Scarabaeiformia

Figures 1–4.Polyphylla (Granida) simoni sp. n. 1 Habitus of holotype male (length 22.0 mm), dorsal view 2 Detail of pronotum, dorsal view 3 Male genitalia, dorsal view 4 The same, right lateral view, shaded area indicates overlapping part of left paramere. Scale bar: 5 mm for Figs 3–4.

Figures 4–7. Characters of the left hindleg of Parochodaeus species, ventral view 4 Parochodaeus dentipes with arrow indicating acute tooth near apex of metafemur; metatibia broad 5 Parochodaeus jujuyus, showing greatly enlarged first metatarsomere 6 Parochodaeus procelipes, narrow metatibia 7 Parochodaeus stupendus, with large metatrochanteral tooth, large ventral tooth at the metafemoral apex (second, dorsal tooth obscured), and strongly curved metatibia.

Figure 1.Cretohypna cristata Yan, Nikolajev & Ren gen. et sp. n., holotype, registration No. CNU-COL-NN2011003, a protibia b mesotibia c body in dorsal view d metatibia and metatarsus.

Figures 1–8.Oblique and lateral habitus views of Bolboceras: 1–4 Bolboceras inaequale (1–2 major specimen from Kolkata, length 11 mm 3–4 minor specimen from Dhanjuri, length 9 mm) 5–6 duplicatum, holotype 7–8 orissicum, holotype. Scale lines 1 mm.

Figures 1–12.Peltonotus species dorsal habitus. 1 Peltonotus animus, male 2–3 Peltonotus cybele, male and female (respectively) 4 Peltonotus favonius, male 5–7 Peltonotus fujiokai, males (showing variation) 8 Peltonotus fujiokai, female 9 Peltonotus karubei, male 10–11 Peltonotus malayensis, male and female (respectively) 12 Peltonotus morio, male.

Figures 1–10.1–3 Habitus of males 4 punctation of elytral disc 5 aedeagus in lateral view and parameres in dorsal view 6 serration of medial margins of parameres 7–10 aedeagus in ventral view 1, 4–7 Stenosternus costatus Karsch, holotype 2 Aegidium colombianum Westwood 3 Orphnus macleayi Laporte de Castelnau 8 Hybalus cornifrons (Brullé) 9 Aegidium parvulum Westwood 10 Orphnus compactus Petrovitz.

Figure 1.Gyronotus perissinottoi sp. n. A Holotype male, dorsal habitus B Allotype female, dorsal habitus C Male pygidium, ventral side D Female pygidium, ventral side E left F right and G ventral sides of aedeagus. Photo: Mickaël François.

Figure 3.A–D Neoserica bansongchana Ahrens, Liu & Fabrizi sp. n. (holotype), E–H Neoserica septemfoliata Moser (lectotype) I–L Neoserica gaoligongshanica Ahrens, Liu & Fabrizi sp. n. (holotype) A, E, I Aedeagus, left side lateral view C, G, K Aedeagus, right side lateral view B, F, J parameres, dorsal view D, H, L Habitus. Scale: 0.5 mm.

Figures 1–4.Syntypes of Onthophagus cervicornis Kirby, 1825. 1 Dorsal habitus of the male 2 Dorsal habitus of the female 3 Labels of the male (Kirby handwriting) 4 Labels of the female.

Figure 1.Eriopeltastes (Eriopeltastes) ornatus Ricchiardi, sp. n. Holotype ♂: a habitus in dorsal view b habitus in ventral view c parameres in frontal view d parameres in lateral view (Photo & drawing: Enrico Ricchiardi).

Figures 1–4.Primary types of Apterocyclus honoluluensis and its synonyms, with labels. 1 Apterocyclus honoluluensis Waterhouse, lectotype 2 Apterocyclus deceptor Sharp, holotype 3 Apterocyclus feminalis Sharp, holotype 4 Apterocyclus varians Sharp, holotype.

Figures 1–5.Morphological features of Parachorius semsanganus sp. n.: 1–3 paratype, aedeagus 1 aedeagus lateral view 2 aedeagus apical view 3 aedeagus dorsal view 4 male holotype, habitus 5 male paratype, hind leg, arrow indicates teeth on inner tibial margin.

