA sequence information. Molecular Phylogenetics and Evolution, 15: 269-282. PDF at Wolfgang Wüster, School of Biological Sciences, Bangor University. Consultado el 7 de abril de 2008.
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Crotalus viridis: Brief Summary ( шпански; кастиљски )

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Crotalus viridis es una especie de serpiente venenosa nativa del oeste de Estados Unidos, suroeste de Canadá, y norte de México. En la actualidad, dos subespecies son reconocidas (véase más abajo).​

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Crotalus viridis ( баскиски )

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Crotalus viridis Crotalus generoko animalia da. Narrastien barruko Viperidae familian sailkatuta dago.

Erreferentziak

  1. (Ingelesez)IUCN 2012. IUCN Red List of Threatened Species. Version 2012.2. www.iucnredlist.org. 2012ko urriaren 20an eskuratua.
  2. The Species 2000 and ITIS Catalogue of Life

Kanpo estekak

Ikus, gainera

(RLQ=window.RLQ||[]).push(function(){mw.log.warn("Gadget "ErrefAurrebista" was not loaded. Please migrate it to use ResourceLoader. See u003Chttps://eu.wikipedia.org/wiki/Berezi:Gadgetaku003E.");});
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Crotalus viridis: Brief Summary ( баскиски )

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Crotalus viridis Crotalus generoko animalia da. Narrastien barruko Viperidae familian sailkatuta dago.

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Crotalus viridis ( француски )

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Crotalus viridis est une espèce de serpents de la famille des Viperidae[1]. La sous-espèce Crotalus viridis viridis est également nommée Crotale des prairies.

Répartition

 src=
Aire de répartition de l'espèce Crotalus viridis selon l'UICN (consulté le 22 février 2013).

Cette espèce se rencontre[1] :

Sa présence est incertaine dans l'Oregon.

Description

 src=
Crotalus viridis nuntius
CarrizoPlainRattlesnake.jpg

Ce serpent venimeux et vivipare[1] dépasse la taille de 100 cm, avec un maximum connu de 151 cm (Klauber, 1937).

Liste des sous-espèces

Selon Reptarium Reptile Database (13 février 2014)[2] :

Publications originales

  • Klauber, 1935 : A new subspecies of Crotalus confluentus, the prairie rattlesnake. Transactions of the San Diego Society of Natural History, vol. 8, no 13, p. 75-87 (texte intégral).
  • Rafinesque, 1818 : Further accounts of discoveries in natural history in the western states. The American Monthly Magazine and Critical Review, vol. 4, no 5, p. 39-42 (texte intégral).

Notes et références

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Crotalus viridis: Brief Summary ( француски )

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Crotalus viridis est une espèce de serpents de la famille des Viperidae. La sous-espèce Crotalus viridis viridis est également nommée Crotale des prairies.

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Žalioji barškuolė ( литвански )

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Binomas Crotalus viridis

Žalioji barškuolė (lot. Crotalus viridis, angl. Prairie rattlesnake) – barškuolių (Crotalus) genties gyvatė, priklausanti angių (Viperidae) šeimai, duobagalvių angių (Crotalinae) pošeimiui. Labai nuodinga.

Aprašymas

Kūnas apie 1 m ilgio, žaliai ir pilkos spalvos, išilgai nugaros su tamsiomis apskritomis skersinėmis dėmėmis.

Paplitimas

Žalioji barškuolė paplitusi JAV nuo Oklahomos ir Kanzaso iki Ramiojo vandenyno pakrančių. Šiaurėje arealas pasiekia Kanadą, o pietuose – Meksiką. Gyvena įvairiuose biotopuose. Žinomi 9 porūšiai.

Mityba

Gaudo smulkius graužikus (prerinius šuniukus, kiškių jauniklius), taip pat paukščius, varles, driežus ir vabzdžius. Prerinių šuniukų urvus naudoja slėptuvei ir žiemojimui. Vikiteka

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Žalioji barškuolė: Brief Summary ( литвански )

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Žalioji barškuolė (lot. Crotalus viridis, angl. Prairie rattlesnake) – barškuolių (Crotalus) genties gyvatė, priklausanti angių (Viperidae) šeimai, duobagalvių angių (Crotalinae) pošeimiui. Labai nuodinga.

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Crotalus viridis ( португалски )

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Crotalus viridis é uma espécie de serpente venenosa da família Viperidae, presente no Canadá, Estados Unidos da América, e México.[1]

Referências

  1. a b Frost, D.R.; Hammerson, G.A.; Santos-Barrera, G. (2007). «Crotalus viridis». Lista Vermelha de Espécies Ameaçadas. 2007: e.T64339A12771847. doi:. Consultado em 19 de novembro de 2021
  2. McDiarmid RW, Campbell JA, Touré TA. 1999. Snake Species of the World: A Taxonomic and Geographic Reference, Volume 1. Washington, District of Columbia: Herpetologists' League. 511 pp. ISBN 1-893777-00-6 (series). ISBN 1-893777-01-4 (volume).
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Crotalus viridis: Brief Summary ( португалски )

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Crotalus viridis é uma espécie de serpente venenosa da família Viperidae, presente no Canadá, Estados Unidos da América, e México.

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서부방울뱀 ( корејски )

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서부방울뱀(학명: Crotalus viridis)은 살무사과 살무사아과 방울뱀속에 속하는 유독성 의 일종이다. 미국·캐나다 남부·로키산맥·멕시코 북부에 걸쳐서 서식한다. 승명아종인 대초원방울뱀(Crotalus viridis viridis)을 포함한 2갈래의 아종으로 나눠진다.[3]

각주

  1. Crotalus . 미국 통합 분류학 정보 시스템(Integrated Taxonomic Information System, ITIS).
  2. Frost, D.R.; Hammerson, G.A.; Santos-Barrera, G. (2007). “Crotalus viridis”. 《IUCN 적색 목록》 (IUCN) 2007: e.T64339A12771847. doi:10.2305/IUCN.UK.2007.RLTS.T64339A12771847.en.
  3. Crotalus viridis . 미국 통합 분류학 정보 시스템(Integrated Taxonomic Information System, ITIS).
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서부방울뱀: Brief Summary ( корејски )

добавил wikipedia 한국어 위키백과

서부방울뱀(학명: Crotalus viridis)은 살무사과 살무사아과 방울뱀속에 속하는 유독성 의 일종이다. 미국·캐나다 남부·로키산맥·멕시코 북부에 걸쳐서 서식한다. 승명아종인 대초원방울뱀(Crotalus viridis viridis)을 포함한 2갈래의 아종으로 나눠진다.

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