Mirabilis albida (Walt.) Heimerl

In the eastern half of the continent, Mirabilis albida is reasonably uniform, usually erect, with lanceolate or narrowly oblong leaves, and fruits with large, wartlike tubercles on ribs and sulci. These fruits are very similar to those of M. nyctaginea, except they are usually yellowish brown rather than reddish brown. On the western plains, M. albida intergrades with M. linearis. R. Spellenberg (1998), in attempting to maintain a fairly uniform M. albida, provided a table distinguishing among it, M. oblongifolia, and M. melanotricha. Distinguishing leaf forms of M. oblongifolia as circumscribed by C. F. Reed (1969) from western races of M. albida becomes untenable, as proposed by B. L. Turner (1993b). Individual specimens are sometimes very different, but in a series of populations distinctions merge. The minute glandular hairs beneath the tufts of larger hairs and the presence of large, shelflike tubercles are fairly consistent throughout the eastern half of the continent. In the southwestern mountains, where M. oblongifolia in the broad sense occurs, and in the northern Rocky Mountains, where M. lanceolata occurs (as these phases in the complex have been named), both features are either inconsistent or absent, the fruits becoming much less warty and more like those of M. linearis. Some phases, such as M. comata, and Allionia pratensis, seem to form reasonably recognizable geographically and ecologically restricted populations and may be worthy of taxonomic recognition, perhaps at the infraspecific level. The type of M. hirsuta has the general form of broad-leaved plains races of M. albida is lightly hispid on basal parts. The exceedingly hispid, broad-leaved forms commonly identified as M. hirsuta from near the eastern base of the Rocky Mountains are here recognized as M. rotundifolia; the hispid narrow-leaved phases from the southwestern plains are included within M. linearis.

Stems 1 -many, erect to decumbent, few or highly branched, sparsely to densely leafy in basal ¹/2 or throughout, 0.8-15 dm, glabrous to puberulent basally in 2 lines or throughout, hairs often awned, or stems villous and often viscid, or sometimes hirsute, hair types often mixed, spreading pubescent. Leaves ascending to spreading at 10-90°; petiole 0-4 cm; blade green to glaucous blue-gray, linear-lanceolate to ovate-lanceolate, ovate, or deltate, 1-11 × 0.6-2.5 cm in lanceolate leaves, 2-9 × 1-6.5 cm in ovate leaves, thin and fleshy to thick and coriaceous, base cuneate to round, truncate, or cordate, apex acute, obtuse, or round, surfaces glabrous or viscid-puberulent, viscid-villous, or hirsute. Inflorescences axillary or terminal, few branched, ± evenly forked and open, or, when axillary, often consisting only of single involucres (and then flowers commonly cleistogamous); peduncle 1-25 mm, puberulent with curled hairs, hispid, villous, or viscid-villous, crosswalls of hairs pale; involucres pale green or sometimes blushed with purple when young, widely bell-shaped, 4-7 mm in flower, 5-15 mm in fruit, sparsely to densely pubescent with small curled hairs or long spreading hairs, often viscid, 50-80% connate, lobes ovate, triangular-ovate, broadly ovate, or occasionally round. Flowers (1-)3 per involucre; perianth white, pink, or deep red-violet, 0.8-1.5 cm. Fruits brown to dark brown with pale tan, brown, or dark brown ribs, obovoid to narrowly obovate and tapering at both ends, 3.5-5.5 mm, pubescent with tufted spreading hairs 0.1-0.5 mm, with or without minute glandular hairs; ribs round or round-angled, (0.3-)0.7-1.5 times width of sulci, 0.5-1 times as wide as high, with tall shelflike tubercles (eastern part of range), smooth or somewhat rugose or moderately tuberculate (western part of range); sulci with prominent, pale, shelflike tubercles (eastern), minutely rugose or with small low warts (western). 2n = 58.

Alta., B.C., Man., Ont., Que., Sask.; Ala., Ariz., Ark., Calif., Colo., Conn., Ga., Ill., Ind., Iowa, Kans., Ky., La., Maine, Mass., Mich., Minn., Miss., Mo., Nebr., Nev., N.J., N.Mex., N.Y., N.Dak., Ohio, Okla., Pa., S.C., S.Dak., Tenn., Tex., Utah, Wis., Wyo.; Mexico.

Flowering late summer-early fall.

Dry meadows, sandy prairies, hillsides, rocky slopes; 0-2600m.

