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Lifespan, longevity, and ageing

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Maximum longevity: 30 years (wild) Observations: One study in the wild estimated age for 18 individuals with the oldest estimated to be around 30 years (Stacey and Arnold 1999).
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Joao Pedro de Magalhaes
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de Magalhaes, J. P.
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Associations

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Other than humans, there are no known natural predators of Irrawaddy dolphins. Humans are responsible for a large number of deaths because Irrawaddy dolphins are often caught in nets or harmed through destructive fishing practices (e.g., dynamite fishing). Otherwise, they are typically considered the top predator in their river ecosystems.

Known Predators:

  • humans (Homo sapiens)
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Koss, M.; L. Mahan and S. Merrill 2012. "Orcaella brevirostris" (On-line), Animal Diversity Web. Accessed April 27, 2013 at http://animaldiversity.ummz.umich.edu/site/accounts/information/Orcaella_brevirostris.html
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Melissa Koss, San Diego Mesa College
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Lucretia Mahan, San Diego Mesa College
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Sam Merrill, San Diego Mesa College
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Paul Detwiler, San Diego Mesa College
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Tanya Dewey, University of Michigan-Ann Arbor
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Morphology

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Unlike other dolphins, Irrawaddy dolphins lack a beak and have flexible necks. The flexibility in the neck causes visible creases behind the head. Additionally, Irrawaddy dolphins have bulging heads, with the forehead extending past the mouth, broad triangular, paddle-like, pectoral fins, and small, triangular dorsal fins set approximately two-thirds of the body length along the back. Skin coloration varies from slate-blue to slate-gray with a lighter underside. The face and head are similar in appearance to beluga whales (Delphinapterus leucas). Irrawaddy dolphins have narrow, pointed, peg-like teeth about 1 cm in length in both the upper and lower jaws. Body mass varies from 114 kg to 143 kg and the length ranges from 146 cm to 275 cm. Males tend to be larger in mass and length, with a larger dorsal fin. Irrawaddy dolphins lack a cardiac sphincter and the stomach is subdivided into compartments.

Range mass: 114 to 133 kg.

Average mass: 124 kg.

Range length: 146 to 275 cm.

Average length: 210 cm.

Other Physical Features: endothermic ; homoiothermic; bilateral symmetry

Sexual Dimorphism: male larger

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Koss, M.; L. Mahan and S. Merrill 2012. "Orcaella brevirostris" (On-line), Animal Diversity Web. Accessed April 27, 2013 at http://animaldiversity.ummz.umich.edu/site/accounts/information/Orcaella_brevirostris.html
author
Melissa Koss, San Diego Mesa College
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Lucretia Mahan, San Diego Mesa College
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Sam Merrill, San Diego Mesa College
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Paul Detwiler, San Diego Mesa College
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Tanya Dewey, University of Michigan-Ann Arbor
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Life Expectancy

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The oldest recorded Irrawaddy dolphins, all of which were found dead in fishing nets, were estimated to be 28 years old. Several individuals of this age have been found, all entangled in nets, so it is believed that Irrawaddy dolphins can live longer. Due to the small population size of Irrawady dolphins and subsequent difficulty in tracking the species, no further information is known regarding average lifespan.

Range lifespan
Status: wild:
28 to 30 years.

Average lifespan
Status: wild:
32 years.

Typical lifespan
Status: wild:
5 to 32 years.

Average lifespan
Status: wild:
16 years.

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Koss, M.; L. Mahan and S. Merrill 2012. "Orcaella brevirostris" (On-line), Animal Diversity Web. Accessed April 27, 2013 at http://animaldiversity.ummz.umich.edu/site/accounts/information/Orcaella_brevirostris.html
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Melissa Koss, San Diego Mesa College
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Lucretia Mahan, San Diego Mesa College
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Sam Merrill, San Diego Mesa College
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Paul Detwiler, San Diego Mesa College
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Tanya Dewey, University of Michigan-Ann Arbor
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Habitat

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Irrawaddy dolphins prefer coastal areas, particularly muddy, brackish waters at river mouths and deltas, and do not appear to venture far offshore. Most sightings have been made within 1.6 km of the coastline, but some have been reported in waters greater than 5 km from shore. Some populations are apparently restricted to fresh water, e.g. Chilka Lake, India, and Songhkla, Thailand.

Range depth: 2.5 to 18.0 m.

Habitat Regions: tropical ; saltwater or marine ; freshwater

Aquatic Biomes: rivers and streams; coastal ; brackish water

Other Habitat Features: riparian ; estuarine

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Koss, M.; L. Mahan and S. Merrill 2012. "Orcaella brevirostris" (On-line), Animal Diversity Web. Accessed April 27, 2013 at http://animaldiversity.ummz.umich.edu/site/accounts/information/Orcaella_brevirostris.html
author
Melissa Koss, San Diego Mesa College
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Lucretia Mahan, San Diego Mesa College
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Sam Merrill, San Diego Mesa College
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Paul Detwiler, San Diego Mesa College
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Tanya Dewey, University of Michigan-Ann Arbor
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Distribution

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Irrawaddy dolphins (Orcaella brevirostris) are found throughout much of the tropical and subtropical Indo-Pacific region; along the coasts of India, Bangladesh, Myanmar (Burma), Thailand, Cambodia (Kampuchea), Lao Peoples' Democratic Republic (Laos), Socialist Republic of Vietnam, Malaysia, Singapore, Brunei Darrusalam, and Indonesia.

