Lamproscatella bimaculata Hendel 1933

Lamproscatella (Lamproscatella) bimaculata Hendel

Lamproscatella sibilans (in part) of American authors [misidentification; not Haliday, 1833:175].—Sturtevant and Wheeler, 1954:176 [review, as species of Scatella].—Wirth and Stone, 1956:475 [key, distribution in California].—Wirth, 1965:756 [catalog].

Lamproscatella bimaculata Hendel, 1933:53.—Papp, 1975:115 [review, figures of head, wing, male terminalia].

DIAGNOSIS.—Specimens of this species are distinguished from those of similar congeners by the following combination of characters: Specimens averaging smaller than those of L. brunnipennis or L. occidentalis, coloration of facial pollenosity appearing white (white appearance becomes discolored, light golden brown in older and worn specimens); shape of fifth tergum of males subtriangular; merger of surstyli with ventral margin of epandrium indistinguishable; cercal cavity large; posteroventral surface of surstyli with secondary lobe; gonite slender, not produced ventrally.

DESCRIPTION.—Small to moderately small shore flies, length 1.72 to 2.07 mm (averaging 1.85 mm); description generally as for specimens of L. occidentalis except as follows.

Head: Head width-to-height ratio averaging 1:0.69; eye width-to-height ratio averaging 1:1.05; eye-to-cheek ratio averaging 1:0.17. Face tending to be whitish gray, but coloration variable.

Thorax: Coloration of mesonotum mostly grayish brown, slightly subshiny, thinly pollinose. Wing length-to-width ratio averaging 1:0.49; costal vein ratio averaging 1:0.155; M1+2 vein ratio averaging 1:0.63.

Abdomen: Fifth tergum of male subtriangular in dorsal view, apex distinctly pointed or acutely rounded; cercal cavity (Figure 39) large, subequal to one-half length of combined epandrium and fused surstyli; surstyli in posterior view broadly produced as subtruncate ventral processes, each bearing a secondary, acutely pointed anteromedial lobe; gap between surstyli about one-half width of surstylus near apex; aedeagus (Figure 40) acutely pointed apically; aedeagal apodeme sublunate in lateral view; gonite not produced ventrally.

TYPE MATERIAL.—Two syntype males. According to Hendel (1933:54), both males were collected from “Donau-Auen der Lobau bei Wien, Juni.” Hendel also stated that the types were in his collection which was later given to the Naturhistorisches Museum Wien. Through Dr. Ruth Lichtenberg, I was sent the specimens of L. bimaculata from the Wien collection, but none could be identified as either of the two syntype males. One male specimen did bear a single label indicating that it was from the Hendel collection but lacked a locality label.

