Mordella

Sommaire

• Mordella abbreviata Solier, 1851
• Mordella aculeata Linnaeus, 1758
• Mordella acuticauda Champion, 1891
• Mordella adipata Lea, 1917
• Mordella adnexa Ermisch, 1970
• Mordella aeneosignata Blair, 1931
• Mordella aeruginosa Champion, 1891
• Mordella africana Franciscolo, 1955
• Mordella albiventris Maeklin, 1875
• Mordella albodispersa Píc, 1933
• Mordella alboguttata Solier, 1851
• Mordella albomacaculata Lucas, 1857
• Mordella albonotata Maeklin, 1875
• Mordella albopicta Champion, 1891
• Mordella alboscutellaris Píc, 1937
• Mordella alboscutellata Lea, 1895
• Mordella albosparsa Gemminger, 1870
• Mordella albosuturalis Liljeblad, 1922
• Mordella ambrensis Píc, 1942
• Mordella amoena Maeklin, 1875
• Mordella analis Champion, 1891
• Mordella andina Fairmaire & Germain, 1863
• Mordella angeliquae (Leblanc, 2002)
• Mordella angulata LeConte, 1878
• Mordella anticelunulata Píc, 1936
• Mordella apicalis Píc, 1929
• Mordella apicata Lea, 1902
• Mordella apicetestacea Píc, 1942
• Mordella apicicornis Champion, 1891
• Mordella apphabetica Lea, 1917
• Mordella approximata Píc, 1923
• Mordella aradasiana Patti, 1840
• Mordella arcuata Champion, 1891
• Mordella arcuatefasciata Píc, 1941
• Mordella argenteoguttata Champion, 1922
• Mordella argenteonotata Píc, 1936
• Mordella argenteosuturalis Píc, 1937
• Mordella argentifera Fairmaire, 1849
• Mordella argentipunctata Curtis, 1845
• Mordella argyropleura Franciscolo, 1942
• Mordella armeniaca Odnosum, 2004
• Mordella astuta Champion, 1891
• Mordella atrata Melsheimer, 1845
• Mordella atricolor Champion, 1891
• Mordella atripes Píc, 1936
• Mordella atroapicalis Píc, 1927
• Mordella auratonotata Píc, 1941
• Mordella aureofasciata Píc, 1917
• Mordella aurolineata Ray, 1936
• Mordella auromaculata Chûjô, 1935
• Mordella auronotata Lea, 1917
• Mordella auropubescens Ray, 1936
• Mordella auroviolacea Ray, 1936
• Mordella australis Boisduval, 1835
• Mordella badiipennis Champion, 1891
• Mordella baeri Píc, 1936
• Mordella baidensis Blackburn, 1891
• Mordella banahaosa Píc, 1926
• Mordella basifulva Quedenfeldt, 1886
• Mordella batjanensis Píc, 1941
• Mordella batteni (Plaza Infante, 1986)
• Mordella bella Waterhouse, 1878
• Mordella biauronotata Lea, 1931
• Mordella bicolor Fairmaire & Germain, 1863
• Mordella bicoloriceps Píc, 1936
• Mordella bicoloripes Píc, 1923
• Mordella bicoloriventris Píc, 1936
• Mordella bidenominata Píc, 1936
• Mordella bifasciata Fabricius, 1801
• Mordella biformis Champion, 1891
• Mordella binhana Píc, 1928
• Mordella binotata Píc, 1941
• Mordella bipunctata Germain, 1827
• Mordella biquadrinotata Píc, 1936
• Mordella bisbisignata Píc, 1931
• Mordella bisignatipennis Píc, 1942
• Mordella bistrimaculata Píc, 1936
• Mordella bistrinotata Píc, 1928
• Mordella bivittata Ray, 1930
• Mordella blanchardi Solier, 1851
• Mordella blanda Lea, 1917
• Mordella borealis LeConte, 1862
• Mordella bovieri Píc, 1920
• Mordella brachyura Mulsant, 1856
• Mordella brancuccii Horák, 1995
• Mordella brasiliana Maeklin, 1875
• Mordella braueri Kolbe, 1910
• Mordella brevicornis Méquignon, 1946
• Mordella brevis Lea, 1902
• Mordella brevistylis Liljeblad, 1922
• Mordella bribiensis Lea, 1921
