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The Varunidae are a family of thoracotrematan crabs. The delimitation of this family, part of the taxonomically confusing Grapsoidea, is undergoing revision. For a long time, they were placed at the rank of subfamily in the Grapsidae, but they appear to be closest to Macropthalmus and the Mictyridae, which are usually placed in the Ocypodoidea. It may thus be better to merge the latter superfamily with the Grapsoidea, retaining the latter name as it is older.[2]
That notwithstanding, the revision of the Grapsoidea (in the narrow but apparently still paraphyletic sense) is also not fully completed, as many taxa remain to be restudied. Already, several former Grapsidae genera have been moved to the Varunidae, and others are tentatively placed here pending detailed study. Among the most noteworthy of these is the Chinese mitten crab, Eriocheir sinensis.[2] The genus Xenograpsus, formerly included in the Varunidae, is now placed in its own family, the Xenograpsidae.[1]
The following genera are included:[3]
The Varunidae are a family of thoracotrematan crabs. The delimitation of this family, part of the taxonomically confusing Grapsoidea, is undergoing revision. For a long time, they were placed at the rank of subfamily in the Grapsidae, but they appear to be closest to Macropthalmus and the Mictyridae, which are usually placed in the Ocypodoidea. It may thus be better to merge the latter superfamily with the Grapsoidea, retaining the latter name as it is older.
That notwithstanding, the revision of the Grapsoidea (in the narrow but apparently still paraphyletic sense) is also not fully completed, as many taxa remain to be restudied. Already, several former Grapsidae genera have been moved to the Varunidae, and others are tentatively placed here pending detailed study. Among the most noteworthy of these is the Chinese mitten crab, Eriocheir sinensis. The genus Xenograpsus, formerly included in the Varunidae, is now placed in its own family, the Xenograpsidae.
The following genera are included:
Asthenognathinae Stimpson, 1858 Asthenognathus Stimpson, 1858 Globihexapus † Schweitzer & Feldmann, 2001 Cyclograpsinae H. Milne-Edwards, 1853 Austrohelice K. Sakai, Türkay & Yang, 2006 Chasmagnathus De Haan, 1833 Cyclograpsus H. Milne-Edwards, 1837 Helicana K. Sakai & Yatsuzuka, 1980 Helice De Haan, 1833 Helograpsus Campbell & Griffin, 1966 Metaplax H. Milne-Edwards, 1852 Miosesarma † Karasawa, 1989 Neohelice K. Sakai, Türkay & Yang, 2006 Paragrapsus H. Milne-Edwards, 1853 Parahelice K. Sakai, Türkay & Yang, 2006 Pseudohelice K. Sakai, Türkay & Yang, 2006 Gaeticinae Davie & N. K. Ng, 2007 Gaetice Gistel, 1848 Gopkittisak Naruse & P. F. Clark, 2009 Sestrostoma Davie & N. K. Ng, 2007 Thalassograpsinae Davie & N. K. Ng, 2007 Thalassograpsus Tweedie, 1950 Varuninae H. Milne-Edwards, 1853 Acmaeopleura Stimpson, 1858 Brachynotus De Haan, 1833 Cyrtograpsus Dana, 1851 Eriocheir De Haan, 1835 Grapsodius Holmes, 1900 Hemigrapsus Dana, 1851 Neoeriocheir T. Sakai, 1983 Noarograpsus N. K. Ng, Manuel & Ng, 2006 Orcovita Ng & Tomascik, 1994 Otognathon Ng & Stevcic, 1993 Parapyxidognathus Ward, 1941 Platyeriocheir N. K. Ng, J. Guo & Ng, 1999 Pseudogaetice Davie & N. K. Ng, 2007 Pseudograpsus H. Milne-Edwards, 1837 Ptychognathus Stimpson, 1858b Pyxidognathus A. Milne-Edwards, 1879 Scutumara Ng & Nakasone, 1993 Tetragrapsus Rathbun, 1918 Utica White, 1847 Varuna Milne-Edwards, 1830