Carlana eigenmanni (Meek 1912)

Lives exclusively near the shoreline, river backwaters or other stagnant biotopes between 35 and 85 m elevation. Found near vegetation, feeding mainly on filamentous algae and occasionally on aquatic insects (Ref. 36880).

Lives exclusively near the shoreline, river backwaters or other stagnant biotopes between 35 and 85 m elevation. Found near vegetation, feeding mainly on filamentous algae and occasionally on aquatic insects (Ref. 36880).

Carlana eigenmanni (Meek)

Cheirodon eigenmantii Meek, 1912:70 [original description; La Junta, Costa Rica].

Carlia eigenmanni.—Meek, 1914:108 [new genus; listed from La Junta].

Carlana eigenmanni.—Strand, 1926:54 [new name for preoccupied Carlia].

Rhoadsia eigenmanni.—Eigenmann and Myers, 1929:463 [description of specimen from La Junta], pl. 73: fig. 1 [photograph].—Grey, 1947 [correction of FMNH number of type].—Böhlke, 1958:178 [compared with Hyphessobrycon tortuguerae; discussion of generic status].—Miller, 1966:785 [listed from Lake Nicaragua to Atlantic slope of Costa Rica fide Rivas and Bussing].—Bussing, 1967:236 [listed from Costa Rica].—Astorqui, 1971:20 [listed from Rio Tisla and Rio Osayo, Nicaragua; description].

MATERIAL EXAMINED

*FMNH 7683, holotype, a male 52.8 mm, Costa Rica: La Junta, S. E. Meek, Apr. 1912.

*FMNH 43150, Costa Rica: La Junta, S. E. Meek, 8 Apr. 1912; 1 specimen 56.4 mm.

*FMNH 7861, Costa Rica: La Junta, S. E. Meek, 17 Apr. 1913; 3 specimens 46.4–53.9 mm.

*USNM 74240, Costa Rica: La Junta, S. E. Meek; 1 specimen 53.6 mm. (maxillary, premaxillary and dentary of right side cleared and stained).

*CAS(SU) 61380, Nicaragua: Rio Frio, about 1.5 mi above San Carlos, in bayou with sluggish current, G. S. Myers, 12 Feb. 1963; 4 specimens 21.0–26.9 mm (1 cleared and stained).

*CAS(SU) 61369, Nicaragua: Rio Frio, 1.5 mi above San Carlos, 0.4 mi above Esperanza, small sluggish stream into Rio Frio, G. S. Myers; 8 specimens 21.6–35.8 mm.

*GCRL 5126, Nicaragua: Rio Osayo at IAH, 9 km N of Costa Rica border, L. R. Rivas and I. Astorqui, 13 July 1960; 2 specimens 40.0–43.1 mm.

*GCRL 5125, Nicaragua: Rio Isala at road from Managua to Juigalpa, 8 km W of Juigalpa, L. R. Rivas and I. Astorqui, 28 June 1960; 1 specimen 37.2 mm.

Meek (1912) in describing Cheirodon eigenmanni listed 1 specimen, FMNH 7583, 67 mm; collected April 1912 at La Junta; subsequently (1914), when he placed the species in Carlia, he listed 6 specimens from the type-locality. Grey (1947) corrected the number on the type to 7683 and listed 1 unnumbered paratype. We presume that the specimen Grey considered as paratype is FMNH 43150, because it is the only other FMNH specimen collected April 1912. However, since Meek mentioned no paratypes in the original description we do not accept paratype status for any specimens.

Eigenmann and Myers (1929) listed a specimen 73.0 mm from La Junta, coll. Meek, and stated that it had minute hooks on the anal-fin rays. We have been unable to locate their specimen after searching the CAS, FMNH, and USNM collections.

In the following description, proportions for juveniles are given first, followed by adults, separated by a semicolon. In listing meristics, counts for juveniles and adults are combined. Juveniles in this case are arbitrarily defined by the presence of only 2 maxillary teeth and consist of 6 specimens from CAS(SU) 61369 and 4 specimens from CAS(SU) 61380. All specimens are included in tables.

