Comprehensive Description
(
anglais
)
fourni par Smithsonian Contributions to Zoology
Eviota prasina (Klunzinger)
Eleotris prasinus Klunzinger, 1871:481 [type-locality: Koseir, Red Sea].
Allogobius viridis Waite, 1904:177, pl. 23: fig. 3 [type-locality: Lord Howe Island].
Eviota verna Smith, 1958:139, pl. I(J,K,L): fig. 1 [type-locality: Aldabra].
MATERIAL EXAMINED.—907 specimens from 30 major localities, totaling 404 males, 331 females, 172 juveniles and unsexed specimens; total size range 7.0–30.9; largest male 30.9, largest female 26.5; smallest gravid female 10.9.
Lectotype: NFIS 1693, (11.6), female; Koseir, Red Sea, B. Klunzinger.
Other Material: RED SEA POPULATION: GULF OF AQABA: USNM 191716, 1 (14.8), female; Elat, 5–9 Sep 1960, E. Clarke. ANSP 83366, 2 (14.3–16.1), males; Elat, 26 Apr 1950, H. Steinitz, 2050. HUJ E62/39, 1 (14.7), male; Elat, 4 May 1960. USNM 218003, 7 (8.5–15.5), 2 juv., 4 males (15.5), 1 female (10.9); Marsa Muqabila, 29 Jul 1969, V. G. Springer, 69-8. AMS I.20061–002, 2 (11.8, ca. 14.0), males; Ras Burqa, 23 Jul 1969, V. G. Springer, 69–7. DIFNEIN ISLAND: USNM 218004, 2 (13.2, 13.7), 1 juv., 1 male (13.7); south shore Ethiopia, 15 Aug 1969, V. G. Springer, 69-15. DAHLAK ARCHIPELAGO: HUJ E62/3678g, 4 (11.7–14.3), females; Um Aabak, 6 Apr 1962, E. Clark. HUJ E62/507, 39 (7.0–14.9), 17 juv., 10 males (14.9), 12 females (13.8); Entedebir, 7 Apr 1962. HUJ E62/4313, 25 (8.0–15.6), 3 juv., 13 males (15.6), 9 females (14.5); Entedebir, 7 Apr 1962. ZUBAIR ISLAND: USNM 218005, 12 (10.9–15.2), 6 males (15.2), 6 females (14.0); 30 Sep 1967, E. Clark. WESTERN INDIAN OCEAN POPULATION: KENYA: RUSI 3865, 5 (ca. 15.6–19.5), 2 males (19.5), 3 females (15.9); Shimoni; questionable paratypes of E. verna Smith. MOZAMBIQUE: RUSI 3864, 7 (11.0–17.6), 3 males (17.6), 4 females (14.1); Wamezi I.; questionable paratypes of E. verna Smith. RUSI 3867, 2 (16.8, 15.9), male and female; Baixa Pinda; questionable paratypes of E. verna Smith. MADAGASCAR: UMMZ 186039, 22 (16.8–20.4), 11 males (20.4), 11 females (18.6); Tulear, 24 Jul 1964, J. Bardach and L. A. Mauge, B64-24. GRAND COMORE ISLAND: CAS 33256, 3 (11.5–16.4), males; 9 Feb 1975, J. E. McCosker, 75-4. CAS 33465, 15 (9.5–17.7), 2 juv., 6 males (17.7), 7 females (16.6); 13 Feb 1975, J. E. McCosker, 75-8. CAS 33052, 1 (17.5), female; 8 Feb 1975, J. E. McCosker, 75-2. CAS 33604, 19 (7.6–17.8), 5 juv., 9 males (17.8), 5 females (15.5); 15 Feb 1975, J. E. McCosker, 75-11. ALDABRA: RUSI 255, holotype of Eviota verna Smith, (18.6), male. USNM 219261, 9 (12.5–17.9), 7 males (17.9), 2 females (14.1); 3 Sep 1967, H. A. Fehlmann, 67-66. USNM 219266, 2 (15.5–15.6), males; 26 Aug 1967, H. A. Fehlmann, 67-62. USNM 219264, 2 (14.6, 14.3), male and female; 10 Sep 1967, H. A. Fehlmann, 67-74. USNM 219245, 1 (12.0), females; 6 Sep 1967, H. A. Fehlmann, 67-69. USNM 219258, 1 (17.8), male; 4 Sep 1967, H. A. Fehlmann, 