Biogeographic Regions: oriental (Native ); indian ocean (Native ); pacific ocean (Native )

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Koss, M.; L. Mahan and S. Merrill 2012. "Orcaella brevirostris" (On-line), Animal Diversity Web. Accessed April 27, 2013 at http://animaldiversity.ummz.umich.edu/site/accounts/information/Orcaella_brevirostris.html
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Melissa Koss, San Diego Mesa College
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Lucretia Mahan, San Diego Mesa College
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Sam Merrill, San Diego Mesa College
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Paul Detwiler, San Diego Mesa College
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Tanya Dewey, University of Michigan-Ann Arbor
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Trophic Strategy

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Irrawaddy dolphins feed on fish, cephalopods, and crustaceans. Irrawaddy dolphins sometimes spit water while feeding, which may be used to herd fish.

Researchers have documented that in the Irrawaddy (or Ayerarwady) River, Myanmar, these dolphins engage in cooperative fishing with cast-net fishermen. Fishermen search for dolphins and call them by tapping a lahai kway, wooden key, on the sides of their boats. One or two lead dolphins then swim in smaller and smaller semi-circles herding the fish towards the shore. During cooperative fishing, the dolphins often dive deeply with their flukes aloft just after the net is cast and create turbulence under the surface around the outside of the net. The dolphins seem to benefit from the fishing by preying on fish that are confused by the sinking net, and those trapped around the edges of the lead line or stuck in the mud bottom just after the net is pulled up.

Animal Foods: fish; eggs; mollusks; aquatic crustaceans

Primary Diet: carnivore (Piscivore , Eats non-insect arthropods, Molluscivore )

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Koss, M.; L. Mahan and S. Merrill 2012. "Orcaella brevirostris" (On-line), Animal Diversity Web. Accessed April 27, 2013 at http://animaldiversity.ummz.umich.edu/site/accounts/information/Orcaella_brevirostris.html
author
Melissa Koss, San Diego Mesa College
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Lucretia Mahan, San Diego Mesa College
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Sam Merrill, San Diego Mesa College
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Paul Detwiler, San Diego Mesa College
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Tanya Dewey, University of Michigan-Ann Arbor
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Behavior

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Vocalizations of Irrawaddy dolphins in captivity within the Mahakam River, Indonesia included a single-component sonar signal, with the majority of frequencies captured at 60 kHz. Pulse trains were consistent with rates repeated at 40 to 60 kHz. Researchers theorize that Irrawaddy dolphins have a narrow sonar field. Little is known or recorded regarding courtship communication or other social signals.

Communication Channels: visual ; acoustic ; chemical

Perception Channels: visual ; tactile ; acoustic ; ultrasound ; chemical

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Koss, M.; L. Mahan and S. Merrill 2012. "Orcaella brevirostris" (On-line), Animal Diversity Web. Accessed April 27, 2013 at http://animaldiversity.ummz.umich.edu/site/accounts/information/Orcaella_brevirostris.html
author
Melissa Koss, San Diego Mesa College
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Lucretia Mahan, San Diego Mesa College
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Sam Merrill, San Diego Mesa College
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Paul Detwiler, San Diego Mesa College
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Tanya Dewey, University of Michigan-Ann Arbor
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Conservation Status

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Currently, the most immediate threat facing Irrawaddy dolphins is drowning in gill nets. The threat of gill net entanglement occurs primarily during the dry season (December to May), when dolphins settle in deep water pools. Dynamite and electric fishing occur in some important habitats. These activities are causing depletion of the dolphin's fish supply and noise from the explosions is potentially dangerous to dolphins. Due to the small population size and their narrow distribution, it is quite possible that dam construction anywhere within their habitat might critically endanger populations. Furthermore, uncontrolled tourism can harass dolphins in important habitats during the dry season and interfere with normal activities, such as feeding, resting, and socializing. Overfishing, collisions with boats and injuries from boat propellers are also threats to their survival.

US Federal List: no special status

CITES: appendix i

State of Michigan List: no special status

IUCN Red List of Threatened Species: critically endangered

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Koss, M.; L. Mahan and S. Merrill 2012. "Orcaella brevirostris" (On-line), Animal Diversity Web. Accessed April 27, 2013 at http://animaldiversity.ummz.umich.edu/site/accounts/information/Orcaella_brevirostris.html
author
Melissa Koss, San Diego Mesa College
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Lucretia Mahan, San Diego Mesa College
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Sam Merrill, San Diego Mesa College
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Paul Detwiler, San Diego Mesa College
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Tanya Dewey, University of Michigan-Ann Arbor
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Benefits

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The only way that Irrawaddy dolphins may have a negative economic impact on humans is that they share a common food source. Some fisherman believe the dolphins are pests because they reduce their catch, although this is unlikely.

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Koss, M.; L. Mahan and S. Merrill 2012. "Orcaella brevirostris" (On-line), Animal Diversity Web. Accessed April 27, 2013 at http://animaldiversity.ummz.umich.edu/site/accounts/information/Orcaella_brevirostris.html
author
Melissa Koss, San Diego Mesa College
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Lucretia Mahan, San Diego Mesa College
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Sam Merrill, San Diego Mesa College
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Paul Detwiler, San Diego Mesa College
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Tanya Dewey, University of Michigan-Ann Arbor
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Benefits

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In the Irrawaddy River these dolphins engage in cooperative fishing with cast-net fishermen. Irrawaddy dolphins increase the amount of fish the fishermen catch; therefore, they are of economic value. Additionally, Irrawaddy dolphins have brought ecotourism to communities in Indonesia and India.