DESCRIPTION.—Standard length of specimens examined 21.0–29.8; 34.3–56.4 mm. Body elongate, compressed laterally; greatest body depth 32.0–39.5; 35.8–41.2. Predorsal body profile convex or slightly convex with a slight concavity at nape. At anterior dorsal-fin base, body profile angles ventrally to posterior termination of dorsal-fin base. Body profile straight or somewhat convex between posterior dorsal-fin base and adipose-fin base. Posterior to adipose fin, body profile straight or slightly concave to dorsal procurrent caudal-fin rays. Distance from eye to dorsal-fin origin 38.9–41.5; 39.6–41.2; distance from dorsal origin to end of caudal peduncle 49.0–52.4; 51.9–56.3. Ventral profile in juveniles rounded to anus, steepest inclination ventral to jaws; profile protruding ventrally its greatest distance midway between pelvic-fin insertion and anal-fin origin. Ventral profile in adults somewhat convex to a vertical from pectoral-fin base, then becomes more convex to pelvic-fin base; posterior to pelvic-fin base, body profile convex to anal-fin origin. Body profile along anal-fin bases straight or slightly concave in juveniles, straight or somewhat convex in adults; between posterior anal-fin termination and ventral procurrent caudal-fin rays, body profile concave. Caudal peduncle depth 8.3–10.1; 9.5–11.6; caudal peduncle length 8.7–10.9; 9.5–10.5.

Head length 24.9–27.6; 23.1–25.2. Eye diameter 10.8–12.7; 8.5–10.8. Snout length 4.9–6.9; 6.1–7.1. Least bony interorbital width 7.2–9.0; 8.4–8.9. Maxillary short in juveniles, long in adults, sloping ventrally and posteriorly, forming an angle of about 50–60 degrees to longitudinal body axis. Upper-jaw length 7.5–8.2; 8.6–11.9. All teeth uniserial. Maxillary of juveniles with 2 proximal teeth having 5–8 cusps (median cusps may be slightly enlarged); maxillary of adults with 2 proximal teeth as in juveniles but also with up to 8 more conical teeth along its ventral margin. Premaxillary teeth with anterior cutting edge beginning nearer to tooth base than cusps on posterior cutting edge; apices of teeth angled toward midline in adults. Dentary teeth 7–9, broad, with 8–12 cusps (except posteriormost 2 or 3 small teeth with 3 to 5 cusps); cutting edge straight or slightly rounded with median cusps broader, becoming narrower and smaller laterally. No teeth on vomer, palatines, or pterygoids.

Fontanel moderately large in juveniles, that part anterior to the epiphyseal bar a little more than two-thirds length of fontanel posterior to bar; in adults, that part anterior to bar about one and one-half times as long as width of fontanel immediately posterior to bar. Gill rakers moderate, 14–18, usually 16. Circumorbital bones well ossified, in juveniles iniraorbital 3 with a broad naked area ventrally and posteriorly, in adults it contacts preopercle below but with a naked area posteriorly.

Scales moderately large, cycloid with concentric circuli and 0–5 radii on exposed posterior field. Lateral line incomplete with 5–9, usually 8–9, perforated scales; lateral line with a slight ventral curve. Lateral scales 35–41, usually 38 (holotype with 38); these scales not always regularly arranged, therefore counts not always accurate. Scales above lateral line 8–9, usually 8; scales below lateral line 7–8, usually 7. Predorsal scales about 17. Axillary scale present dorsal to pelvic-fin origin; caudal fin scaleless; 6 or 7 scales forming a sheath along anterior base of anal fin.

Dorsal fin with 2 unbranched rays and 8–9, usually 9, branched rays. Dorsal-fin origin anterior to anal-fin origin, posterior to pelvic-fin origin, nearer eye than caudal-fin base. Distance from tip of snout to dorsal-fin origin 53.4–56.7; 52.0–55.0. Third or fourth ray of dorsal fin longest, with posterior rays shorter, forming a slightly convex posterior margin to fin; longest rays extended as a filament in large adults, length of longest ray 29.0–33.2; 31.4–44.0.

Anal fin with 4 unbranched rays and 26–31, usually 29, branched rays. Origin of anal fin behind midpoint of standard length 58.5–61.7; 57.6–61.1. Fourth through eighth or ninth rays slightly longer than posterior rays, forming a protruding fin margin anteriorly and a straight margin posteriorly. Adult males with dorsally recurved or straight spinules on posterolateral surface of posterior branches of anal-fin rays; these spinules usually on third and fourth unbranched rays and on first through sixth to twelfth branched rays. Spinules occur from fin base to near fin tip in first through fourth to seventh branched rays (spinules sometimes sparsely scattered toward distal ray tip), spinules on posterior successive rays cover less area, usually present from anal-fin base to first ray branch or to about midway between ray base and first ray branch. One to several spinules per bony ray segment. (Note: These are not hooks, but spinules as described in Weitzman and Thomerson, 1970.)

