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Figures 1-6. Daphniola eptalophos sp n. photographed in ethanol. Apical view 1, alive specimen carrying egg capsules with an embryo on last body whorl and inside umbilicus (photographed in water) 2 operculum 3 soft body, head with tentacles and penis in situ 4-5 penis 6. A background square represents 1 mm2 in Figs 1, 4, 5. Scale bar 1 mm and 0.5 mm in Figs 2 and 3 respectively. Black arrow points the penis in Figs 4-5 and the outgrowth of penis in Fig. 6.
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Peter Glöer, Vladimir Pešić
Zookeys
Figure 9.Pseudamnicola georgievi sp. n.: shell.
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Vladimir Pešić, Peter Glöer
Zookeys
Figure 2.Karucia sublacustrina sp. n. a–b shells (a holotype b paratype) c head with penis in situ (arrow showing outgrowth on the left side of penis) d penis. Abbreviations: e eye, p penis, s snout, t tentacle.
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Robert Hershler, Hsiu-Ping Liu, Corbin Bradford
Zookeys
Figure 3.Shells, opercula and radula, Pyrgulopsis licina sp. n. A Holotype, USNM 850347 B, C Shells, USNM 1204732, USNM 1188732 D, E Opercula (outer, inner sides), USNM 850348 F Portion of radular ribbon, USNM 850348 G Central teeth, USNM 850348 H Lateral and inner marginal teeth, USNM 850348. Scale bars A–C 1.0 mm; D, E 100 µm; F, H 10 µm; G 2 µm.
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Robert Hershler, Hsiu-Ping Liu, Corbin Bradford
Zookeys
Figure 4.Penes (dorsal, ventral surfaces). A, B Pyrgulopsis licina sp. n., USNM 850346 C, D Pyrgulopsis perforata sp. n., BellMNH 20891 E, F Pyrgulopsis sanchezi sp. n., USNM 883361 G, H Pyrgulopsis micrococcus, BellMNH 20663 I, J Pyrgulopsis turbatrix, USNM 860699 K, L Pyrgulopsis turbatrix, USNM 883373. Scale bars A–C 250 µm; D–L 500 µm. Pd penial duct Pf penial filament Pl penial lobe Tg terminal gland.
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Canella Radea, Aristeidis Parmakelis, Vassilis Papadogiannis, Despoina Charou, Kostas A. Triantis
Zookeys
Figure 3.Hydrobioids collectedduring the survey in mainland and insular Greece. A Daphniola exigua (dorsal and ventral view) B Isimerope semele (dorsal and ventral view, Megali Vrysi) C Pseudoislamia balcanica (dorsal and ventral view, Ag. Sophia) D Pseudamnicola pieperi (Olympos) E Radomaniola cf. curta (spring of Louros river) F Radomaniola cf. curta (Ag. Sophia spring) G Trichonia trichonica H Pseudobithynia eubooensis. Scale bar 1 mm.
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Abraham S.H. Breure, Jonathan D. Ablett
Zookeys
Figure 25.Drymaeus species. A Drymaeus (Mesembrinus) aurifluus (Pfeiffer, 1857), lectotype NHMUK 1975502 (H = 21.8) B Drymaeus (Mesembrinus) bugabensis (Martens, 1893), lectotype NHMUK 1901.6.22.958 (H = 26.3) C Drymaeus (Mesembrinus) dubius (Pfeiffer, 1853), lectotype NHMUK 1975519 (H = 27.1) D Drymaeus (Mesembrinus) (?) rufescens (J.E. Gray, 1825), lectotype of Bulimus erusbescens Pfeiffer, 1847 NHMUK 1975562 (H = 24.2) E Drymaeus (Mesembrinus) conicus da Costa, 1907, lectotype NHMUK 1907.11.21.32 (H = 17.5) F Drymaeus (Mesembrinus) granadensis (Pfeiffer, 1848), lectotype of Bulimus demotus Reeve, 1850 NHMUK 1975504 (H = 31.8) G Drymaeus (Mesembrinus) vexillum (Wood, 1828), lectotype of Bulimus keppelli Pfeiffer, 1853 NHMUK 1975538 (H = 33.2) H Drymaeus (Mesembrinus) attenuatus varicosus (Pfeiffer, 1853), lectotype NHMUK 1975466 (H = 35.5) I–K Drymaeus (Drymaeus) signifer (Pfeiffer, 1855), lectotype NHMUK 1975216 (H = 32.9). All enlarged.
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Robert Hershler, Victoria Ratcliffe, Hsiu-Ping Liu, Brian Lang, Claire Hay
Zookeys
Figure 3.Shells, opercula and radula, Pyrgulopsis marilynae n. sp. A Holotype, USNM 1135068 B Shell, USNM 1135067 C, D Opercula (outer, inner sides), USNM 1135067 E Portion of radular ribbon, USNM 1135067 F Central teeth, USNM 1135067 G Lateral and inner marginal teeth, USNM 1135067. Scale bars A, B 1.0 mm; C, D 100 µm; E 20 µm, F, G 10 µm.
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Figures 7-10. Daphniola eptalophos sp. n. shell images from SEM Shell habitus 7, 8 protoconch 9 teleoconch 10 Scale bar 0.5 mm in Figs 7, 9 and 0.05 mm in Figs 8,10.
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Peter Glöer, Vladimir Pešić
Zookeys
Figure 10.Kaskakia khorrasanensis sp. n. a shell b penis in situ c–d penis (c: dorsal view, d: ventral view).