67-67. BPBM 22571, 3 (14.8–17.5), 2 males (17.5), 1 female (14.8); 18 Aug 1967, H. A. Fehlmann, 67-56. AMS I.20808-001, 8 (13.1–16.5), 4 males (16.5), 4 females (ca. 14.8); 25 Aug 1967, H. A. Fehlmann, 67-61. USNM 219235, 5 (13.1–16.1), 2 males (16.1), 3 females (15.0); 14 Sep 1967, H. A. Fehlmann, 67-76. USNM 219269, 2 (15.2–15.4), females; 24 Aug 1967, H. A. Fehlmann, 67-60. USNM 219247, 1 (16.4), female; 8 Sep 1967, H. A. Fehlmann, 67-71. AMIRANTES ISLANDS: ANSP 141211, 19 (7.3–16.8), 10 juv., 5 males (16.8), 4 females (13.7); D'Arros I., 6 Mar 1964, J. E. Böhlke, F-91. SEYCHELLES ISLANDS: ANSP 141209, 2 (13.8, 14.2), males; Mahé, 4 Feb 1964, J. E. Böhlke, F-23. ANSP 141210, 12 (11.8–17.1), 8 males (17.1), 4 females (16.3); Mahé, 10 Feb 1964, J. E. Böhlke, F-37. ANSP 141207, 29 (10.2–16.5), 18 males (16.5), 11 females (14.8); Mahé, 2 Feb 1964, J. E. Böhlke, F-17. ANSP 138925, 15 (11.4–15.8), 10 males (15.8), 15 females (14.9); Mahé, 31 Jan 1964, J. E. Böhlke, F-13. USNM 209208, 1 (17.5), male; Mahé, 12 Sep 1954, J.L.B. Smith; questionable paratype of E. verna Smith. RUSI 3866, 8 (14.2–16.4), 3 males (16.4), 5 females (16.1); Mahé. ANSP 141208, 27 (8.7–18.2), 10 juv., 8 males (18.2), 9 females (14.9); Anonyme I., 4 Feb 1964, J. E. Böhlke, F-21. AGALEGA ISLANDS: USNM 219248, 1 (15.8), male; North I., 19 Apr 1976, V. G. Springer, 76-29. USNM 219270, 1 (14.6), male; North I., 17 Apr 1976, V. G. Springer, 76-24. ST. BRANDON SHOALS (collected by V. G. Springer in 1976): USNM 219234, 1, (14.8), female; 30 Mar, 76-1. USNM 219229, 12 (15.8–20.4), 6 males (19.7), 6 females (20.4); 1 Apr, 76-4. USNM 219237, 1 (18.7), female; 2 Apr, 76-6. USNM 219257, 39 (7.9–17.8), 1 juv., 18 males (17.8), 20 females (15.2); 3 Apr, 76-7. AMS I.20804-001, 1 (16.8), male; 5 Apr, 76-9. USNM 219231, 8 (10.0–16.8), 1 juv., 3 males (16.8), 4 females (14.9); 7 Apr, 76-11. USNM 219228, 8 (9.5–19.1), 3 juv., 5 males (19.1); 11 Apr, 76-17. MAURITIUS: RUSI 2276, 19 (10.1–14.9), 2 juv., 7 males (13.9), 10 females (14.9); 26 Mar 1971, T. H. Fraser, THF-SA-43. RUSI 2182, 11 (9.9–16.4), 4 juv., 4 males (16.4), 3 females (14.0); 7 Mar 1971, T. H. Fraser, THF-SA-30. CHAGAS ARCHIPELAGO, DIEGO GARCIA ATOLL (collected by H. A. Fehlmann in 1967): USNM 219254, 3 (11.6–12.6), 2 males (12.6), 1 female (12.1); 11 Jun, 67-2. USNM 219250, 1 (13.8), male; 15 Jun, 67-7. USNM 219259, 1 (ca. 13.5), male; 22 Jun, 67-16. USNM 219251, 5 (12.4–14.8), 4 males (14.8), 1 female (12.4); 9 Jul, 67-38. USNM 219249, 4 (13.6–16.1), 3 males (16.1), 1 female (13.6); 24 Jul, 67-52. SRI LANKA: TRINCOMALEE. (Collected in 1969): USNM 219255, 2 (16.1, 14.9), male and female; 30 Jun, W. F. Smith-Vaniz, 69-125. USNM 219256, 4 (12.8–14.8), 1 male (12.8), 3 females (14.8); 26 Jun, W. F. Smith Vaniz, 69-119. USNM 219262, 1 (15.2), male; 1 Jul, W. F. Smith-Vaniz, 69-127. USNM 219253, 1 (16.0), male; 27 Jun, W. F. Smith-Vaniz, 