^Wahlberg, N., Leneveu, J., Kodandaramaiah, U., Peña, C., Nylin, S., Freitas, A. V., & Brower, A. V. (2009). Nymphalid butterflies diversify following near demise at the Cretaceous/Tertiary boundary. Proceedings of the Royal Society of London B: Biological Sciences, 276(1677), 4295-4302. (英文)
Brower, A. V. Z. 2000 Phylogenetic relationships among the Nymphalidae (Lepidoptera) inferred from partial sequences of the wingless gene. Proc. R. Soc. Lond. B 267, 1201-1211.
Freitas, A. V. L. & Brown, K. S. J. 2004 Phylogeny of the Nymphalidae (Lepidoptera). Systematic Biology 53, 363-383.
Lamas G ed. 2004. Atlas of Neotropical Lepidoptera. Checklist: Part 4A Hesperioidea - Papiionoidea. Gainesville: Scientific Publishers/Association of Tropical Lepidoptera.
Wahlberg, N., Weingartner, E. & Nylin, S. 2003 Towards a better understanding of the higher systematics of Nymphalidae (Lepidoptera: Papilionoidea). Mol. Phylogenet. Evol. 28, 473-484.
寿建新 周尧 李宇飞. 世界蝴蝶分類名錄. 中國: 陝西科學技術出版社. 2006-04-01. ISBN 9787536936768(中文(简体)).