Pectoral fin with 1 unbranched ray and 9–12, usually 12, branched rays. Pectoral fins reach to or just anterior to pelvic-fin origin. Distance from tip of snout to dorsal end of pectoral-fin base 26.9–29.8; 24.4–26.6; length of pectoral fin from base to tip of longest ray 17.8–21.3; 19.6–21.2.

Pelvic fin with 1 unbranched ray and 7–8, usually 7, branched rays, distal tip reaching slightly beyond anal-fin origin. Distance from tip of snout to pelvic-fin origin 45.3–48.5; 41.6–45.9; pelvic-fin length 16.7–19.3; 16.7–19.8. First or second through fifth pelvic-fin rays with small spinules (occasionally only on first branched ray); 1 or 2 spinules per bony ray segment.

Caudal fin with 10/9 principal rays (9/8 in 1 specimen); fin forked, not split to base. Caudal fin symmetrical, without hooks or spinules. Dorsal and ventral procurrent caudal rays equal in size.

Precaudal vertebrae 18–19, usually 18, total vertebrae 36–38, usually 37.

Color in alcohol: Ground color light yellowish brown or pale brown. An extremely thin dark line extends along midline. Melanophores under scale margins form reticulate pattern dorsal to midline; melanophores scattered over nape and head; below midline melanophores present to just above anal-fin base and occasionally along myomere junctions between midline and anal-fin base. Melanophores usually less numerous in smaller specimens. Adults with a series of about 8–10 vaguely defined longitudinal bars along body, formed by patches of larger melanophores under scales. Young with 2 small diffuse humeral spots composed of large melanophores; adults with 1 humeral spot under 5 or 6 scales present in area of eighth or ninth lateral scales, posterior to faint “pseudotympanum.” Young with a dark blotch on caudal peduncle, extending slightly onto caudal fin; adults with a bar along midline from just anterior to a vertical from adipose fin, widening at end of caudal peduncle, then extending as a thin line to distal tip of middle caudal-fin rays. Small melanophores scattered on interradial membrane of all fins, often concentrated along fin rays, not on them, except in middle caudal-fin rays. Small specimens with cheek silvery; peritoneum silvery except where intestine is visible, forming an inverted dark gray “comma” mark.

Range: According to Miller (1966; fide Bussing and Rivas), C. eigenmanni is found from Lake Nicaragua to the Atlantic slope of Costa Rica. The type locality is La Junta, on the Rio Reventazon at an altitude of approximately 65 meters (Meek, 1914).

COMPARISONS.—The generic placement of this species has been discussed above. Carlana eigenmanni can be immediately distinguished from all other Central American characids with a single premaxillary tooth series by its number (26–31) of branched anal-fin rays (see Table 3), shape of the dentary teeth, and from all except Phenagoniates macrolepis by the presence of a fully toothed maxillary in adults. See Figures 17–21 for illustration of changes occurring during growth.

Phenagoniates Eigenmann and Wilson

Phenagoniates Eigenmann and Wilson, 1914, in Eigenmann, Henn, and Wilson, 1914:2 [type-species Phenagoniates wilsoni by monotypy].

Phanagoniates Eigenmann, 1915:43 [invalid spelling emendation].

Monotypic, separated from other Central American cheirodontin genera by an elongate body, large tricuspid teeth, a long anal fin (42–50 branched rays, all others treated here with 31 or fewer), origin of anal fin anterior to dorsal-fin origin, low number of pelvic-fin rays (i,5 versus i,7 in all others), and no adipose fin.

A search of the literature indicates that Phenagoniates may be related to Leptagoniates Boulenger, differing only in the latter's possession of a small adipose fin and possession of a complete lateral line. We have examined Xenagoniates Myers and found it to possess an adipose fin and ectopterygoid teeth. We also examined Paragoniates, which also may be related to Phenagoniates, but which has an adipose fin, is stouter, and has a long, slender fully toothed maxillary. Reasonable estimation of relationships of these genera would entail an investigation beyond the scope of this paper. We do not consider Phenagoniates close to other genera discussed in this paper and believe that it lends credence to our conviction that the Cheirodontinae is not a natural group.