Positive Impacts: food ; ecotourism

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Koss, M.; L. Mahan and S. Merrill 2012. "Orcaella brevirostris" (On-line), Animal Diversity Web. Accessed April 27, 2013 at http://animaldiversity.ummz.umich.edu/site/accounts/information/Orcaella_brevirostris.html
author
Melissa Koss, San Diego Mesa College
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Lucretia Mahan, San Diego Mesa College
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Sam Merrill, San Diego Mesa College
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Paul Detwiler, San Diego Mesa College
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Tanya Dewey, University of Michigan-Ann Arbor
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Associations

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Irrawaddy dolphins are top ecosystem predators, feeding on fish, cephalopods, and crustaceans. They are hosts for parasites including roundworms (Anisakis simplex), flukes (Braunina cordiformis), and tapeworms (Monorygma delphini). Healthy dolphin populations indicate a healthy marine ecosystem.

Ecosystem Impact: keystone species

Commensal/Parasitic Species:

  • roundworms (Anisakis simplex)
  • flukes (Braunina cordiformis)
  • tapeworms (Monorygma delphini)
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Koss, M.; L. Mahan and S. Merrill 2012. "Orcaella brevirostris" (On-line), Animal Diversity Web. Accessed April 27, 2013 at http://animaldiversity.ummz.umich.edu/site/accounts/information/Orcaella_brevirostris.html
author
Melissa Koss, San Diego Mesa College
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Lucretia Mahan, San Diego Mesa College
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Sam Merrill, San Diego Mesa College
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Paul Detwiler, San Diego Mesa College
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Tanya Dewey, University of Michigan-Ann Arbor
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Reproduction

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Little is currently known about mating systems in Irrawaddy dolphins. Since the animals live in small groups averaging between three to six individuals, it is presumed that breeding happens outside of those groups. The breeding season is between December and June; however, little is known about mating behavior. As in other dolphin species, it can be presumed that males mate with multiple females and compete over mates.

Mating System: polygynous

There is relatively little information on reproduction in Irrawaddy dolphins. The mating season in the northern hemisphere is reported to be December to June. The calving season seems to be from June to August, with noted exceptions. Data from Chilka Lake, India shows a low rate of breeding and production of only a single calf in three years. Additionally, gestation is approximately nine months, but data from two captive births estimate the gestation period at fourteen months. Captive born Irrawaddy dolphin calves measured 96 cm in length and weighed 12.3 kg. In the first seven months the calves increased in length by 59% and 266% in weight. Calves begin eating fish around six months and are fully weaned at about two years old. Adult length is achieved between three and five years old. Rate of sexual maturation is thought to be positively correlated with growth rate.

Breeding interval: Females may not reproduce yearly, in one population females give birth every 3 years.

Breeding season: Irrawaddy dolphins breed in December through June.

Average number of offspring: 1.

Range gestation period: 9 to 14 months.

Average weaning age: 24 months.

Range age at sexual or reproductive maturity (female): 3 to 6 years.

Range age at sexual or reproductive maturity (male): 3 to 6 years.

Key Reproductive Features: iteroparous ; seasonal breeding ; gonochoric/gonochoristic/dioecious (sexes separate); sexual ; viviparous

Average birth mass: 12300 g.

Average number of offspring: 1.

Irrawaddy dolphins are not completely weaned until 2 years. From birth to approximately seven months old, the calf survives solely on the nutrition from the mother. For the following seven months the calves stay within the pod and continue to receive nourishment from the mother while also eating fish. It is presumed the calves learn to prey on fish by copying behavior of the mother's and pod mates. There is little information on the rearing of calves. Currently it is not known if both males and females are involved in the upbringing of calves. Like most mammals, females invest heavily in their young.

Parental Investment: female parental care ; pre-fertilization (Provisioning, Protecting: Female); pre-hatching/birth (Provisioning: Female, Protecting: Female); pre-weaning/fledging (Provisioning: Female, Protecting: Female); pre-independence (Provisioning: Female, Protecting: Female); post-independence association with parents; extended period of juvenile learning

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Koss, M.; L. Mahan and S. Merrill 2012. "Orcaella brevirostris" (On-line), Animal Diversity Web. Accessed April 27, 2013 at http://animaldiversity.ummz.umich.edu/site/accounts/information/Orcaella_brevirostris.html
author
Melissa Koss, San Diego Mesa College
author
Lucretia Mahan, San Diego Mesa College
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Sam Merrill, San Diego Mesa College
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Paul Detwiler, San Diego Mesa College
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Tanya Dewey, University of Michigan-Ann Arbor
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Biology

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Believed to be reincarnated humans by some of the people of Laos (2), Irrawaddy dolphins are less active than many other dolphins with only the uppermost dorsal surface of the animal becomes visible during a slow rolling dive; they make only occasional low leaps and never bow-ride. Feeding together in groups of usually less than six, but as many as 15 (5), the Irrawaddy dolphin can dive for up to 12 minutes to feed on bony fish, crustaceans, cephalopods and fish eggs. Irrawaddy dolphins are known to spit water to herd fish, and have even been reported to stun large fish with a blow from the lower jaw, only to play with them before casting them aside (2). In some areas of Asia, fishermen consider the Irrawaddy dolphin to be a competitor for fish, but in other areas the fishermen attract them to the boat and encourage them to drive fish into the nets where the dolphins also benefit by preying on fish whose movements are confused by the nets and those that are momentarily trapped around the edges or in the mud (6). Irrawaddy dolphins communicate with clicks, creaks and buzzes (7) at a dominant frequency of about 60 kilohertz which is thought to be used for echolocation (8). Little is known about the reproductive biology of the Irrawaddy dolphin, but it is thought to breed between April and June in the Mahakam River, and gestation is estimated at 14 months and weaning after two years (2). This dolphin species is known to carry out daily migrations from the Semayang Lake in eastern Borneo to the Mahakam River, returning to the lake in the evening. In Indonesia, Irrawaddy dolphins move into tributaries at high water and into the main river during low water (4).
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Conservation