• Mordella brincki Franciscolo, 1965
• Mordella bruchi Píc, 1930
• Mordella brusteli (Leblanc, 2002)
• Mordella caatellanii Franciscolo, 1949
• Mordella calodema Lea, 1917
• Mordella calopasa Lea, 1917
• Mordella caloptera Lea, 1917
• Mordella canellina Fairmaire, 1897
• Mordella capillosa Liljeblad, 1945
• Mordella cara Blackburn, 1893
• Mordella caracaensis Píc, 1929
• Mordella carinata Smith, 1883
• Mordella carnoti Píc, 1937
• Mordella caroli Lea, 1896
• Mordella castaneipennis Fairmaire & Germain, 1863
• Mordella castellanii Franciscolo, 1949
• Mordella cata Champion, 1891
• Mordella celebensis Píc, 1923
• Mordella charkrabandhui Chûjô, 1964
• Mordella chevrolati Champion, 1891
• Mordella chrysophora Lea, 1917
• Mordella cinerea Gebl, 1830
• Mordella cinereoatra Liljeblad, 1945
• Mordella cinereoplagiata Blair, 1931
• Mordella cineroatra Liljeblad, 1945
• Mordella cingulata Champion, 1891
• Mordella clavicornis Kirby, 1818
• Mordella communis Waterhouse, 1878
• Mordella compacta Fairmaire, 1895
• Mordella comptei Plaza, 1977
• Mordella confusa Maeklin, 1875
• Mordella conjuncta Shchegolvera-Barovskaya, 1931
• Mordella conradti Píc, 1936
• Mordella consobrina Maeklin, 1875
• Mordella conspecta Lea, 1917
• Mordella corporaali Píc
• Mordella corvina Lea, 1917
• Mordella crassipes Champion, 1891
• Mordella cristovallensis Montrouzier, 1855
• Mordella cuneata Lea, 1902
• Mordella cuneiformis Ermisch, 1968
• Mordella curta Píc, 1927
• Mordella curticornis Ermisch, 1968
• Mordella curtipennis Píc, 1936
• Mordella curvipalpis Stshegolvera-Barovskaja, 1930
• Mordella decemguttata Fabricius, 1801
• Mordella decorata Maeklin, 1875
• Mordella defectiva Walker, 1859
• Mordella denudata Fairmaire, 1906
• Mordella deserta Casey, 1885
• Mordella detracta Pascoe, 1876
• Mordella dilaticornis Champion, 1891
• Mordella dimidiata Champion, 1891
• Mordella disparilis Champion, 1917
• Mordella distincta Lea, 1895
• Mordella divergens Ray, 1939
• Mordella diversepubens Píc, 1941
• Mordella diversinotata Ray, 1936
• Mordella diversiventris Píc, 1942
• Mordella dodonaeae Montrouzier, 1860
• Mordella dumbrelli Lea, 1895
• Mordella duplicata Schilsky, 1895
• Mordella durvillei Boisduval, 1835
• Mordella dybasi Ray, 1944
• Mordella elegantula Csiki, 1915
• Mordella elongatula McLeay, 1887
• Mordella enerosa Píc, 1936
• Mordella erythrocephala Champion, 1891
• Mordella erythrura Fairmaire & Germain, 1863
• Mordella evanescens Norman, 1949
• Mordella fairmairei Píc, 1908
• Mordella fallaciosa Shchegolvera-Barovskaya, 1931
• Mordella fasciata Thomson, 1864
• Mordella fascifera LeConte, 1878
• Mordella felix Waterhouse, 1877
• Mordella festiva Lea, 1895
• Mordella flavicans McLeay, 1887
• Mordella flavifrons Champion, 1891
• Mordella flavimana Marseul, 1876
• Mordella flaviventris Smith, 1883
• Mordella flavofasciata Champion, 1891
• Mordella flavolineata Champion, 1891
• Mordella flavonotata Champion, 1891
• Mordella flavopunctata Castelnau, 1833
• Mordella flexuosa Fairmaire & Germain, 1863
• Mordella fluctuosa Champion, 1891
• Mordella fruhstorferi Píc, 1936
• Mordella fuliginosa Maeklin, 1875
• Mordella fulvonotata Maeklin, 1875
• Mordella fulvopilosa Píc, 1936