Phenagoniates macrolepis (Meek and Hildebrand)

Roeboides macrolepis Meek and Hildebrand, 1913:84 [original description; Rio Cupe, Boca de Cupe, Panama].—Grey, 1947:183 [type listed], fig. 53 [holotype figured].

Phenagoniates wilsoni Eigenmann in Eigenmann, Henn, and Wilson, 1914:2 [original description; Manigru].

Phanagoniates wilsoni.—Eigenmann, 1915:43 [description; spelling emended], pl. V: fig. 1.

Phanagoniates macrolepis.—Meek and Hildebrand, 1916:272 [description; P. wilsoni considered a synonym].—Eigenmann, 1922:128 [listed from Atrato and Tuyra basins].—Breder, 1927:177 [listed; variation; range], 163 [in key].—Jordan, Evermann, and Clark, 1930:95 [listed].—Hildebrand, 1938:249 [range].

Phenagoniates macrolepis.—Myers, 1942:90 [compared with Xenagoniates bondi; note on spelling of Phenagoniates].—Schultz, 1944:311 [listed from several localities in Venezuela].—Miller, 1966:784 [listed; range].

MATERIAL EXAMINED

*FMNH 7590, holotype, 45.1 mm, Panama: Rio Cupe, Boca de Cupe, Darien, S. E. Meek and S. F. Hildebrand, 12 Feb. 1912.

*USNM 78665, Panama: Rio Cupe, Boca de Cupe, Darien, S. E. Meek and S. F. Hildebrand, 24 Feb. 1912; 3 specimens 34.9–41.0 mm.

USNM 78661, Panama: Rio Cupe, Cituro, Darien, S. E. Meek and S. F. Hildebrand, 25 Feb. 1912; 1 specimen 25.9 mm (condition poor).

*USNM 78662, Panama, Rio Yape, Darien, S. E. Mark and S. F. Hildebrand, 6 Mar. 1912; 1 specimen 42.4 mm.

*USNM 78664, Panama: Rio Aruza, Darien, S. E. Meek and S. F. Hildebrand, 26 Feb. 1912; 1 specimen 39.7 mm.

*USNM 208517, Panama: Rio Sambu, Sta. BNW–20, approx. 7°52′N, 78°O5′W, 25 Mar. 1967; 4 specimens, all damaged, approximately 30.0–43.0 mm (1 cleared and stained).

*USNM 208547, Panama: Rio Membrillo, Sta. BNW–9, approx. 8°41′N, 77°41′W, 22 Mar. 1967; 3 specimens 41.4–42.1 mm (1 damaged and unmeasurable).

*USNM 208546, Panama: Rio Morte at Hydro Station, Sta. BNW–8, approx. 8°54′N, 77°33′W, 16 Mar. 1967; 3 specimens 33.4–33.6 mm (1 damaged and unmeasurable).

*FMNH 56541, Colombia: Certegui, C. Wilson, 1913; 1 specimen 23.8 mm (paratype of P. wilsoni).

*FMNH 56542 Colombia: Truando, C. Wilson, 1913; 1 specimen 30.3 mm (paratype of P. wilsoni).

We have examined paratypes of Phenagoniates wilsoni and agree with previous authors that P. wilsoni is synonymous with P. macrolepis.

Grey (1947) listed 5 paratypes of P. macrolepis; this is an error as no paratypes were designated in the original description. In one lot of P. macrolepis (USNM 78661) there are 2 fishes; 1 of these is P. macrolepis, and the other specimen is in poor condition and not identified here. This small fish is a “cheirodontin,” with a single row of elongate conical teeth on the premaxillary and another similar row along the entire anterior border of the maxillary. The dentary teeth are elongate, with a large median cusp and 2 smaller cusps, 1 on each side. The anal-fin base is short and the ventral procurrent caudal-fin rays are elongate. The specimen is too damaged for an accurate identification or description.

Specimens used in description are included in tables.

DESCRIPTION.—Standard length of specimens examined 33.4–45.1 mm. Body elongate, compressed laterally; greatest body depth 26.0–29.8. Predorsal body profile somewhat convex; profile arches dorsally just anterior to dorsal-fin base. Body profile straight or very slightly convex between posterior dorsal-fin base termination and procurrent caudal-fin rays; adipose fin absent. Distance from eye to dorsal-fin origin 44.8–47.0; distance from dorsal-fin origin to end of caudal peduncle 45.1–49.1. Ventral body profile gently rounded from jaws to anus; steepest inclination ventral to jaws. Ventral body profile protrudes ventrally its greatest distance just anterior to or just behind pelvic-fin insertion. Body profile along anal-fin base straight or very slightly convex; between posterior anal-fin termination and procurrent caudal-fin rays, body profile concave. Caudal-peduncle depth 8.7–10.2, peduncle length 5.8–7.1.