Allionia albida Walter, Fl. Carol., 84. 1788; A. coahuilensis Standley; A. comata Small; A. divaricata Rydberg; A. hirsuta Pursh; A. oblongifolia (A. Gray) Small; A. pauciflora (Buckley) Standley; A. pratensis Standley; A. pseudaggregata (Heimerl) Standley; A. pumila Standley; Mirabilis ciliata (Standley) Standley; M. coahuilensis (Standley) Standley; M. comata (Small) Standley; M. eutricha Shinners; M. grayana (Standley) Standley; M. hirsuta (Pursh) MacMillan; M. lanceolata (Rydberg) Standley; M. oblongifolia (A. Gray) Heimerl; M. pauciflora (Buckley) Standley; M. pseudaggregata Heimerl; M. pumila (Standley) Standley; Oxybaphus albidus (Walter) Sweet; O. coahuilensis (Standley) Weatherby; O. comatus (Small) Weatherby; O. hirsutus (Pursh) Sweet; O. pauciflorus Buckley; O. pseudaggregatus (Heimerl) Weatherby; O. pumilus (Standley) Standley

Allionia coahuilensis Standley, Contr. U. S. Nat. Herb. 12: 347
1909.
Allionia Greggii Standley, Contr. U. S. Nat. Herb.^ 12: 348 1909 nwhnJyhv* mn.huilpMMis: Weatherbv. Proc. Am. Acad. 45: 4^0. J.yiu. Stems few, from a thick woody root, erect, 6-10 dm. high, simple or sparsely branched below the inflorescence, stout, glaucescent, densely puberulent below, short-pilose above; petioles stout, 1.5 cm. long or shorter, very short in the uppermost leaves, pilose; leaf -blades ovate, ovate-oblong, or deltoid-lanceolate, 2.5-5 cm. long, 0.7-2 cm. wide, obtuse or truncate at the base and decurrent, narrowed to the obtuse or acutish apex, entire or undulate, succulent, thick-coriaceous when dry, glaucescent beneath, densely puberulent or short-pilose; inflorescence cymose-paniculate, copiously branched, the branches rather slender, mostly alternate, densely viscid-pilose, bearing numerous reduced bractlike sessile leaves; involucres numerous, 3-flowered, short-pedunculate, 4-5 mm. long at anthesis, about 8 mm. long in fruit, densely viscid-pilose with fulvous hairs, the lobes shorter than the tube, ovate-orbicular, broadly obtuse; perianth 10-12 mm. long, sparsely pilose; stamens 5, exserted; fruit obovoid, 4-5 mm. long, brownish-olivaceous, minutely pilose, the angles broad, smooth, the sides rugulose or short-tuberculate; seed broadly obovoid, 2-3 mm. long, pale yellowish-brown.
Type locality: Saltillo, Coahuila. Distribution: Vicinity of the type locality.

Allionia coahuilensis Standley, Contr. U. S. Nat. Herb. 12: 347
1909.
Allionia Greggii Standley, Contr. U. S. Nat. Herb. 12: 348. 1909. Oxybaphus coahuilensis Weatherby, Proc. Am. Acad. 45: 425. 19 1U. Stems few, from a thick woody root, erect, 6-10 dm. high, simple or sparsely branched below the inflorescence, stout, glaucescent, densely puberulent below, short-pilose above; petioles stout, 1.5 cm. long or shorter, very short in the uppermost leaves, pilose; leaf-blades ovate, ovate-oblong, or deltoid-lanceolate, 2.5-5 cm. long, 0.7-2 cm. wide, obtuse or truncate at the base and decurrent, narrowed to the obtuse or acutish apex, entire or undulate, succulent, thick-coriaceous when dry, glaucescent beneath, densely puberulent or short-pilose; inflorescence cymose-paniculate, copiously branched, the branches rather slender, mostly alternate, densely viscid-pilose, bearing numerous reduced bractlike sessile leaves; involucres numerous, 3-flowered, short-pedunculate, 4-5 mm. long at anthesis, about 8 mm. long in fruit, densely viscid-pilose with fulvous hairs, the lobes shorter than the tube, ovate-orbicular, broadly obtuse; perianth 10-12 mm. long, sparsely pilose; stamens 5, exserted; fruit obovoid, 4-5 mm. long, brownish-olivaceous, minutely pilose, the angles broad, smooth, the sides rugulose or short-tuberculate ; seed broadly obovoid, 2-3 mm. long, pale yellowish-brown.
Type locality: Saltillo, Coahuila. Distribution: Vicinity of the type locality.