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Robert Hershler, Hsiu-Ping Liu, Corbin Bradford
Zookeys
Figure 4.Penes (dorsal, ventral surfaces). A, B Pyrgulopsis licina sp. n., USNM 850346 C, D Pyrgulopsis perforata sp. n., BellMNH 20891 E, F Pyrgulopsis sanchezi sp. n., USNM 883361 G, H Pyrgulopsis micrococcus, BellMNH 20663 I, J Pyrgulopsis turbatrix, USNM 860699 K, L Pyrgulopsis turbatrix, USNM 883373. Scale bars A–C 250 µm; D–L 500 µm. Pd penial duct Pf penial filament Pl penial lobe Tg terminal gland.
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Robert Hershler, Hsiu-Ping Liu, Corbin Bradford
Zookeys
Figure 9.Shells, opercula and radula, Pyrgulopsis turbatrix. A Paratype, USNM 860699. B–D Shells, SBMNH uncat., USNM 853508, USNM 853510 E, F Opercula (outer, inner sides), USNM 860699 G Portion of radular ribbon, USNM 860699 H Central teeth, USNM 860699 I Lateral and outer marginal teeth, USNM 860699. Scale bars A–D 1.0 mm; E, F 250 µm; G 20 µm; H, I 10 µm.
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Canella Radea, Aristeidis Parmakelis, Vassilis Papadogiannis, Despoina Charou, Kostas A. Triantis
Zookeys
Figure 3.Hydrobioids collectedduring the survey in mainland and insular Greece. A Daphniola exigua (dorsal and ventral view) B Isimerope semele (dorsal and ventral view, Megali Vrysi) C Pseudoislamia balcanica (dorsal and ventral view, Ag. Sophia) D Pseudamnicola pieperi (Olympos) E Radomaniola cf. curta (spring of Louros river) F Radomaniola cf. curta (Ag. Sophia spring) G Trichonia trichonica H Pseudobithynia eubooensis. Scale bar 1 mm.
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Abraham S.H. Breure, Jonathan D. Ablett
Zookeys
Figure L18.Labels. i Bulimus deshayesi Pfeiffer, 1845 ii Bulimus devians Dohrn, 1863 iii Bulinus discrepans Sowerby I, 1833 iv Bulimus dubius Pfeiffer, 1853 v Bulimus dombeyanus ‘Férussac’ Pfeiffer, 1846.
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Robert Hershler, Victoria Ratcliffe, Hsiu-Ping Liu, Brian Lang, Claire Hay
Zookeys
Figure 4.Penes (dorsal, ventral surfaces). A, B Pyrgulopsis marilynae n. sp., USNM 883175 C, D Pyrgulopsis similis n. sp., USNM 1135065 E, F Pyrgulopsis gilae, BellMNH 20898. Scale bars A–F 200 µm. Dg2 dorsal gland along left edge Dg3 dorsal gland along right distal edge Ipg inner (left) penial gland Opg outer (right) penial gland Tg terminal gland Vg ventral gland.
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Figures 11-13.Female genitalia of Daphniola species.Daphniola eptalophos sp. n. female genitalia drawn from the only one female individual found among dissected specimens 11 Daphniola louisifemale genitalia re-drawn from Falniowski and Szarowska (2000) 12 Daphniola exiguafemale genitalia re-drawn from Radoman (1983) 13 Scale bar 0.5 mm.
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Peter Glöer, Vladimir Pešić
Zookeys
Figure 11.Sarkhia sarabensis nov. sp.a shell b, c penis in situ.
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Robert Hershler, Hsiu-Ping Liu, Corbin Bradford
Zookeys
Figure 2.Map showing the distribution of mtDNA clades A–F with color codes matching those in Fig. 1.
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Canella Radea, Aristeidis Parmakelis, Vassilis Papadogiannis, Despoina Charou, Kostas A. Triantis
Zookeys
Figure 1.Map showing the administrative units of Greece where “hydrobioid” localities were sampled during the fieldwork of this study.
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Robert Hershler, Victoria Ratcliffe, Hsiu-Ping Liu, Brian Lang, Claire Hay
Zookeys
Figure 1.Map showing the distribution of mtDNA clades I-III.
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Figures 14-16. Shells of Daphniola species. a, apical view, b, ventral view, c. frontal view Daphniola eptalophos sp. n. (Agoriani) 14 Daphniola exigua (Marathonas, Attica) 15 Daphniola louisi (Kessariani, Attica) 16 A background square represents 1 mm2.
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Robert Hershler, Hsiu-Ping Liu, Corbin Bradford
Zookeys
Figure 5.Photographs of habitats. A Spring south of Clay Pits, Ash Meadows, Nye County, Nevada, type locality of Pyrgulopsis licina sp. n. (photograph taken on 7/VII/1986) B Purgatory Spring, Ash Meadows, Nye County, Nevada, type locality of Pyrgulopsis sanchezi (15/XI/2011) C, D Uppermost spring east of Scotty’s Castle, Death Valley, Inyo County, California, type locality of Pyrgulopsis perforata sp. n. (18/IV/1980).
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Canella Radea, Aristeidis Parmakelis, Vassilis Papadogiannis, Despoina Charou, Kostas A. Triantis
Zookeys
Figure 3.Hydrobioids collectedduring the survey in mainland and insular Greece. A Daphniola exigua (dorsal and ventral view) B Isimerope semele (dorsal and ventral view, Megali Vrysi) C Pseudoislamia balcanica (dorsal and ventral view, Ag. Sophia) D Pseudamnicola pieperi (Olympos) E Radomaniola cf. curta (spring of Louros river) F Radomaniola cf. curta (Ag. Sophia spring) G Trichonia trichonica H Pseudobithynia eubooensis. Scale bar 1 mm.
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Robert Hershler, Victoria Ratcliffe, Hsiu-Ping Liu, Brian Lang, Claire Hay
Zookeys
Figure 2.Bayesian tree based on the combined (COI, NDI) dataset. Posterior probabilities for nodes are given when >95%. Specimen codes are from the Table 1.