69-120. USNM 219271, 9 (10.8–18.3), 2 juv., 6 males (18.3), 1 female (13.5); 30 Sep, P. C. Heemstra, 69-278. GULF OF THAILAND: CAS 43808, 39 (17.0–24.2), 12 males (24.2), 27 females (19.5); 8°26′31″N, 100°45′31″E, 25 Jan 1960, R. Bolin, 60-26, GVF Reg. 2037. CAS 43810, 1 (18.8), male; 8°26′N, 100°45′E, 26 Jan 1960, R. Bolin, 60-28, GVF Reg. 2039. INDONESIA-PHILLIPPINE POPULATION: PHILIPPINE ISLANDS: USNM 161214, 1 (16.0), male; Batan, 22 Jul 1909, Albatross. USNM 161215, 4 (13.6–16.6), 3 males (16.6), 1 female (15.4); Batan, 5 Jun 1909, Albatross. USNM 161216, 1 (14.5), female; Batan, 22 Jul 1909, Albatross. USNM 161213, 2 (16.4, 16.9), females; Panay, 4 Feb 1908, Albatross. SU 51566, 4 (ca. 8.6–13.6), 1 juv., 1 male (13.5), 2 females (13.6); Panay, 26 Jun 1953, G. W. Barlow. PALAU ISLANDS: CAS 43765, 7 (11.0–14.5), 1 juv., 5 males (14.5), 1 female (12.0); Kayangel I., 8 Oct 1956, H. A. Fehlmann, sta 161, GVF Reg. 947. INDONESIA: SU 30041, 10 (11.8–15.0), 4 males (15.0), 6 females (14.9); Sangi I., 24 Jun 1929, A. W. Herre, USNM 219246, 5 (11.3–19.2), 4 males (19.2), 1 female (16.4); Moluccas, Ambon, 13 Mar 1974, V. G. Springer, 74-12. AMS I.20803-001, 1 (16.4), male; same data as above. USNM 210962, 2 (15.8–16.2), females; Moluccas, Saparua, 20 Jan 1973, V. G. Springer, Rumphius Exp. 1, sta IP-1. USNM 211039, 1 (17.4), male; Moluccas, Saparua, 18 Jan 1973, V. G. Springer, Rumphius Exp. 1, sta IP-3. NEW GUINEA: USNM 219230, 2 (14.1, 15.9), males; Trobriand I., 17 Jun 1970, B. B. Collette, 1522. SOUTH CHINA SEA-JAPAN POPULATION: SOUTH CHINA SEA: CAS 43799, 16 (7.4–14.6), 11 juv., 3 males (14.6) 2 females (13.2); Pratas Reef, 23 May 1958, R. L. Bolin. Taiwan, Yeh-lin, collected 16–18 May 1968 by V. G. Springer: USNM 219265, 19 (18.7–26.2), 12 males (26.2), 7 females (22.6); VGS 68-24. USNM 219252, 8 (19.5–23.1), 7 males (23.1), 1 female (19.5); VGS 68-25. USNM 219260, 4 (18.1–21.7), females; VGS 68-26. AMS I.20809-001, 5 (17.6–21.7), 2 males (21.7), 3 females (21.4); VGS 68-27. USNM 219267, 6 (15.7–19.0), 4 males (19.0), 2 females (16.9); VGS 68-28. RYUKYU ISLANDS: ANSP 85044, 1 (13.1), female; Aguni Shima, 2 Aug 1945, E. R. Tinkham. ANSP 89822, 1 (17.8), female; Aguni Shima, 29 Jul 1945, E. R. Tinkham. ANSP 85043, 1 (15.2), female; Aguni Shima, 27 Jul 1945, E. R. Tinkham. USNM 219268, 1 (23.9), male; Okinawa, Sate, 9 Apr 1965, C. R. Johnson. USNM 219233, 1 (18.0), male; Okinawa, Sate, 23 Jan 1965, C. R. Johnson. USNM 71452, 132 (9.5–20.5), 15 juv., 65 males (20.5), 50 females (19.9); Okinawa, Luchu I., 1906, Albatross. JAPAN: USNM 219232, 31 (12.1–26.5), 8 juv., 15 males (26.1), 8 females (26.5); Tanegashima I., 1906, Albatross. USNM 219263, 66 (8.3–14.7), 50 juv., 11 males (13.9), 5 females (14.7); Tanegashima I., 1906, Albatross. CAS 43748, 1 (21.6), female; Tanabe Bay, 17 Sep 1955, R. Rofen, sta 238 J-2, GVF Reg. 748. FMNH 83873, 23 (11.8–22.9), 