Head length 19.5–21.8. Eye diameter 8.0–9.5. Snout length 4.6–4.9. Least bony interorbital width 5.6–6.2. Maxillary sloping ventrally and posteriorly, forming an angle of 70–80 degrees to longitudinal body axis; upper-jaw length 6.3–6.9. All teeth tricuspid (except for smallest dentary and maxillary teeth, which are conical), with median cusp larger; all teeth in a single series. Premaxillary, anterior maxillary, and dentary teeth arise perpendicular to jaw bones and then curve posteriorly just proximal to cusps. Maxillary with 8–11, usually 10, teeth; premaxillary with 6–7, usually 7, teeth (holotype with 7); dentary with 10–12 teeth. No teeth present on vomer, palatines, or pterygoids.

Fontanel moderate, length of that part anterior to epiphyseal bar just over one-third length of that part posterior to bar. Gill rakers short, 12–14 (holotype not counted). Circumorbital bones well ossified, infraorbital 3 wide, contacting preopercle ventrally and posteriorly.

Scales moderately large, cycloid with concentric circuli, and about 2–6 radii on exposed posterior field. Lateral line with a slight ventral curve, incomplete, with 10–15 perforated scales. Lateral scales about 41–42 (holotype damaged); scales above lateral line 6–7, usually 7; scales below lateral line 6–7. Predorsal scales about 20–22. Scale sheath along entire base of anal fin. Large axillary scale present dorsal to pelvic-fin insertion; caudal fin scaled at base.

Dorsal fin with 2 unbranched rays and 7–8 branched rays. Dorsal-fin origin posterior to anal-fin origin, about equidistant from eye and caudal base. Distance from tip of snout to dorsal-fin origin 55.4–59.3. Third, fourth, or fifth dorsal-fin ray longest with rays posterior to fifth abruptly shorter; length of longest ray 22.8–25.4.

Anal fin with 4 unbranched rays and 45–50, usually 49, branched rays. First unbranched ray not always visible externally. Origin well anterior to midpoint of standard length 41.5–45.5. Fourth through eighth or ninth anal-fin rays slightly longer than successive posterior rays, forming a slightly protruding margin anteriorly and a straight margin posteriorly. No anal-ray hooks present.

Pectoral fin with 1 unbranched ray and 11–12, usually 11, branched rays. Pectoral fins reach well beyond pelvic-fin insertion, almost to anal-fin origin. Distance from tip of snout to dorsal end of pectoral-fin base 20.2–22.4 and length of pectoral fin from base to tip of longest ray 17.1–21.7.

Pelvic fin with i,5 rays in all specimens, distal tip reaching just to anal-fin origin. Distance from tip of snout to pelvic-fin insertion 33.0–34.8; pelvic-fin length 8.8–10.2. No pelvic-ray hooks present.

Caudal fin with 10/9 principal rays in all specimens; fin forked, not split to base. Caudal-fin lobes symmetrical. No hooks on caudal-fin rays. Dorsal and ventral procurrent caudal rays usually of equal size.

Precaudal vertebrae 13–14, usually 13. Total vertebrae 41–43, usually 42.

Color in alcohol: Ground color pale brown, nape dark brown. Cheeks with few melanophores; tip of lower jaw often dark brown. Back and sides above midline with reticulate pattern of small melanophores. Broad stripe of small melanophores present from just behind opercle to caudal peduncle, extending to tips of middle caudal-fin rays; ventral to stripe, melanophores sparsely distributed along belly and sides, more concentrated posteriorly (vaguely reticulate pattern anteriorly, following myomere junctions above anal fin). Dorsal fin with a few melanophores along rays and with melanophores scattered on interradial membranes, more concentrated distally. Caudal fin with melanophores on borders of rays and on interradial membranes, more concentrated on middle caudal rays. Anal fin with melanophores on interradial membranes. Pectoral and pelvic fins with a few melanophores on borders of fin rays.

Range: Phenagoniales macrolepis is found in Central America only in eastern Panama, in the drainages of the Rio Sambu, Rio Tuira, and Rio Chucunaque (a major tributary of the Rio Tuira), in Darien Province. The species was found by Eigenmann (1915) in the Atrato basin in Colombia. Phenagoniates macrolepis is actually a northern South American fish which has invaded the extreme eastern drainages of Panama; it does not seem common, judging from collections we have examined (this could simply be due to insufficient sampling).