• Mordella fulvosignata Fairmaire & Germain, 1863
• Mordella fumosa Fairmaire & Germain, 1863
• Mordella funerea Pascoe, 1876
• Mordella funesta Fald, 1837
• Mordella fuscipilis Champion, 1895
• Mordella fuscocinerea Fall, 1907
• Mordella geniculata Píc, 1928
• Mordella gounellei Píc, 1924
• Mordella goyasensis Píc, 1929
• Mordella grandis Liljeblad, 1922
• Mordella graphiptera Champion, 1895
• Mordella gratiosa Plaza & Compte, 1981
• Mordella griseopubens Píc, 1923
• Mordella griseosuturalis Píc, 1936
• Mordella gutianshana (Fan & Yang, 1995)
• Mordella guttatipennis Píc, 1927
• Mordella guyanensis Píc, 1929
• Mordella haemorrhoidalis Fabricius, 1801
• Mordella haitiensis Ray, 1939
• Mordella hamata Fabricius, 1801
• Mordella hamatilis McLeay, 1887
• Mordella hananoi Nakane & Nomura, 1950
• Mordella hananomi Kônô, 1928
• Mordella heros Palm, 1823
• Mordella h-fasciata Lea, 1895
• Mordella hieroglyphica Fairmaire & Germain, 1863
• Mordella himalayana Horák, 1995
• Mordella hoanensis Píc, 1941
• Mordella hoberlandti Horák, 1985
• Mordella hofferi Horák, 1979
• Mordella holomelaena Apfelbeck, 1914
• Mordella holosericea Solier, 1851
• Mordella homochroa Fairmaire, 1895
• Mordella hoshihanamima Franciscolo, 1965
• Mordella hubbsi Liljeblad, 1922
• Mordella huetheri Ermisch, 1956
• Mordella humeralis Waterhouse, 1878
• Mordella humeropicta Ermisch, 1962
• Mordella ignaciosa Píc, 1936
• Mordella ignota Lea, 1895
• Mordella immaculata Smith, 1883
• Mordella incisa Lea, 1931
• Mordella incisura Lea, 1931
• Mordella inclusa (Germar, 1813)
• Mordella inconspicua Lea, 1895
• Mordella indata Statz, 1952
• Mordella infranotata Píc, 1937
• Mordella infrasignata Píc, 1941
• Mordella innotatopyga Píc, 1936
• Mordella inornata Lea, 1902
• Mordella inornatipennis Píc, 1948
• Mordella insidiosa Lucas, 1849
• Mordella insignata Píc, 1941
• Mordella insulata LeConte, 1859
• Mordella inusitata Blackburn, 1893
• Mordella invisitata Liljeblad, 1945
• Mordella iridea Lea, 1917
• Mordella israelsoni Plaza, 1977
• Mordella javana Píc, 1941
• Mordella kannegietri Píc, 1941
• Mordella kanoi Kônô, 1932
• Mordella kanpira Takakuwa, 1985
• Mordella knulli Liljeblad, 1922
• Mordella kreusei Philippi, 1864
• Mordella kuatunensis Ermisch, 1968
• Mordella lachrymosa Lea, 1931
• Mordella lacsonensis Píc, 1922
• Mordella lapidicola Wickham, 1909
• Mordella latefasciata Fairmaire, 1905
• Mordella latejuncta Píc, 1924
• Mordella latemaculata Ray, 1944
• Mordella latenotata Píc, 1923
• Mordella lateplagiata Fairmaire, 1895
• Mordella laterufescens Píc, 1936
• Mordella latesignata Píc, 1936
• Mordella laticornis Ray, 1946
• Mordella latipennis Píc, 1941
• Mordella latithorax Ray, 1939
• Mordella leai Csiki, 1915
• Mordella lebisi Píc, 1937
• Mordella legionensis Plaza & Compte, 1981
• Mordella lemoulti Píc, 1936
• Mordella leonensis Píc, 1932
• Mordella lepida Redtdenbacher, 1868
• Mordella leucaspis Küster, 1849
• Mordella leucocephla Quedenfeldt, 1886
• Mordella leucogramma Champion, 1891
• Mordella leucographa Champion, 1893
• Mordella leucopleura Lea, 1931
• Mordella leucospila Fairmaire, 1883
• Mordella leucosticta Germain, 1848
• Mordella leucostigma Fairmaire, 1863