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Very few provisions have been made to conserve the dolphins or their habitat. The Irrawaddy dolphin is protected by law in Laos, Vietnam and Cambodia. Direct taking of cetaceans is prohibited in Bangladesh, India, Laos, Malaysia, Myanmar and Thailand. (9). In December 2005, the Department of Fisheries in Myanmar, established a protected area for Irrawaddy dolphins along a 74 kilometre segment of the Ayeyarwady River. Protective measures in the area include requiring fishermen to immediately release dolphins if found alive and entangled in their nets, and prohibiting the trade and catching or killing of dolphins and the use of electricity fishing and gill nets that obstruct the water-course, are more than 300 feet long, or spaced less than 600 feet apart (11). The problem of live captures for the aquarium trade has largely been solved by the uplisting of the species to Appendix I of the Convention on International Trade in Endangered Species (CITES) which prohibits international trade. Some captive breeding of this species has been successful (2).
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Description

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A particularly distinctive dolphin, the Irrawaddy has a rounded head with no beak and a flexible neck, causing visible creases behind the head (2). Although most closely related to the orca (4), the Irrawaddy dolphin is similar in body form to the beluga whale, but darker in colour, with a pale to dark grey back and a light underside. The dorsal fin is small, triangular and rounded, and the flippers are long and broad (2).
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Habitat

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Inhabits coastal, brackish and freshwaters of the tropical and sub-tropical Indo-Pacific (2).
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Range

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The Irrawaddy dolphin has a patchy distribution in the shallow, coastal waters of the Indo-Pacific from the Philippines to northeastern India. Freshwater subpopulations occur in three river systems: the Mahakam of Indonesia, the Ayeyarwady (previously Irrawaddy) of Burma, and the Mekong of Laos, Cambodia and Viet Nam. It is also found in completely or partially isolated brackish water bodies such as Chika Lake in India and Songkhla Lake in Thailand (1).
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Status

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The Irrawaddy dolphin is classified as Vulnerable (VU) on the IUCN Red List (1) and is listed on Appendix I of CITES (3). Geographically isolated populations in the Ayeyarwady River (Myanmar), Mahakam River (Indonesia), Malampaya Sound (Philippines), Mekong River (Lao PDR, Cambodia and Vietnam) and Songkhla Lake (India) are classified as Critically Endangered on the IUCN Red List (1).
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Threats

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The major threat to the Irrawaddy dolphin is incidental entanglement in gillnets (5) (9), but it is not believed to be at risk of imminent extinction (2). In the Ayeyarwady River it is also at risk from electrocution and prey depletion from electric fishing (10). Other issues affecting this dolphin include increasing pollution, construction of dams and the build-up of silt and sedimentation following severe erosion (2). Fishing with explosives also results in dolphin casualties (4). Most live captures are for the oceanarium trade in Asia (4), and hunting of this species is rare, occurring only in parts of India to harvest oil for the treatment of rheumatism (2).
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Deskripsi Umum Orcaella Brevirostris

provided by EOL authors
Pesut atau lumba-lumba air tawar adalah spesies mamalia air yang menghuni wilayah perairan tawar di India, Indocina, Filipina dan Kalimantan. Spesies ini pertama kali dideskripsikan oleh Sir Richard Owen tahun 1866 berdasarkan satu spesiemen yang ditemukan tahun 1852, di pelabuhan Vishakhapatnum di pantai timur India. Kadang-kadang pesut terdaftar dalam beragam famili yang terdiri dari pesut itu sendiri pada Monodontidae dan dalam Delphinapteridae. Sekarang ada persetujuan bahwa pesut termasuk famili Delphinidae. Spesies ini secara genetis terhubung dengan paus pembunuh. Nama spesies brevirostris berasal dari bahasa Latin yang berarti berparuh pendek. Tahun 2005, analisis genetik menunjukkan bahwa lumba-lumba sirip pendek Australia merupakan spesies kedua dari genus Orcaella. Orcaella brevirostris disebut juga Irrawaddy dolphin dalam bahasa Inggris, dan di Spanyol Delfín del Irrawaddy. Pesut jenis ini dulu populasinya sangat banyak di daerah sungai Mahakam Kalimantan. Tapi sekarang telah diklaim sebagai satwa langka karena populasinya yang berkurang drastis. Pesut memiliki tubuh yang berwarna kelabu hingga biru tua, bagian bawahnya berwarna lebih pucat dan tidak memiliki pola khas. Sirip punggungnya kecil dan membulat di tengah punggung. Penampilan pesut mirip dengan beluga, meski lebih berkerabat dengan orka. Spesies ini mempunyai melon (jaringan berlemak dan berminyak di kepala) fungsinya untuk ekolokasi. Sirip punggung yang terletak dua pertiga posterior di punggung, pendek, tumpul, dan segitiga. Sirip tangan panjang dan lebar. Secara keseluruhan ia berwarna cerah, namun lebih putih di bawah tubuh daripada di punggung. Pesut dewasa beratnya lebih dari 130 kg dan panjangnya 2,3 m psaat dewasa. Panjang maksimum yang tercatat adalah jantan 2,75 m dari Thailand. Pesut mencapai kedewasaan seksual pada 7 sampai 9 tahun. Di belahan bumi utara, perkawinan dilaporkan berlangsung pada bulan Desember sampai Juni. Masa hamilnya 14 bulan, melahirkan seekor anak setiap 2 hingga 3 tahun. Saat lahir panjangnya 1 m dan beratnya 10 kg dan umur pesut dapat mencapai 30 tahun.
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Diagnostic Description

