↑Liem, K. F. & Stewart, D. J., 1976. Evolution of the scale-eating cichlid fishes of Lake Tanganyika: a generic revision with a description of a new species. Bulletin of the Museum of Comparative Zoology v. 147 (núm. 7): 319-350, Pls. 1-2.
↑Boulenger, G. A., 1898. Report on the fishes recently obtained by Mr. J. E. S. Moore in Lake Tanganyika. Proceedings of the General Meetings for Scientific Business of the Zoological Society of London 1898 (pt 3): 494-497.