Allionia grayana Standley, sp. nov
Oxybaphus nyctagineus latifolius A. Gray, in Torr. Bot. Mex. Bound. Surv. 174, in part. 1859. Allionia nyctaginea latifolia Coult. Contr. U. S. Nat. Herb. 2: 352, in part. 1894. Allionia latifolia Standley, Contr. U. S. Nat. Herb. 12: 350, excludmg synonym. 1909.
Stems erect, 3-12 dm. high, simple or sparsely branched below the inflorescence, glaucescent, glabrous below or bifariously puberulent; petioles slender or stout, 0.5-2 cm. long, the uppermost leaves often sessile; leaf-blades deltoid or rounded-deltoid, sometimes broadly ovate, 2.5-5.5 cm. long, 1.5-3.5 cm. wide, subcordate, truncate, or rounded at the base, rounded or very obtuse at the apex or rarely acute, entire or subundulate, subcoriaceous when dry, often glaucescent beneath, sparsely pilose or glabrous, often ciliolate; inflorescence cymosepaniculate, usually much branched, the branches slender, mostly opposite, viscid-pilose, usually bearing numerous reduced bractlike leaves; involucres numerous, 3-flowered, slender-pedunculate, 4r-5 mm. long in anthesis, 10-14 mm. long in fruit, densely viscid-pilose, the lobes roundedovate, commonly rounded at the apex but sometimes acutish; perianth about 10 mm. long, sparsely pilose, pink; stamens usually 5, exserted; fruit broadly obovoid, 4 mm. long, darkbrown, finely pilose, the angles narrow, tuberculate, the sides finely and closely tuberculate; seed broadly obovoid, 2.5-3 mm. long, pale-brown.
Type collected in western Texas in 1849, Charles Wright 603 (U. S. Nat. Herb. no. 22766). Distribution: Western Texas.

Allionia grayana Standley, sp. nov
Oxybaphus nyctagineus latifolius A. Gray, in Torr. Bot. Mex. Bound. Surv. 174, in part. 1859. Allionia nyctaginea latifolia Coult. Contr. U. S. Nat. Herb. 2: 352, in part. 1894. Allionia latifolia Standley, Contr. U. S. Nat. Herb. 12: 350, excluding synonym. 1909.
Stems erect, 3-12 dm. high, simple or sparsely branched below the inflorescence, glaucescent, glabrous below or bifariously puberulent; petioles slender or stout, 0.5-2 cm. long, the uppermost leaves often sessile; leaf-blades deltoid or rounded-deltoid, sometimes broadly ovate, 2.5-5.5 cm. long, 1.5-3.5 cm. wide, subcordate, truncate, or rounded at the base, rounded or very obtuse at the apex or rarely acute, entire or subundulate, subcoriaceous when dry, often glaucescent beneath, sparsely pilose or glabrous, often ciliolate; inflorescence cymosepaniculate, usually much branched, the branches slender, mostly opposite, viscid-pilose, usually bearing numerous reduced bractlike leaves; involucres numerous, 3-flowered, slender-pedunculate, 4^5 mm. long in anthesis, 10-14 mm. long in fruit, densely viscid-pilose, the lobes roundedovate, commonly rounded at the apex but sometimes acutish; perianth about 10 mm. long, sparsely pilose, pink; stamens usually 5, exserted; fruit broadly obovoid, 4 mm. long, darkbrown, finely pilose, the angles narrow, tuberculate, the sides finely and closely tuberculate; seed broadly obovoid, 2.5-3 mm. long, pale-brown.
Type collected in western Texas in 1849, Charles Wright 603 (U. S. Nat. Herb. no. 22766). Distribution: Western Texas.

