More info on this topic. More info for the terms:
climax,
forest,
hardwood,
successionObligate Climax Species
Spruce pine is very shade tolerant. Seedlings and saplings can grow
where available light is as low as 250 foot-candles (2,778 lux) and are
common in many parts of the southern mixed-hardwood forest where light
intensity is less than 1000 foot-candles (11,111 lux) at noon on a
summer day [
2]. It is usually only found in late succession hardwood
stands of magnolia, beech, or other climax species. In these stands, it
is represented by all stages of growth [
2,
15,
16]. Where it is found in
younger seral stands, it has usually become established in the shade of
loblolly or shortleaf pines [
9,
16]. According to Platt and Schwartz
[
24], however, spruce pine appears to capitalize on large-scale
disturbance caused by hurricanes. They state that advance recruits in
localized light gaps that are capable of rapid growth at high light
intensities will capture space in the canopy following large-scale
disruption by hurricanes. Hirsh and Platt found that age structures of
spruce pine tend to consist of discrete age classes corresponding to
dates of hurricanes [
25].