11 juv., 7 males (22.2), 5 females (22.9); Aikawa Rikuzen, 1906, Albatross. LORD HOWE-NORFOLK ISLANDS POPULATION: LORD HOWE ISLAND: AMS I.5880–5884, syntypes of Allogobius viridis Waite, 13 (19.9–30.5), 7 males (30.5), 6 females (24.6); Feb 1903, Waite and McCulloch. USNM 219302, 2 (23.8, 22.4), male and female; removed from AMS I.5880–5884. AMS I.17368-050, 4 (12.6–21.7), 2 males (21.7), 2 females (15.8); Feb 1973, LHI 73-49. BPBM 17569, 2 (15.9, 16.1), 1 juv., 1 female (16.1); 7 Feb 1973, D. Hoese, LHI 73-22. NORFOLK ISLAND: AMS IB.5365, 1 (20.6), female; ANSP 75301, 21 (8.5–25.9), 11 unsexed, 8 males (25.9), 2 females (20.9); Kingston, Feb 1952, M. Laird.
Tentative Identifications: CAS 43815, 10 (14.0–17.9), 7 males (17.9), 3 females (16.5); Apo I., Negros Oriental, 10 May 1960, D. Empeso, D-2, GVF Reg. 2667. USNM 219236, 1 (21.5), female; Taiwan, 5 May 1968, V. G. Springer, 68-18.
DIAGNOSIS.—Pectoral rays branched, usually the eleventh through the sixteenth; spinous dorsal fin elongate or filamentous in males; fifth pelvic ray usually absent, rudimentary when present; genital papilla in male highly fimbriate; a dark circular to chevron-shaped spot on middle of caudal peduncle, about 3 scale rows from caudal fin base; usually 2 weak to moderately developed, dark spots laterally on occipital area, obscure or intense in some populations; 2 weak spots usually on pectoral base; dark subcutaneous bars on posterior portion of trunk number 4/5 or 5/5.
DESCRIPTION.—Comparisons of certain data by populations are given in Tables 9–12. Dorsal fin VI-I,8(5), VI-I,9(158), VI-I,10(78), VI-I,11(1); anal fin I,7(1), I,8(91), I,9(2); pectoral fin 14(3), 15(25), 16(88), 17(85), 18(25), 19(1); pelvic fin I,4(191), I,4 + a rudiment (41); fourth ray of pelvic fin with 4–14 branches averaging 8.0; segments between consecutive branches of the fourth pelvic fin ray number 1–6, usually 1–3, average 1.9; pelvic fin membrane reduced; branched caudal fin rays 11(1), 12(31), 13(31), 14(6); segmented caudal fin rays 16(4), 17(90); lateral scale rows 23(15), 24(73), 25(2); transverse scale rows 6(16), 7(40), 8(3): scales with about 27–35 ctenii, 11–16 primary radii; breast scaleless.
First dorsal fin spine filamentous in males, maximum length, when depressed, may reach to base of seventh dorsal fin ray; second spine rarely elongate; no spinous dorsal elongation in females; pelvic fins usually reach area just anterior to origin of anal fin.
The cephalic sensory pore system is pattern 2. The cutaneous papilla system is pattern B.
Genital papilla in male highly fimbriate along lateral edges, the tip moderately indented and fringed, papilla extends to the first anal spine; female papilla bulbous, reaching at most to anal spine, the tip with 4–6 fingerlike projections.
Gravid females range in size from 10.9–22.6 mm SL.
Vertebrae 10(24) precaudal and 16(23) caudal, total 26(23).