provided by FAO species catalogs
The lrrawaddy dolphin resembles the finless porpoise, but unlike that species, it has dorsal fin. The fin is small and triangular, with a bluntly rounded tip, and is set just behind midback. The large flippers have curved leading edges and rounded tips. The head is blunt, with no beak; the mouthline is straight, and there may be a visible neck crease. The U-shaped blowhole is open toward the front, the reverse of the situation in most dolphin species. The back and sides of lrrawaddy dolphins are grey to bluish grey; the belly is somewhat lighter. Tooth counts are 17 to 20 (upper) and 15 to 18 (lower) in each row. The teeth have slightly expanded crowns. Can be confused with: lrrawaddy dolphins can be confused with finless porpoises or dugongs in the parts of their range where these species overlap. When a clear view is obtained, lrrawaddy dolphins are distinguishable because neither of the other species has a dorsal fin.
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Marine mammals of the world. Jefferson, T.A., S. Leatherwood & M.A. Webber - 1993. FAO species identification guide. Rome, FAO. 320 p. 587 figs. . 
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Food and Agriculture Organization of the UN
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Size

provided by FAO species catalogs
This is a relatively small dolphin: adults range from 2 to 2.75 m. Scant evidence indicates that the length at birth is about 1 m.
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Marine mammals of the world. Jefferson, T.A., S. Leatherwood & M.A. Webber - 1993. FAO species identification guide. Rome, FAO. 320 p. 587 figs. . 
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Food and Agriculture Organization of the UN
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Brief Summary

provided by FAO species catalogs
Groups of fewer than 6 individuals are most common, but sometimes up to 15 dolphins are seen together. lrrawaddy dolphins have been seen in the same area as bottlenose dolphin and Indo-Pacific hump-backed dolphin. lrrawaddy dolphins are not particularly active, but do make low leaps on occasion. They are not known to bowride. The calving season is not well known. Some calves appear to have been born from June to August, but 1 captive female gave birth in December. Fishes, cephalopods, and crustaceans are taken as food. They sometimes spit water while feeding, apparently to herd fish.
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Marine mammals of the world. Jefferson, T.A., S. Leatherwood & M.A. Webber - 1993. FAO species identification guide. Rome, FAO. 320 p. 587 figs. . 
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Food and Agriculture Organization of the UN
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Benefits

provided by FAO species catalogs
Shark gillnets in Australia and fish traps and other types of nets throughout the range are known to take some lrrawaddy dolphins. Some small-scale hunting by local people probably occurs in many areas of its range. IUCN: Insufficiently known.
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Marine mammals of the world. Jefferson, T.A., S. Leatherwood & M.A. Webber - 1993. FAO species identification guide. Rome, FAO. 320 p. 587 figs. . 
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Irrawaddy dolphin

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The Irrawaddy dolphin (Orcaella brevirostris) is a euryhaline species of oceanic dolphin found in scattered subpopulations near sea coasts and in estuaries and rivers in parts of the Bay of Bengal and Southeast Asia. It closely resembles the Australian snubfin dolphin (of the same genus, Orcaella), which was not described as a separate species until 2005. It has a slate blue to a slate gray color. Although found in much of the riverine and marine zones of South and Southeast Asia, the only concentrated lagoon populations are found in Chilika Lake in Odisha, India and Songkhla Lake in southern Thailand.[4]

Taxonomy

Irrawaddy dolphin skeleton specimen exhibited in Museo di storia naturale e del territorio dell'Università di Pisa

One of the earliest recorded descriptions of the Irrawaddy dolphin was by Sir Richard Owen in 1866 based on a specimen found in 1852, in the harbour of Visakhapatnam on the east coast of India.[5] It is one of two species in its genus. It has sometimes been listed variously in a family containing just itself and in the Monodontidae and Delphinapteridae. Widespread agreement now exists to list it in the family Delphinidae.

Etymology and local names

The species name brevirostris is from the Latin meaning short-beaked. It is very closely related to the Australian snubfin dolphin (Orcaella heinsohni). The two snubfin dolphins were only recognised as separate species in 2005 when a genetic analysis showed that the population found along the coast of northern Australia forms a second species in the genus Orcaella. The Orcaella dolphins are close relatives of the oceanic dolphins in Globicephalinae subfamily.

Vernacular names for the Irrawaddy dolphin include:[6][5][7][8]: 1 [9]: 105 

  • Thai: โลมาอิรวดี loma irawadi, โลมาหัวบาตร loma hua bat ("alms-bowl dolphin", due to the shape of their heads)
  • Odia: ଶିଶୁମାର sisumāra, ଭୁଆସୁଣୀ ମାଛ bhuāsuṇi mācha (lit. oil-yielding dolphin), lagoon area local name- ଖେରା kherā
  • Tagalog: lampasut
  • Bengali: শুশুক shushuko
  • Kutai Malay: pesut mahakam, ikan pesut
  • Khmer: ផ្សោត ph’sout
  • Lao: ປາຂ່າ pa’kha
  • Sarawak Malay: empesut, pesut
  • Burmese: ဧရာဝတီ လင်းပိုင် eyawadi lăbaing