Allionia albida Walt. Fl. Car. 84. 1788
Calymenia albida ~Nutt. Gen. 1: 26. 1818. Oxybaphus albidus Sweet, Hort. Brit. 2: 429. 1827. Calymenia granulala Raf. Aut. Bot. 15. 1840.
Oxybaphus angustifolius Choisy, in DC. Prodr. 13 2 : 433, in part. 1849. Not 0. angustifolius Sweet. 1827.
Mirabilis nyctaginea albida Heimerl, Beitr. Syst. Nyct. 22. 1897. Mirabilis albida Heimerl, Ann. Cons. Jard. Geneve 5: 182. 1901. Mirabilis albida uni flora Heimerl, Ann. Cons. Jard. Geneve 5: 182. 1901. Allionia bracteata Rydb. Bull. Torrey Club 29: 690. 1902.
Allionia decumbens Rydb. Bull. Torrey Club 29: 692. 1902. Not A. decumbens Spreng. 1825. Allionia lanceolata Standley, Contr. U. S. Nat. Herb. 12: 355, in part. 1909. Not A. lanceolata Rydb. 1902.
Allionia lanceolata unifiora Standley, Contr. U. S. Nat. Herb. 12: 355. 1909.
Stems solitary or few, 2—12 dm. high, erect or decumbent, stout or slender, simple or sparsely branched below the inflorescence, glaucous or glaucescent, glabrous or bifariously puberulent, the internodes elongate or often very short; petioles 5 mm. long or shorter, stout, usually wanting; leafblades mostly narrowly lanceolate, sometimes linear to ovate, 4.5—10 cm. long, 0.3-2.8 cm. wide, acute to longattenuate at the base, gradually narrowed to the obtuse or acutish apex, entire or subsinuate, usually thick and succulent, bright-green, or often very glaucous beneath., glabrous or very sparsely short-pilose; inflorescence often wholly of axillary involucres in young plants, in mature plants cymose-paniculate, copiously branched, the branches slender or stout, opposite or alternate, puberulent or shortly viscid-pilose, often bearing very numerous reduced bractlike leaves; involucres 1-3-flowered, slender-pedunculate, about 4 mm. long at anthesis, 8-14 mm. long in fruit, densely or sparsely viscidpilose, the lobes short, rounded; perianth 8-10 mm. long, pale-pink, sparsely pilose; stamens 3-5, exserted; fruit obovoid, 5 mm. long, dark-olivaceous, short-hirtellous, the angles broad and coarsely tuberculate or composed of distinct tubercles, the sides also tuberculate; seed obovoid, 3.5 mm. long, pale yellowish-brown.
Typej locality: South Carolina.
Distribution: Meadows and hillsides, South Carolina and Georgia to Tennessee, Kansas, and Texas.

Allionia albida Walt. Fl. Car. 84. 1788
Calymenia albida Nutt. Gen. 1: 26. 1818. Oxybaphus albidus Sweet, Hort. Brit. 2: 429. 1827. Calymenia granulata Raf. Aut. Bot. 15. 1840.
Oxybaphus angustifolius Choisy, in DC. Prodr. 13 2 : 433, in part. 1849. Not 0. angustifolius
Sweet. 1827. Mirabilis nyctaginea albida Heimerl, Beitr. Syst. Nyct. 22, 1897. Mirabilis albida Heimerl, Ann. Cons. Jard. Geneve 5: 182. 1901. Mirabilis albida uni flora Heimerl, Ann. Cons. Jard. Geneve 5: 182. 1901. Allionia bracleata Rydb. Bull. Torrey Club 29: 690. 1902.
Allionia decumbens Rydb. Bull. Torrey Club 29: 692. 1902. Not A. decumbens Spreng. 1825. Allionia lanceolata Standley, Contr. U. S. Nat. Herb. 12: 355, in part. 1909. Not A. lanceolata
Rydb. 1902. Allionia lanceolata uniflora Standley, Contr. U. S. Nat. Herb. 12: 355. 1909.
Stems solitary or few, 2-12 dm. high, erect or decumbent, stout or slender, simple or sparsely branched below the inflorescence, glaucous or glaucescent, glabrous or bifariously puberulent, the internodes elongate or often very short ; petioles 5 mm. long or shorter, stout, usually wanting; leaf-blades mostly narrowly lanceolate, sometimes linear to ovate, 4.5-10 cm. long, 0.3-2.8 cm. wide, acute to long-attenuate at the base, gradually narrowed to the obtuse or acutish apex, entire or subsinuate, usually thick and succulent, bright-green, or often very glaucous beneath, glabrous or very sparsely short-pilose; inflorescence often wholly of axillary involucres in young plants, in mature plants cymose-paniculate, copiously branched, the branches slender or stout, opposite or alternate, puberulent or shortly viscid-pilose, often bearing very numerous reduced bractlike leaves; involucres 1-3-flowered, slender-pedunculate, about 4 mm. long at anthesis, 8-14 mm. long in fruit, densely or sparsely viscid-pilose, the lobes short, rounded; perianth 8-10 mm. long, pale-pink, sparsely pilose; stamens 3-5, exserted; fruit obovoid, 5 mm. long, dark-olivaceous, short-hirtellous, the angles broad and coarsely tuberculate or composed of distinct tubercles, the sides also tuberculate; seed obovoid, 3.5 mm. long, pale yellowish-brown.
Type locality: South Carolina.
Distribution: Meadows and hillsides, South Carolina and Georgia to Tennessee, Kansas, and Texas.