COMPARISONS.—Phenagoniates macrolepis is a distinctive species which cannot be confused with any other recorded Central American cheirodontin. Its long anal fin, dorsal-fin position, and large tricuspid teeth set it clearly apart from the other species discussed in this work.

Saccoderma Schultz

Saccoderma Schultz, 1944:314 [type-species Saccoderma melanostigma Schultz, 1944, by original designation].

We have found only 1 collection of Saccoderma species from Central America. There are 15 specimens of Saccoderma in SU 48851 (Panama: Veraguas Prov. vicinity of Santiago. Coll. T. D. White, 2 March 1956), along with 7 specimens of Cheirodon dialcpturus. Caudal squamation and dentary teeth place these 15 specimens in Saccoderma, but since they are immature we were unable to establish their specific identity.

We illustrate the caudal-fin squamation (Figure 26) of S. hastata (FMNH 56383, Colombia, Soplaviento), as an aid for future identification of the genus from Panama. (Note that the lower [ventral] caudal-fin rays of the holotype of S. hastata are somewhat malformed and swollen; this is not true of other specimens examined). The dentary teeth (Figure 25) and modified saclike caudal-fin scales (Figure 26) place Saccoderma clearly apart from other genera included in this paper. In addition, adult males of at least 3 species in the genus have antrorse hooks on the rays of the lower caudal-fin lobe.

Loftin (1965) states that he found Saccoderma species in Panama on the Atlantic slope in the Chagres basin and on the Pacific slope from the Rio Bayano basin and coastal streams near the Canal Zone. We were unable to find Saccoderma in Loftin's Panama material at the ANSP or the National Museum of Natural History.

Sumario

Las especies conocidas de peces quirodontinos de la américa central, caracinos con solo una fila de dientes premaxilaries, están revisados. La clave para su identificación está procedida. La revista incluye miembros de los géneros Cheirodon, Carlana, Phenagoniates, y Saccoderma. Todas estas especies están describidas y están illustradas con excepción de la forma de Saccoderma de la américa central que solo se conoce en la forma juvenil y puede ser que pertenezcan a una especie indescribable. Dos neuvas especies de Cheirodon están describidas, C. dialepturns de las cuencas pacificas del oeste central y oeste de Panamá, incluyendo la parte de la Provincia Coclé oeste, hasta al Río Coto, Costa Rica, y C. mitopterus del Río Coclé del Norte, una cuenca atlántica de Panamá central.

Rhoadsia eigenmanni se encuentra en Carlana, el nombre génerico disponible por el ya utilizado Carlia. Los géneros Odontostilbe, Pseudocheirodon, y Compsura están sinonimozados con Cheirodon basado en una reexaminación de sujetos usados antes para definir estos géneros asi como una nueva evaluación del concepto del género de peces caracinos. Finalmente, unas razones están presentadas para considerar los Cheirodontinae polyphylético.

Carlana eigenmanni és una espècie de peix de la família dels caràcids i de l'ordre dels caraciformes.

Carlana is a genus of freshwater fish in the family Characidae. It contains the single species Carlana eigenmanni, which is found in Costa Rica, Nicaragua and Panama. The maximum standard length is 5.4 centimetres (2.1 in).

C. eigenmanni is found in freshwater environments in Central America in the Pacific and Atlantic drainages from Nicaragua to Panama. This species specifically lives close to the shoreline, river backwaters, and motionless region of habitats. They can be found near vegetation. They feed mainly on algae and sometimes on aquatic insects.

The fish is named in honor of ichthyologist Carl H. Eigenmann (1863-1927), who increased the knowledge of the Characins.

Carlana eigenmanni es una especie de peces de la familia Characidae en el orden de los Characiformes, es la única especie del género Carlana.

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Carlana eigenmanni Carlana generoko animalia da. Arrainen barruko Actinopterygii klasean sailkatzen da, Characidae familian.

艾氏頭脂鯉，為輻鰭魚綱脂鯉目脂鯉亞目脂鯉科的其中一個種。分布於中美洲尼加拉瓜及巴拿馬淡水流域，體長可達5.4公分，棲息在海拔35至85公尺的溪流、洄水區，以藻類及水生昆蟲為食，生活習性不明。