• Mordella limbata Waterhouse, 1878
• Mordella lineatipyga Champion, 1891
• Mordella lineatonotata Ray, 1936
• Mordella longecaudata Fairmaire, 1891
• Mordella longehumeralis Píc, 1936
• Mordella longicauda Roubal, 1921
• Mordella longipalpis Ray, 1946
• Mordella lottini Boisduval, 1835
• Mordella luctuosa Solier, 1851
• Mordella lunulata Helmuth, 1865
• Mordella luteodispersa Píc, 1936
• Mordella luteoguttata Blanch, 1843
• Mordella luteomaculata Píc, 1928
• Mordella luteonotata Píc, 1916
• Mordella luteopyga Píc, 1929
• Mordella luteosignata Píc, 1941
• Mordella luteosuturalis Píc, 1936
• Mordella maceki (Horák, 1985)
• Mordella maculipennis Ray, 1936
• Mordella madagascariensis Csiki, 1915
• Mordella major Píc, 1936
• Mordella malleri Píc, 1936
• Mordella marginata Melsheimer, 1845
• Mordella marmorata Fabricius, 1801
• Mordella maroniensis Píc, 1924
• Mordella mastersi Lea, 1895
• Mordella maxima Píc, 1948
• Mordella mcnamarae Lea, 1931
• Mordella mediolineata Píc, 1936
• Mordella melaena Germar, 1824
• Mordella melanocephala Fairmaire & Germain, 1863
• Mordella melanozosta Fairmaire, 1906
• Mordella mendesensis Píc, 1936
• Mordella meridionalis Méquignon, 1946
• Mordella mesoleuca Lea, 1929
• Mordella metallica Champion, 1891
• Mordella metasternalis Lea, 1917
• Mordella mexicana Champion, 1891
• Mordella micacea Píc, 1941
• Mordella militaris Ray, 1939
• Mordella minor Píc, 1942
• Mordella minutissima Píc, 1942
• Mordella mixta Fabricius, 1801
• Mordella mongolica Ermisch, 1964
• Mordella moorei Perroud, 1864
• Mordella moscovia Roubal, 1921
• Mordella moscoviensis Roubal, 1921
• Mordella multiguttata Waterhouse, 1878
• Mordella nana Fairmaire & Germain, 1863
• Mordella nesiotica Ray, 1949
• Mordella niasensis Píc, 1941
• Mordella nigra Fairmaire & Germain, 1863
• Mordella nigrans McLeay, 1887
• Mordella nigricolor Píc, 1931
• Mordella nigricornis Shchegolvera-Barovskaya, 1931
• Mordella nigripes Píc, 1941
• Mordella nigroapicalis Píc, 1941
• Mordella nigroguttata Fairmaire, 1899
• Mordella nigromaculata Champion, 1891
• Mordella nigropilosa Statz, 1952
• Mordella nigroterminata Blair, 1922
• Mordella niveoscutellata Nakane & Nomura, 1950
• Mordella norfolcensis Lea, 1917
• Mordella notatipennis Lea, 1917
• Mordella novemguttata Montrouzier, 1855
• Mordella novemmaculata Lea, 1902
• Mordella novemnotata Ray, 1944
• Mordella obliqua LeConte, 1878
• Mordella obliquirufa Lea, 1917
• Mordella obscuripennis McLeay, 1887
• Mordella octodecimmaculata Lea, 1895
• Mordella octoguttata Montrouzier, 1855
• Mordella octolineata Champion, 1891
• Mordella octonotata Píc, 1936
• Mordella ogloblini Píc, 1930
• Mordella ornata Waterhouse, 1878
• Mordella ornatopallida Reitt, 1911
• Mordella ovalistica McLeay, 1887
• Mordella pagdeni Píc, 1935
• Mordella palembanga Píc, 1941
• Mordella pallida Lea, 1895
• Mordella palmae Emery, 1876
• Mordella palmai Méquignon, 1946
• Mordella panamensis Champion, 1891
• Mordella parva Champion, 1895
• Mordella pauli Píc, 1924
• Mordella pauper Maeklin, 1875
• Mordella peregrinator Champion, 1917
• Mordella phungi Píc, 1923
• Mordella picta Chevrolat, 1829
• Mordella plagiata Mannh, 1849
• Mordella plaumanni Píc, 1936