COLOR IN PRESERVATION.—The intensity of prominent color marks varies among individuals within a population as well as among the 5 populations that we recognize (Tables 9–12). There are also differences related to sexual dichromatism and quality of preservation. Based on color marks and meristic characters the 5 populations recognized are tentatively assigned to racial levels of differentiation.
The prominent color pattern of the species (Figure 46a–e) consists of a dark caudal peduncle spot; usually 2 occipital spots and 2 weak pectoral spots; small weak spots on dorsal midline, often obscure; cheek and predorsal areas often spotted or with scattered chromatophores; outer four-fifths of first dorsal fin usually dark; second dorsal fin dusky to dark; anal fin dark, more so than second dorsal fin; caudal fin dusky, with fine spots mostly in upper, basal portion; scale pockets with weak dark margins; 5 dark spots along ventral midline of posterior portion of trunk coalesced with subcutaneous bars.
In order to analyze the significance of numerous color marks on the head and trunk, we evaluated and recorded, by collection and population, the presence, size, and intensity of each mark. These data are reviewed and those indicating populational divergence are discussed below.
Occipital Spots: Two dark spots dorsolaterally behind eye (usually a dense cluster of small to large chromatophores), the anterior one smaller, vertically elongate and often integrated with chromatophores forming a bandlike arrangement of spots over head behind eyes; the posterior spot is round, larger and usually not integrated with transverse band on head. Spots weak to moderate except in fish from Madagascar, Mauritius, and in some from St. Brandon Shoals in the southern portion of the west Indian Ocean population where the chormatophores are coalesced to form an intense black spot. These spots may be obscure in some specimens from Lord Howe and Norfolk Islands. Spots weaker in females.
Cheek Spots: About 4–7 irregular clusters of small to large chromatophores on cheek and anterior part of opercle, usually equal in intensity to occipital spots. Spots more intense in Lord Howe-Norfolk population and less intense in some southern insular areas of the western Indian Ocean. Sometimes the spots are replaced by more or less uniformly scattered chromatophores.
Predorsal Area: Area from origin of first dorsal fin anterior to eyes is usually spotted, with weak to moderate clusters of chromatophores, often suggestive of transverse bars; spots sometimes replaced by scattered chromatophores and sometimes area is pale, except for spots persisting just behind eyes.
Spots along Dorsal Midline of Trunk: A linear series of about 13 dark spots extending from origin of first dorsal fin posteriorly to area just before procurrent caudal rays, present in all populations but faint or obscure in most specimens, particularly the Lord Howe–Norfolk population.
Spots on Pectoral Base: Two dark spots on upper and lower part of pectoral base, consisting of clusters of large chromatophores separated by an area of equally large but paler chromatophores or, less commonly, by an area of fine dark chromatophores, or by a pale unpigmented area; spots sometimes replaced by scattered chromatophores, particularly in the Lord Howe-Norfolk population.
Caudal Peduncle Spot: A conspicuous black circular or chevron-shaped spot, equal to size of pupil or larger, located medially on peduncle, about 3 scale rows anterior to base of caudal fin, overlying dark, posteriormost subcutaneous bar. Caudal peduncle spot small and weak in Red Sea population.
Dark Spots on Ventral Midline: A series of 5 spots, mostly subcutaneous, extend from origin of anal fin posteriorly on peduncle to area below midcaudal peduncle spot.
Subcutaneous Bars: Lower posterior part of trunk with 5 dark subcutaneous bars coalesced with ventral midline spots and somewhat fainter than spots; upper portion of trunk with 4 or 5 bars; when 5 bars are present they are usually continuous with lower bars, but they are sometimes discontinuous along midline of trunk and the upper portion is staggered anteriorly; this condition occurs more so at the first bar above origin of anal fin, or at the fourth bar; the Japanese and Indonesia-Philippine populations almost always have 4 bars in the upper trunk series, the third upper bar located between the third and fourth lower bars. All specimens from Thailand and 1 specimen from Sri Lanka also have a 4 upper–5 lower bar ratio. These bars are usually weak to moderately developed, obscure in some. There are 3 lower and 2 upper subcutaneous bars on belly area, usually weaker than those on peduncle, except in gravid females, where the lower bars are wide and prominent.
Scale Pigmentation: Scale pockets margined with dark chromatophores, weakly developed but more evident on upper part of trunk and more pronounced in males.