The word pesut comes from hearing the sound made from the dolphin exhaling from its blowhole.[8]: 1 [10]: 11 

Description

1878 illustration

The Irrawaddy dolphin's colour is grey to dark slate blue, paler underneath, without a distinctive pattern. The dorsal fin is small and rounded behind the middle of the back. The forehead is high and rounded; the beak is lacking. The front of its snout is blunt. The flippers are broad and rounded. The finless porpoise (Neophocaena phocaenoides) is similar and has no back fin; the humpback dolphin (Sousa chinensis) is larger, and has a longer beak and a larger dorsal fin.[5] It ranges in weight from 90 to 200 kg (200 to 440 lb) with a length of 2.3 m (7.5 ft) at full maturity.[11][12] Maximum recorded length is 2.75 m (9.0 ft) of a male in Thailand.[6]

The Irrawaddy dolphin is similar to the beluga in appearance, though most closely related to the killer whale. It has a large melon and a blunt, rounded head, and the beak is indistinct. Its dorsal fin, located about two-thirds posterior along the back, is short, blunt, and triangular.[12] Dorsal fin shapes differ from one Irrawaddy dolphin to another.[12] The flippers are long and broad.[13] These dolphins are usually two-toned, with the back and sides being gray to bluish-gray and the belly lighter.[13] Unlike any other dolphin, the Irrawaddy's U-shaped blowhole is on the left of the midline and opens towards the front of the dolphin.[13] Its short beak appears very different from those of other dolphins, and its mouth is known for having 12-19 peg-like teeth on each side of the jaws.[13]

Behaviour

Closeup of an Irrawaddy dolphin jumping in the Mekong River

Communication is carried out with clicks, creaks, and buzzes at a dominant frequency of about 60 kilohertz, which is thought to be used for echolocation. Bony fish and fish eggs, cephalopods, and crustaceans are taken as food. Observations of captive animals indicate food may be taken into the mouth by suction. Irrawaddy dolphins are capable of squirting streams of water that can reach up to 1.5 metres (4 ft 11 in); this distinct behaviour has been known for herding fish into a general area for hunting.[14] They do this sometimes while spyhopping and during feeding, apparently to expel water ingested during fish capture or possibly to herd fish. Some Irrawaddy dolphins kept in captivity have been trained to do spyhopping on command. The Irrawaddy dolphin is a slow swimmer, but swimming speeds of 20–25 kilometres per hour (12–16 mph) were reported when dolphins were being chased in a boat.[15] In a year, females can sometimes swim as fast as average of 45 km, males can swim up to average of 100 km.[8]: 2 

Most Irrawaddy dolphins are shy of boats, not known to bow-ride, and generally dive when alarmed. They are relatively slow moving but can sometimes be seen spyhopping and rolling to one side while waving a flipper and occasionally breaching. They are generally found in groups of 2-3 animals, though sometimes as many as 25 individuals have been known to congregate in deep pools. Groups of fewer than six individuals are most common, but sometimes up to 15 dolphins are seen together.[15][16] Traveling and staying in groups not only enables Irrawaddy dolphins to hunt, but it also creates and maintains social bonds and allows copulation to occur.[17] There are a few groups of dolphins in Brunei Bay seen near the mouths of Temburong and Aloh Besar rivers which are tame and gather around fisherman boats that catch fish during the fishing season from around December to February.[10]

It surfaces in a rolling fashion and lifts its tail fluke clear of the water only for a deep dive. Deep dive times range from 30 to 150 seconds to 12 minutes. When 277 group dives were timed (time of disappearance of the last dolphin in the group to the emergence of the first dolphin in the group) in Laos, mean duration was 115.3 seconds with a range of 19 seconds to 7.18 minutes.[6]

Interspecific competition has been observed when Irrawaddy dolphins were forced inshore and excluded by more specialized dolphins. When captive humpback dolphins (Sousa chinensis) and Irrawaddy dolphins were held together, reportedly the Irrawaddy dolphins were frequently chased and confined to a small portion of the tank by the dominant humpbacks. In Chilika Lake, local fishers say when Irrawaddy dolphins and bottlenose dolphins meet in the outer channel, the former get frightened and are forced to return toward the lake.[5]

Mating

A female or male dolphin will attempt to pursue a mate for about a few minutes. They intertwine facing their bellies together and begin to copulate for 40 seconds. Once copulation has occurred, the dolphins will break away from each other and set off in different directions.[17]

Reproduction

1878 illustration of a foetus in the uterus
Mother with calf from Sundarban, Bangladesh

These dolphins are thought to reach sexual maturity at seven to nine years. In the Northern Hemisphere, mating is reported from December to June. Its gestation period is 14 months; cows give birth to a single calf every two to three years. Length is about 1 m (3.3 ft) at birth. Birth weight is about 10 kg (22 lb). Weaning is after two years. Lifespan is about 30 years.

Feeding

There are plenty of food items that this dolphin feeds upon. They include fish, crustaceans, and cephalopods. During foraging periods, herds of about 7 dolphins will circle around prey and trap their victim. These prey entrapments occur slightly below the water surface level.[17]

Habitat and subpopulations

Irrawaddy Dolphin in Chilika Lake, Odisha

Although sometimes called the Irrawaddy river dolphin, it is not a true river dolphin, but an oceanic dolphin that lives in brackish water near coasts, river mouths, and estuaries. It has established subpopulations in freshwater rivers, including the Ganges and the Mekong, as well as the Irrawaddy River from which it takes its name. Its range extends from the Bay of Bengal to New Guinea and the Philippines, although it does not appear to venture off shore. It is often seen in estuaries and bays in Borneo Island, with sightings from Sandakan in Sabah, Malaysia, to most parts of Brunei and Sarawak, Malaysia.[18]: 170  A specimen was collected at Mahakam River in East Kalimantan,[1] the local name pesut mahakam comes from it.[8]: 1 

Presence of the species in Chinese, Taiwanese, and Hong Kong's waters has been questioned as the reported sightings have been considered unreliable,[19] and the easternmost of ranges along Eurasian continent is in Vietnam.