Figures 11–13. Dorsal habitus of Parochodaeus species, males 11 Parochodaeus jujuyus, sp. n.12 Parochodaeus perplexus, sp. n.13 Parochodaeus phoxus, sp. n.

Figure 2.Cretohypna cristata Yan, Nikolajev & Ren gen. et sp. n., line drawings of holotype a dorsal view b ventral view.

Figures 9–15.Forebody and aedeagus of Bolboceras: 9–12 head and pronotum, frontodorsal view 9–10 Bolboceras inaequale (9 major specimen from Kolkata 10 minor specimen from Dhanjuri) 11 duplicatum,holotype 12 orissicum,holotype 13–15 aedeagus, upper side view, of 13 Bolboceras inaequale,major specimen from Kolkata 14 orissicum,holotype 15 duplicatum,holotype. Scale lines 1 mm.

Figures 61–66.Male parameres (with or without phallobase), dorsal and lateral views, in Peltonotus. Male parameres are highly asymmetrical, and we illustrate the lateral view that best assists in identification. 61 Peltonotus fujiokai 62 Peltonotus gracilipodus and Peltonotus podocrassus 63 Peltonotus karubei 64 Peltonotus malayensis 65 Peltonotus morio 66 Peltonotus nasutus.

Figures 11—37.11, 12 Right mandible in dorsal and apical view 13 apex of metafemur 14 head in ventral view 15–23 protibia of males (arrows in figures 15–17 indicate approximate teeth direction) 24–33 metepisternum 34–37 base of prothorax (arrows in figures 34–36 indicate keel separating basal and anterolateral parts of propleurae) 11, 16, 35 Aegidium colombianum Westwood 12, 37 Orphnus declivis Schmidt 13–15, 24, 34 Stenosternus costatus Karsch, holotype 17, 29 Orphnus macleayi Laporte de Castelnau 18, 36 Aegidinus guianensis (Westwood) 19, 27 Aegidiellus alatus (Laporte de Castelnau) 20, 26 Paraegidium costalimai Vulcano et al. 21, 31 Pseudorphnus hiboni Paulian 22, 30 Triodontus itremoi Paulian 23 Chaetonyx robustus Schaum 25 Aegidium parvulum Westwood 28 Orphnus giganteus Paulian 32 Renorphnus clementi (Petrovitz) 33 Hybalus cornifrons (Brullé).

Figure 3.Gyronotus perissinottoi sp. n. in its natural habitat at the Umthamvuna Nature Reserve. Photo: Lynette Clennell.

Figure 7.Distribution of the species of the Neoserica (s.l.) septemlamellata group: Neoserica bansongchana, Neoserica gaoligongshanica, Neoserica guangpingensis, Neoserica plurilamellata, Neoserica sapaensis, Neoserica takakuwai, Neoserica yanzigouensis."?" stands for the not localised records of Neoserica septemfoliata from Yunnan.

Figure 2.Eriopeltastes (Eriopeltastes) ornatus Ricchiardi, sp. n. Paratype ♀: a habitus in dorsal view b habitus in ventral view (Photo: Enrico Ricchiardi).

Figures 12–21.Left protibia and mandibles of Apterocyclus species. 12, 17 Apterocyclus honoluluensis 13, 18 Apterocyclus kawaii 14, 19 Apterocyclus munroi 15, 20 Apterocyclus palmatus 16, 21 Apterocyclus waterhousei.

Map 2. Argentinean distribution of Parochodaeus jujuyus (white circles), Parochodaeus perplexus (orange circles), Parochodaeus cornutus (blue squares), and Parochodaeus dentipes (yellow triangle).

Figure 1.Cretohypna cristata Yan, Nikolajev & Ren gen. et sp. n., holotype, registration No. CNU-COL-NN2011003, a protibia b mesotibia c body in dorsal view d metatibia and metatarsus.

Figures 16–23.Head, full-face view, and pronotum, dorsal view, of Bolboceras: 16–19 Bolboceras inaequale (16–17 major specimen from Kolkata 18–19 minor specimen from Dhanjuri) 20–21 duplicatum,holotype 22–23 orissicum,holotype. Scale lines 1 mm.