Allionia ciliata Standley, Contr. U. S. Nat. Herb. 12: 345
1909.
Oxybaphus aggregates A. Gray, in Torr. Bot. Mex. Bound. Surv. 168. 1859. Not O. aggregates
Vahl, 1806. Allionia deltoidea Standley, Contr. U. S. Nat. Herb. 13: 405. 1911. Oxybaphus ciliatifolius Weatherby, Proc. Am. Acad. 49: 492. 1913.
Stems few, from a slender woody root, 2-4 dm. high, rather slender, copiously branched, glaucous, glabrous or bifariously puberulent, the lower internodes short, the upper elongate; petioles stout, 0.5-2 cm. long, sparsely pilose with stout white hairs; leaf-blades elongatedeltoid, ovate-deltoid, or deltoid-lanceolate, 0.5-1.8 cm. wide, entire or subundulate, succulent, coriaceous when dry, glaucous or glaucescent beneath, sparsely pilose or glabrate, long-ciliate, at least near the base; inflorescence of numerous small dense cymes, or of axillary involucres in young plants, the branches slender, puberulent and pilose; involucres 3-flowered, shortpedunculate, about 4 mm. long at anthesis and 8 mm. in fruit, densely white-pilose, the lobes short, broadly ovate or ovate-orbicular, acute or acutish, at least in age; perianth about 1 cm. long, very sparsely pilose or glabrous, the limb about 15 mm. broad; stamens 5, exserted; fruit broadly obovoid, 3.5-4 mm. long, dark-brown, sparsely and minutely pilose, the angles broad, obscurely rugulose, the sides also slightly rugulose; seed globose-obovoid, 2 mm. long, pale-brown.
Type locality: Smiths Run, western Texas. Distribution : Western Texas and Coahuila.

Allionia lanceolata Rydb. Bull. Torrey Club 29: 691. 1902
Allionia sessilifolia Osterhout, Bull. Torrey Club 32: 611. 1905.
Stems few or solitary, erect or ascending, 4—10 dm. high, stout, simple or sparsely branched below the inflorescence, sparsely puberulent or glabrate below, short-pilose above with mostly viscid hairs, the internodes short or elongate; leaf -blades sessile or on very stout petioles 4 mm. long or shorter, lanceolate or lance-ovate, 3.5—10 cm, long, 1-4 cm. wide, usually rounded at the base but sometimes obtuse or acute, gradually narrowed to the obtuse or acutish apex, entire or subsinuate, bright-green, or glaucescent beneath, glabrous or sparsely short-pilose, usually ciliolate; inflorescence paniculate, much branched, the branches slender, opposite, shortly viscid-pilose; involucres slender-pedunculate, about 4 mm. long at anthesis, in age about 1 cm. long, densely viscid-pilose, the lobes oval, rounded at the apex; flowers usually 3 in each involucre, the perianth about 8 mm. long, sparsely pilose, pink; stamens 5, exserted; fruit obovoid, 5 mm. long, dark-olivaceous, finely hirtellous, the angles narrow, smooth, the sides rugose or short-tuberculate ; seed obovoid, 2.5-3 mm, long, brownish-stramineous.
Typb locality: Estes Park, Larimer County, Colorado. Distribution: Southern Wyoming and northern Colorado.