First Dorsal Fin: Anterior basal portion from first to fifth spine and about one-fifth of height of fin, pale or with a small dark mark near base of first spine, the outer four-fifths dusky to dark; fin barred in some specimens, the dark area being divided into a lower dusky portion and a darker outer portion. Pale basal area larger in females. Usually 2 or 3 small, intense black spots on membrane above pale basal area between second and fifth dorsal spines, commonly present in Red Sea and Japanese populations, absent in some collections from other populations; pronounced in males and juveniles and absent in large females.
Second Dorsal Fin: Dusky to light brown in most specimens, lighter in females. Some specimens in each population have the outer one-third to one-half uniformly shaded in brown, the basal portion with fine brown chromatophores, the rays mostly clear. Some specimens with brownish shading on lower portion, the outer part with less dense chromatophores. Light round spots are present near the midportion of the fin in some collections over the range.
Anal Fin: Dark brown, always darker than second dorsal fin; usually uniformly colored, less often with the basal portion lighter. Fin with light margin. Females with less intense pigmentation.
Caudal Fin: Usually dusky with small dark and light spots on rays of basal area of fin, more so on upper, basal portion.
Pectoral and Pelvic Fins: Membrane pale, the rays of pectoral fin with weak, dark outline.
SEXUAL DIMORPHIC FEATURES.—The prominent features are the fimbriate genital papilla, filamentous first dorsal fin, and the darker coloration of the male. Specifically, the male has darker occipital spots, scale pocket pigmentation, and second dorsal, anal, and caudal fins. The pale, anterior basal portion of first dorsal fin is enlarged in females. The small intense black spots commonly present on the spinous dorsal fin of juveniles and some adult males are lacking in adult females. Females have enlarged and more prominent subcutaneous bars on the belly, especially in gravid specimens. The general body condition, including the fins, is ragged in the females as compared to the males.
GEOGRAPHIC DISTRIBUTION.—This species occurs in the Red Sea, is widely distributed in the western Indian Ocean, Sri Lanka, Gulf of Thailand, Indonesia, and New Guinea, and ranges northward to Japan and southward to the Lord Howe and Norfolk Islands (Figure 45). It has not been taken on the Great Barrier Reef.
- citation bibliographique
- Lachner, Ernest A. and Karnella, Susan J. 1980. "Fishes of the Indo-Pacific genus Eviota with descriptions of eight new species (Teleostei, Gobiidae)." Smithsonian Contributions to Zoology. 1-127. https://doi.org/10.5479/si.00810282.315
Comprehensive Description
(
anglais
)
fourni par Smithsonian Contributions to Zoology
Eviota prasina (Klunzinger)
Eleotris prasinus Klunzinger, 1871:481 [type-locality; Koseir, Red Sea].—Clark, 1968:5 [in part].
Allogobius viridis Waite, 1904:177, pl. 23: fig. 3 [type-locality: Lord Howe Island].
Eviota verna Smith, 1958:139, fig. 1, pl. I: J-L [type-locality: Aldabra].
MATERIAL EXAMINED.—908 specimens from 96 collections taken from the Red Sea eastward to Japan in the north and Lord Howe and Norfolk Islands to the south; size range 7.0.30–9, largest male 30.9, largest female 26.5, smallest gravid female 10.9. Only type specimens and Red Sea material are listed below.
Lectotype: NFIS 1693 (11.6), female; Koseir, Red Sea, C. B. Klunzinger.
Syntypes of Allogobius viridis Waite: AMS I. 5880–5884, 15 (19.9–30.5), 8 males (30.5), 7 females (24.6); Feb 1903, Waite and McCulloch.
Holotype of Eviota verna Smith: RUSI 255 (18.6), male; Aldabra.
