Définition : Organic matter in soil which has reached a point of stability, where it will break down no further and might, if conditions do not change, remain essentially as it is for centuries, or millennia.
Définition : This organism produces this material or substance, either during its life or after death. A produces B if some process that occurs in A has output B.
Définition : A group of species that exploit the same food resources, and/or use the same feeding or foraging methods. http://en.wikipedia.org/wiki/Guild_(ecology)