• Mordella pleurosticta Lucas, 1859
• Mordella plurinotata Blanch, 1853
• Mordella poeciloptera Lea, 1929
• Mordella pondolandiae Franciscolo, 1965
• Mordella postimpressa Píc, 1923
• Mordella postinsignata Píc, 1937
• Mordella pragensis Roubal, 1921
• Mordella pretiosa Champion, 1891
• Mordella priscula Cockerell, 1924
• Mordella promiscua Ermisch, 1942
• Mordella proxima Solier, 1851
• Mordella pseudobrachyura Franciscolo, 1949
• Mordella pubescens Liljeblad, 1921
• Mordella pulchra Lea, 1895
• Mordella pulchripennis Píc, 1936
• Mordella pulverulenta McLeay, 1887
• Mordella punctulata Maeklin, 1875
• Mordella purpurascens Apfelbeck, 1914
• Mordella pustulosa Champion, 1891
• Mordella pygidialis Apfelbeck, 1914
• Mordella pygmaea Champion, 1895
• Mordella quadriguttulata Motschulsky, 1872
• Mordella quadrimaculata Lea, 1917
• Mordella quadrioculata Fairmaire, 1895
• Mordella quadripunctata (Say, 1824)
• Mordella quadripustulata Maeklin, 1875
• Mordella quadrisignata Chevrolat, 1834
• Mordella quatuordecimmaculata McLeay, 1873
• Mordella quelpaertensis Píc, 1941
• Mordella quomoi Franciscolo, 1942
• Mordella ragneri Ermisch
• Mordella reductesignata Píc, 1936
• Mordella reticulata Fairmaire, 1906
• Mordella reynoldsi Ray, 1930
• Mordella robusta Píc, 1936
• Mordella roeri (Batten, 1981)
• Mordella rouyeri Píc, 1920
• Mordella rudebecki Franciscolo, 1965
• Mordella ruficauda Maeklin, 1875
• Mordella ruficollis Waterhouse, 1878
• Mordella ruficornis Shchegolvera-Barovskaya, 1931
• Mordella rufipennis Solier, 1851
• Mordella rufipes Lea, 1895
• Mordella rufoapicalis Píc, 1923
• Mordella rufoaxillaris Fairmaire & Germain, 1860
• Mordella rufocinerea Ray, 1944
• Mordella rufomaculata Champion, 1891
• Mordella rufopectoralis Píc, 1936
• Mordella rufopyga Píc, 1936
• Mordella rufosutralis Píc, 1917
• Mordella rufovittata Champion, 1891
• Mordella rutilicollis Champion, 1891
• Mordella salomonensis Píc, 1929
• Mordella sasajii Takakuwa, 2001
• Mordella savioi Píc, 1924
• Mordella schapleri Píc, 1936
• Mordella scheelei Ermisch, 1941
• Mordella schwarzi Liljeblad, 1945
• Mordella scita Maeklin, 1875
• Mordella scutellaris Fabricius, 1801
• Mordella sellata Champion, 1891
• Mordella semiluctuosa Fairmaire, 1902
• Mordella septemnotata Champion, 1891
• Mordella seriata Champion, 1891
• Mordella sericans Lea, 1917
• Mordella setacea Montrouzier, 1855
• Mordella sexdecimguttata Montrouzier, 1860
• Mordella sexguttata Champion, 1896
• Mordella sexnotata Champion, 1891
• Mordella shirozui Nomura, 1967
• Mordella sicardi Píc, 1942
• Mordella signata Champion, 1891
• Mordella simillina Lea, 1902
• Mordella sinensis Píc, 1917
• Mordella sinuata Fairmaire, 1897
• Mordella sororcula Berg, 1889
• Mordella splendens Miwa, 1933
• Mordella splendidula Falderm, 1837
• Mordella stenopyga Ermisch, 1965
• Mordella sticticoptera Champion, 1891
• Mordella strigipennis Falderm, 1837
• Mordella stygia Wickham, 1914
• Mordella subaenea Ray, 1944
• Mordella subapicalis Píc, 1936
• Mordella subauratonotata Píc, 1924
• Mordella subbasalis Shchegolvera-Barovskaya, 1931
• Mordella subdola Champion, 1891
• Mordella subfasciata Maeklin, 1875
• Mordella submaculata Ray, 1936
• Mordella subnotata Maeklin, 1875