No range-wide survey has been conducted for this vulnerable species; however, the worldwide population appears to be over 7,000. In India, Irawaddy dolphins are mostly found in Chilika Lake. Known subpopulations of Irrawaddy dolphins are found in eight places, listed here in order of population, including conservation status.

Chilka Lake, Odisha, India, habitat of Irrawaddy dolphins
  1. Bangladesh: ~3500 (VU) in coastal waters of the Bay of Bengal[20] and 451 (VU) in the brackish Sundarbans mangrove forest[21]
  2. India: 156 (VU) in the brackish-water Chilika Lake, Odisha.[22] Presence recorded from Sundarbans National Park, West Bengal also.
  3. Laos, Cambodia: 92 (CR) in a 190-km (118-mi) freshwater stretch of the Mekong River[23][24]
  4. Indonesia: ~70 (CR), in a 420-km (260-mi) stretch of the freshwater Mahakam River
  5. Philippines: ~42 (CR) in the brackish inner Malampaya Sound.[25] Researchers are studying the recent discovery of 30-40 dolphins sighted in the waters of Pulupandan and Bago, Negros Occidental, in Western Visayas.[26]
  6. Myanmar (Burma): ~58-72 (CR) in a 370-km (230-mi) freshwater stretch of the Ayeyarwady River
  7. Thailand: less than 50 (CR) in the brackish Songkhla Lake[1]

Interaction with humans

Irrawaddy dolphins have a mutualistic relationship of co-operative fishing with traditional fishers. Fishers in India recall when they would call out to the dolphins, by tapping a wooden key also known as a lahai kway,[27] against the sides of their boats, asking the Irrawaddys to drive fish into their nets.[28] In Burma, in the upper reaches of the Ayeyarwady River, Irrawaddy dolphins drive fish towards fishers using cast nets in response to acoustic signals from them. The fishermen attempt to gain the attention of the dolphins through various efforts such as using a cone-shaped wooden stick to drum the side of their canoes, striking their paddles to the surface of the water, jingling their nets, or making calls that sound turkey-like. A pod of dolphins that agrees to work alongside the fisherman will entrap a school of fish in a semicircle, guiding them towards the boat.[29] In return, the dolphins are rewarded with some of the fishers' bycatch.[30] Historically, Irrawaddy River fishers claimed particular dolphins were associated with individual fishing villages and chased fish into their nets. An 1879 report indicated legal claims were frequently brought into native courts by fishers to recover a share of the fish from the nets of a rival fisher that the plaintiff's dolphin was claimed to have helped fill.[6]

Folk stories

Laotians and Cambodians have a common belief that the Irrawaddy dolphins are reincarnations of their ancestors. Some even claim that the dolphins have saved drowning villagers and protected people from attacks by crocodiles. Their beliefs and experiences have led the people of Laos and Cambodia to live peacefully alongside one another for ages.[31] The West Kalimantan people have a similar story where the dolphins were naughty children that ate a pot of rice reserved for the shaman, but their mouths scalded and jumped into the water to cool themselves, but later transformed into these dolphins.[8]: 5–6 

Buddhist Khmer and Vietnamese fishermen have regarded the Orcaella as a sacred animal. If caught in fishing nets, they release the dolphin from the rest of the catch. In contrast, Muslim Khmer fisherman kill them for food. This has led to the dolphin becoming reputed to recognize the local languages of the area and it approaching areas of the Khmer Muslim community with caution.[32]

Threats

Fishermen with fishnets in Bangladesh

Irrawaddy dolphins are more susceptible to human conflict than most other dolphins that live farther out in the ocean. Drowning in gillnets is the main threat to them throughout their range. Between 1995 and 2001, 38 deaths were reported and 74% died as a result of entanglement in gillnets with large mesh sizes.[33] The majority of reported dolphin deaths in all subpopulations is due to accidental capture and drowning in gillnets and dragnets, and in the Philippines, bottom-set crabnets. In Burma, electrofishing, gold mining, and dam building are also serious and continuing threats. Though most fishers are sympathetic to the dolphins' plight, abandoning their traditional livelihood is difficult for them.[1]

Another identified threat towards the Irrawaddy dolphins was noise pollution from high-speed vessels. This caused the dolphins to dive significantly longer than usual. The Irrawaddy dolphins always changed directions when they encountered these large vessels.[33]

In recent years, Laotians and Cambodians developed techniques of using explosives for fishing. The government of Laos has made use of such tactics illegal, but few regulations have been made in neighboring Cambodia, where explosives are sold in local markets and the practice of using fishnets has been abandoned. The practice of using explosives instead has become very popular and led to a steady decline of populations of fish, and especially the dolphins swimming in the area. Although Laotians may not use explosives, they do use nylon gillnets, which pose another large threat to the survival of the Irrawaddy. Some dolphins accidentally become entangled in the net. Poor fishermen refuse to cut and destroy their nets because it would result in too great of an economic loss to save one Irrawaddy dolphin.[31]

In Laos, a dam across the Mekong River is planned. This could threaten the existence of the endangered Irrawaddy dolphins in downstream Cambodia. Laos's government decision is to forge the dam upstream of the core habitat of the Irrawaddy dolphins. This could precipitate the extinction of this specific species in the Mekong River. The dam builders' proposal is to use explosives to dig out the tons of rock. This will create strong sound waves that could possibly kill the Irrawaddy dolphins due to their highly sensitive hearing structures.