OTHER SPECIMENS EXAMINED.—CANADA: ALBERTA: Lethbridge, 27 Jun 1968, W. W. Wirth (1 ; USNM). UNITED STATES: ARIZONA: Coconino Co., Happy Jack, 9.7 km N, 13 Jun 1971, B. A. Foote (1 ; KSU). CALIFORNIA: Alpine Co., Forestdale Meadow, 17 Jul 1960, G. G. Moore (1 ; UCD); Inyo Co., Bishop, 28 Jul 1940, D. E. Hardy (1 ; USNM); Kern Co., Onyx, 11.2 km E, 12 Jun 1961, H. F. Howden (1 ; CNC), Tehachapi, 14 Sep 1898 (1 ; ANSP); Los Angeles Co., Lovejoy Lake, Mojave Desert, 10 Oct 1934, A. L. Melander (2 ; USNM); Mono Co., Sardine Creek, 6 Jul 1951, A. T. McClay (1 ; UCD), Sonora Pass, 13 Aug 1960 (1 ; CAS); Nevada Co., Hobart Mills, 1.6 km NW, 20 Jun 1974, W. N. Mathis (1; USNM), Hobart Mills, 8.1 km NW Sage Hen, 20 Jun 1954, H. B. James (1 ; WSU), Sage Hen Creek, 20 Jun–15 Jul, 1954–1970, M. T. James, D. S. Chandler, E. C. Sickels, S. F. Casey, C. Goodpasture, E. C. Sickels (3 , 5 ; UCD, USNM); Placer Co., Lake Tahoe, 6300–7000 feet, 30 Aug 1910 (1 ; USNM); Riverside Co., Palm Springs, 14–28 Dec, 1917–1947, J. C. Bradley, R. Coleman (4 ; CU, USNM), Riverside, 11 Apr 1951, A. L. Melander (1 ; USNM), Temecula, 10 May 1945, A. L. Melander (2 , 1 ; USNM); San Bernardino Co., Barton Store, 3 Sep 1950, A. L. Melander (1 ; USNM), Jenks Lake, 18 Aug 1950, A. L. Melander (2 ; USNM), 1000 Springs, 24 Oct 1946, A. L. Melander (1 ; USNM), Victorville, 16 May 1955, W. R. Richards (1 ; CNC); Santa Barbara Co., Santa Barbara, 16 Apr 1951, K. W. Tucker (1 ; UCD); Sierra Co., Webber Lake, 3 Jul 1964, M. E. Irwin (1 ; UCR), Yuba Pass, 7 Jul 1964, M. E. Irwin (1 ; UCR); Sonoma Co., Cloverdale, J. C. Bradley (2 ; CU); Stanislaus Co., Adobe Creek, 19.3 km W Patterson, 25 Apr 1948, H. B. Leech (1 ; CAS); Tulare Co., Crabtree Meadow, 10,550 feet, 29 Jul 1915 (1 ; ANSP), Siberian Outpost, 9,500—10,500 feet, 31 Jul 1915 (1 ; CAS), Tulare, 9 Sep 1898 (1 ; USNM). COLORADO: Clear Creek Co., Mt. Evans, Echo Lake, 10,600 feet, 20 Jul 1961, B. H. Poole (1 ; CNC); Gunnison Co., Monarch Pass, 21 Jun 1940, A. L. Melander (2 ; USNM); Jackson Co., Cameron Pass, 11 Aug 1955, M. T. James (1 ; WSU); Logan Co., Crook, 15 Jun 1940, A. L. Melander (1 ; USNM); Saguache Co., John Smith Ranger Station, Cochetopa National Forest, 3 Jul 1913, A. K. Fischer (1 ; CAS). IDAHO: Gem Co., Emmett, 15 Apr 1938, H. S. Telford (1 ; USNM). NEVADA: Elko Co., Harrison Pass, 25 Jun 1953, A. B. Gurney (1 , 1 ; USNM), Thomas Creek, Ruby Mountains, 21 Aug 1958, E. Mezger (1 ; USNM); Lander Co., Austin, 12 Aug 1940, D. E. Hardy (1 ; KU); Washoe Co., Mt. Rose, 6000 feet, 7 Jul 1972, C. Goodpasture (2 ; UCD), Pyramid Lake, 27 Jun 1972, C. Goodpasture (1 ; UCD). OREGON: Baker Co., Anthony Lake, 21 Jul 1964, K. Goeden (4 ; OSDA); Harney Co., 29 Jun 1953, A. B. Gurney (5 , 3 ; USNM), Idlewild Campground, 25.8 km N Burns 15 Jun–30 Jul, 1972–1975, W. N. Mathis (6 , 3 ; USNM); Lake Co., Ana Reservoir, 24 Sep 1971, W. N. Mathis (1 ; USNM); Umatilla Co., Tollgate, 3.2 km E, 6 Jul 1954, J. J. Davis (1 ; WSU). UTAH: Box Elder Co., Bear River City, 14 Oct 1953, G. F. Knowlton (1 ; USNM); Davis Co., Farmington, 8 Oct 1950, G. F. Knowlton (1 ; USNM), Syracuse, 10 Sep 1933, J. A. Rowe (2 ; 4; ANSP); Duchesne Co., Neola, 12 Jun 1962, G. F. Knowlton (1 ; UMN); Salt Lake Co., Little Cottonwood Canyon, 8000 feet, 23 Jun 1940, A. L. Melander (2 , 2; USNM); Utah Co., Goshen Pond, 2 Mar 1968, W. N. Mathis (1 ; USNM); Weber Co., Hooper, 20 Sep 1933, J. A. Rowe (1 ; ANSP). WASHINGTON: Grant Co., O'Sullivan Dam, 22 May–30 Oct, 1953–1955, M. T. James, H. G. Davis (22 , 21 ; WSU); Kittitas Co., American River, Hell's Crossing, 14 Jul 1954, M. T. James (1 ; WSU); Klickitat Co., Willow Creek, La Crosse-Hooper, 13 May 1951, J. J. Davis (1 ; WSU); Okanogan Co., Boiling Lake, 6000 feet, 27 Jul 1950, C. L. Cooper (1 , 1 ; WSU); Pierce Co., Mt. Rainier, Berkeley Park, 23 Aug 1934 (1 ; USNM), Mt. Rainier, Paradise Park, Aug 1917, A. L. Melander (1 ; USNM), Mt. Rainier, Yakima Park, 14–19 Aug, 1934–1940, A. L. Melander (2 ; USNM). WYOMING: Albany Co., Snowy Range Mountains, Lake Marie, 13 Aug 1957, G. F. Knowlton (1 ; WSU), Snowy Range Mountains, Libby Flats, 13 Jul–13 Aug 1957, G. F. Knowlton (72 , 60; UMN, USNM, WSU), Mirror Lake, 13 Aug 1957, G. F. Knowlton (14 , 8 ; WSU), Silver Lake, 10,000 feet, 13 Aug 1957, G. F. Knowlton (1 ; UMN), Laramie, 8 Aug 1947, P. B. Lawson (9 , 8 ; KU).

GEOGRAPHIC DISTRIBUTION (Figure 41).—This species has a transboreal, Holarctic distribution. In North America, it occurs west of 103° west longitude between 33° and 49° north latitude.