Allionia lanceolata Rydb. Bull. Torrey Club 29: 691. 1902
Allionia sessilifolia Osterhout, Bull. Torrey Club 32: 611. 1905.
Stems few or solitary, erect or ascending, 4^10 dm. high, stout, simple or sparsely branched below the inflorescence, sparsely puberulent or glabrate below, short-pilose above with mostly viscid hairs, the internodes short or elongate; leaf -blades sessile or on very stout petioles 4 mm. long or shorter, lanceolate or lanceovate, 3.5-10 cm. long, 1-4 cm. wide, usually rounded at the base but sometimes obtuse or acute, gradually narrowed to the obtuse or acutish apex, entire or subsinuate, bright-green, or glaucescent beneath, glabrous or sparsely short-pilose, usually ciliolate; inflorescence paniculate, much branched, the branches slender, opposite, shortly viscid-pilose; involucres slender-pedunculate, about 4 mm. long at anthesis, in age about 1 cm. long, densely viscid-pilose, the lobes oval, rounded at the apex; flowers usually 3 in each involucre, the perianth about 8 mm. long, sparsely pilose, pink; stamens 5, exserted; fruit obovoid, 5 mm. long, dark-olivaceous, finely hirtellous, the angles narrow, smooth, the sides rugose or short-tuber culate ; seed obovoid, 2.5-3 mm. long, brownish-stramineous.
TypB locality: Bstes Park, Larimer County, Colorado. Distribution: Southern Wyoming and northern Colorado.

Allionia pauciflora (Buckl.) Standley
Oxybaphus paucifiorus Buckl. Proc. Acad. Phila. 1862: 7. 1862. Allionia gigantea Standley, Contr. U. S. Nat. Herb. 12: 348. 1909. Oxybaphus giganleus Weatherby, Proc. Am. Acad. 49: 492. 1913.
Stems solitary or few, 5-15 dm. high, erect or decumbent, very stout, simple or sparsely branched below the inflorescence, green or glaucescent, densely puberulent throughout, the internodes usually short; petioles 5 mm. long or shorter, very stout, most of the leaves sessile; leaf-blades ovate-oblong to rhombic-ovate, ovate-deltoid, or rounded-ovate, 3.5-10 cm. long, 1.5-7.5 cm. wide, subcordate or rounded at the base and often abruptly short-decurrent, rounded or obtuse at the apex, entire or subsinuate, bright-green, or glaucescent beneath, thick and succulent, densely puberulent or sometimes glabrate; inflorescence paniculate, copiously or sparsely branched, the branches slender or stout, opposite, densely viscid-puberulent or short-pilose, bearing few or numerous reduced bractlike leaves; involucres slenderpedunculate, mostly 3-flowered, about 4 mm. long at anthesis, 8-12 mm. long in fruit, densely or sparsely pilose, the lobes short, rounded; perianth about 8 mm. long, pale-pink, shortpilose, the limb about 12 mm. broad; stamens 3-5, long-exserted; fruit obovoid, 4-5 mm. long, darkolivaceous, hirtellous, the angles broad, coarsely tuber culate or composed of large distinct tubercles, the sides also tuberculate; seed obovoid, 3 mm. long, brownish-stramineous.
Type) locality : On the San Saba River, north of Fort Mason, Texas. Distribution: Oklahoma and Texas.

Allionia pauciflora (Buckl.) Standley
Oxybaphus pauciflorus Buckl. Proc. Acad. Phila. 1862: 7. 1862. Allionia gigantea Standley, Contr. U. S. Nat. Herb. 12: 348. 1909. Oxybaphus giganteus Weatherby, Proc. Am. Acad. 49: 492. 1913.
Stems solitary or few, 5-15 dm. high, erect or decumbent, very stout, simple or sparsely branched below the inflorescence, green or glaucescent, densely puberulent throughout, the internodes usually short; petioles 5 mm. long or shorter, very stout, most of the leaves sessile; leaf-blades ovate-oblong to rhombic-ovate, ovate-deltoid, or rounded-ovate, 3.5-10 cm. long, 1.5-7.5 cm. wide, subcordate or rounded at the base and often abruptly short-decurrent, rounded or obtuse at the apex, entire or subsinuate, bright-green, or glaucescent beneath, thick and succulent, densely puberulent or sometimes glabrate; inflorescence paniculate, copiously or sparsely branched, the branches slender or stout, opposite, densely viscid-puberulent or short-pilose, bearing few or numerous reduced bractlike leaves; involucres slenderpedunculate, mostly 3-flowered, about 4 mm. long at an thesis, 8-12 mm. long in fruit, densely or sparsely pilose, the lobes short, rounded; perianth about 8 mm. long, pale-pink, shortpilose, the limb about 12 mm. broad; stamens 3-5, long-exserted; fruit obovoid, 4-5 mm. long, darkolivaceous, hirtellous, the angles broad, coarsely tuberculate or composed of large distinct tubercles, the sides also tuberculate; seed obovoid, 3 mm. long, brownish-stramineous.
Type locality: On the San Saba River, north of Fort Mason, Texas. Distribution: Oklahoma and Texas.