• Mordella subobliquefasciata Píc, 1936
• Mordella subuniformis Píc, 1937
• Mordella subvittata McLeay, 1887
• Mordella sultiatronotata Píc, 1917
• Mordella sumatrana Píc
• Mordella summermanae Ray, 1939
• Mordella suturalis Fairmaire & Germain, 1863
• Mordella suturella Fairmaire, 1897
• Mordella sydneyana Blackburn, 1893
• Mordella tabulae Franciscolo, 1965
• Mordella tachyporiformis Curtis, 1845
• Mordella tadjikistanica Odnosum, 2002
• Mordella tairuensis Broun, 1880
• Mordella taiwana Nakane & Nomura, 1950
• Mordella t-album Champion, 1891
• Mordella tantilla Champion, 1891
• Mordella tarsalis Lea, 1917
• Mordella teapensis Champion, 1891
• Mordella teitteri Píc, 1936
• Mordella tenella Maeklin, 1875
• Mordella tenuipes Champion, 1891
• Mordella testaceiceps Píc, 1927
• Mordella testaceipes Píc, 1936
• Mordella testaceohumeralis Píc, 1936
• Mordella tetragramma Fairmaire, 1906
• Mordella tetraleucosticta Lea, 1931
• Mordella tetraspilota Champion, 1891
• Mordella tetrastictoptera Lea, 1929
• Mordella theresae Píc, 1948
• Mordella thoracica Solier, 1851
• Mordella tijucaensis Píc, 1936
• Mordella tomentosa Boisduval, 1835
• Mordella tonkinea Píc, 1916
• Mordella tonzalini Píc, 1941
• Mordella triangulifer Champion, 1891
• Mordella trifasciata Píc, 1936
• Mordella trilineata Píc, 1927
• Mordella triloba Say, 1824
• Mordella trilobibasa Lea, 1929
• Mordella trinotata Píc, 1941
• Mordella tripartita Champion, 1891
• Mordella tristicula Maeklin, 1875
• Mordella tristis Lea, 1895
• Mordella tunisica Ermisch, 1969
• Mordella turneri Franciscolo, 1965
• Mordella undosa McLeay, 1887
• Mordella undulata Melsheimer, 1846
• Mordella unifasciata Píc, 1952
• Mordella uniformis Lea, 1895
• Mordella unilineata Píc, 1936
• Mordella uninotata Píc, 1942
• Mordella unisignata Píc, 1942
• Mordella univestis Ermisch, 1965
• Mordella univittata Champion, 1891
• Mordella vadoni Píc, 1937
• Mordella varicornis Champion, 1891
• Mordella varienotata Píc, 1941
• Mordella variesignata Píc, 1936
• Mordella v-aureum Lea, 1902
• Mordella veitchi Blair, 1922
• Mordella velutina Emery, 1876
• Mordella verdensis Píc, 1936
• Mordella verticordiae Lea, 1902
• Mordella vesconis Solier, 1851
• Mordella vestita Emery, 1876
• Mordella v-fasciata Lea, 1895
• Mordella vidua Solier, 1851
• Mordella villiersi Píc, 1950
• Mordella violacescens Phillipi, 1864
• Mordella viridepennis Mulsant, 1856
• Mordella viridescens Costa, 1854
• Mordella viridis Píc, 1930
• Mordella vitiensis Blair, 1922
• Mordella vittacollis Lea, 1917
• Mordella vittata Fabricius, 1801
• Mordella vittaticollis Píc, 1936
• Mordella waterhousei Champion, 1895
• Mordella waterstradi Píc, 1941
• Mordella weisei Schilsky, 1895
• Mordella weiseri Píc, 1930
• Mordella wiburdi Lea, 1895
• Mordella woodi Píc, 1930
• Mordella xanthogastra Fairm & Germain, 1863
• Mordella xanthosticta Champion, 1891
• Mordella yami Nomura, 1967

1. ↑ Linnaeus, C. 1758: Systema Naturae per regna tria naturæ, secundum classes, ordines, genera, species, cum characteribus, differentiis, synonymis, locis, Tomus I. Editio decima, reformata. Holmiæ: impensis direct. Laurentii Salvii. i–ii, 1–824 pp doi: 10.5962/bhl.title.542: page 420