Nontype Material: GULF OF AQABA: USNM 191716, 1 (14.8), female; Elat, 5–9 Sep 1960, E. Clark. ANSP 83366, 2 (14.3, 16.1), males; Elat, 26 Apr 1950, H. Steinitz 2050. HUJ E62/39, 1 (14.7), male; Elat, 4 May 1960. USNM 218003, 7 (8.5–15.5), 2 juv., 4 males (15.5), 1 female (10.9); Marsa Muqabila, 29 Jul 1969, V. G. Springer 69–8. AMS I.20061–002, 2 (11.8, ca. 14.0), males; Ras Burqa, 23 Jul 1969, V. G. Springer 69–7. DIFNEIN ISLAND: USNM 218004, 2 (13.2, 13.7), 1 juv., 1 male (13.7); S shore Ethiopia, 15 Aug 1969, V. G. Springer 69–15. SOUTHERN END OF SINAI PENINSULA: BPBM 18343, 1 (14.6), male; Ras Muhammed, 19 Sep 1974, J. E. Randall. DAHLAK ARCHIPELAGO: HUJ E62/3678g, 4 (11.7–14.3), females; Um Aabak, 6 Apr 1962, E. Clark. HUJ E62/507, 39 (7.0–14.9), 17 juv., 10 males (14.9), 12 females (13.8); Entedebir, 7 Apr 1962. HUJ E62/4313, 25 (8.0–15.6), 3 juv., 13 males (15.6), 9 females (14.5); Entedebir, 7 Apr 1962. ZUBAIR ISLAND: USNM 218005, 12 (10.9–15.2), 6 males (15.2), 6 females (14.0); 30 Sep 1967, E. Clark.
DIAGNOSIS.—Pectoral fin rays branched, usually the eleventh through the sixteenth; spinous dorsal fin elongate or filamentous in males; fifth pelvic fin ray usually absent, rudimentary when present; genital papilla in male highly fimbriate; a dark, circular to chevron-shaped spot on middle of caudal peduncle, about 3 scale rows from caudal fin base; usually 2 weak to moderatedly developed dark spots laterally on occipital area, obscure or intense in some localities; 2 weak spots usually on pectoral fin base; 5 dark subcutaneous spots on ventral midline posterior of anal fin origin.
DESCRIPTION.—Data for certain meristic characters over the range of this species are summarized in Table 1. Other data follow: pelvic fin membrane reduced; branched caudal fin rays 11(1), 12(31), 13(31), 14(6); segmented caudal fin rays 16(4), 17(90); lateral scale rows 23(15), 24(73), 25(2); transverse scale rows 6(16), 7(40), 8(3). Scales with about 27–35 ctenii, 11–16 primary radii.
First dorsal spine in males filamentous, maximum length when depressed, to base of seventh dorsal ray, second spine rarely elongate; no spinous dorsal elongation in females; pelvic fins usually extending just anterior to origin of anal fin; breast scaleless.
The cephalic sensory pore system as in pattern 2. The cutaneous papillae system as in pattern B.
Genital papilla in male highly fimbriate along lateral edges, the tip moderately indented and fringed, papilla extends to first anal spine; female papilla bulbous, reaching at most to anal spine, the tip with 4–6 fingerlike projections.
Gravid females range in size from 10.9 to 22.6 mm SL.
Vertebrae 10(24) precaudal and 16(23) caudal, total 26(23).
COLOR IN PRESERVATION.—The prominent color pattern of the species (Figure 8a-c) consists of a dark caudal peduncle spot, usually 2 occipital spots, and 2 weak pectoral spots; small weak spots on dorsal midline, often obscure; cheek and predorsal areas often spotted or with scattered chromatophores; outer four-fifths of first dorsal fin usually dark; second dorsal fin dusky to dark; anal fin dark, more so than second dorsal fin; caudal fin dusky with fine spots mostly in upper basal portion; scale pockets with weak dark margins; 5 dark spots along ventral midline of posterior trunk integrated with subcutaneous bars. The black marks on the membrane of the spinous dorsal fin (Figure 8a) are well developed and commonly present in the Red Sea and Japanese collections, infrequently present elsewhere.
GEOGRAPHIC DISTRIBUTION.—Aside from the collections listed above for the Red Sea, this species is widely distributed in the western Indian Ocean, Sri Lanka, Gulf of Thailand, Indonesia, New Guinea, northward to Japan, and southward in the Lord Howe and Norfolk Islands, but absent on the Great Barrier Reef.
DISTRIBUTION.—In addition to the collections listed above from the Red Sea, this species has been taken from the Indian Ocean, Sri Lanka, southern Indonesia, Western Australia, Great Barrier Reef, and islands of Oceania.
- citation bibliographique
- Lachner, Ernest A. and Karnella, Susan J. 1978. "Fishes of the genus Eviota of the Red Sea with descriptions of three new species (Teleostei, Gobiidae)." Smithsonian Contributions to Zoology. 1-23. https://doi.org/10.5479/si.00810282.286
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