In several Asian countries, Irrawaddy dolphins have been captured and trained to perform in public aquaria. Their charismatic appearance and unique behaviors, including spitting water, spyhopping, and fluke-slapping, make them very popular for shows in dolphinaria. The commercial motivation for using this dolphin species is high because it can live in freshwater tanks and the high cost of marine aquarium systems is avoided. The region within and near the species' range has developed economically; theme parks, casinos, and other entertainment venues that include dolphin shows have increased.

In 2002, there were more than 80 dolphinariums in at least nine Asian countries.[34]

Collateral deaths of dolphins due to blast fishing were once common in Vietnam and Thailand. In the past, the most direct threat was killing them for their oil.

The IUCN lists five of the seven subpopulations as endangered, primarily due to drowning in fish nets.[1] For example, the Malampaya population, first discovered and described in 1986, at the time consisted of 77 individuals. Due to anthropogenic activities, this number dwindled to 47 dolphins in 2007.[35] In the Mahakam River in Borneo, 73% of dolphin deaths are related to entanglement in gillnets, due to heavy fishing and boat traffic.[36]

Tourism

The Irrawaddy dolphins in Asia are increasingly threatened by tourist activity, such as large numbers of boats circulating the areas in which they live. The development of tours and boats has put a large strain on the dolphins.[37]

Disease

Cutaneous nodules were found present in various vulnerable populations of Irrawaddy dolphins. A more precise estimate of the affected dolphins is six populations. Although the definite fate of this emergent disease is unknown, the species is at risk.[38]

Conservation

The Irrawaddy dolphin's proximity to developing communities makes the effort for conservation difficult.[39] Entanglement in fishnets and degradation of habitats are the main threats to Irrawaddy dolphins. Conservation efforts are being made at international and national levels to alleviate these threats.

International efforts

Protection from international trade is provided by the Convention on International Trade in Endangered Species of Wild Fauna and Flora (CITES). Enforcement, though, is the responsibility of individual countries.[1] While some international trade for dolphinarium animals may have occurred, this is unlikely to have ever been a major threat to the species.

Some Irrawaddy dolphin populations are classified by the IUCN as critically endangered; in Lao PDR, Cambodia, Viet Nam (Mekong River sub-population), Indonesia (Mahakam River sub-population, Borneo), Myanmar (Ayeyarwady/Irrawaddy River sub-population), the Philippines (Malampaya Sound sub-population), and Thailand (Songkhla Lake sub-population). Irrawaddy dolphins in general however, are IUCN listed as an Endangered species, which applies throughout their whole range.[1] In 2004, CITES transferred the Irrawaddy dolphin from Appendix II to Appendix I, which forbids all commercial trade in species that are threatened with extinction.[40]

The UNEP-CMS Action Plan for the Conservation of Freshwater Populations of Irrawaddy dolphins notes that multiple-use protected areas will play a key role for conserving freshwater populations. Protected areas in fresh water could be a particularly effective conservation tool and can facilitate management, due to the fidelity of the species to relatively circumscribed areas. The Action Plan provides details on strategies for mitigating by-catch that includes:

-establishing core conservation areas where gillnetting is banned or severely restricted
-promoting net attendance rules and providing training on the safe release of entangled dolphins
-initiating programs to compensate fishers for damage caused to their nets by entangled dolphins that are safely released
-providing alternative or diversified employment options for gillnet fishers
-encouraging the use of fishing gear that does not harm dolphins, by altering or establishing fee structures for fishing permits to make gillnetting more expensive while decreasing the fees for nondestructive gear
-experimenting with acoustical deterrents and reflective nets.[41]

The Irrawaddy dolphin is listed on both Appendix I and Appendix II of the Convention on the Conservation of Migratory Species of Wild Animals (CMS).[42] It is listed on Appendix I[42] as this species has been categorized as being in danger of extinction throughout all or a significant proportion of their range and CMS Parties strive towards strictly protecting these animals, conserving or restoring the places where they live, mitigating obstacles to migration and controlling other factors that might endanger them, as well on Appendix II[42] as it has an unfavourable conservation status or would benefit significantly from international co-operation organized by tailored agreements.[43]

The species is also covered by the Memorandum of Understanding for the Conservation of Cetaceans and Their Habitats in the Pacific Islands Region (MoU).[44]

National efforts

Several national efforts are resulting in the reduction of threats to local Irrawaddy dolphin subpopulations:

Bangladesh

Satellite image of the Sundarbans

Portions of Irrawaddy dolphin habitat in the Sundarbans mangrove forest of Bangladesh are included within 139,700 ha (539 sq mi) of three wildlife sanctuaries, which are part of the Sunderbans World Heritage Site. The Wildlife Conservation Society is working with the Bangladesh Ministry of Environment and Forests to create protected areas for the 6000 remaining dolphins[7][45]

Cambodia

Irrawaddy dolphins are fully protected as an endangered species under Cambodian fishery law.[46] In 2005, The World Wide Fund For Nature (WWF) established the Cambodian Mekong Dolphin Conservation Project with support from government and local communities. The aim is to support the survival of the remaining population through targeted conservation activities, research, and education.[47] In January 2012, the Cambodian Fisheries Administration, the Commission for Conservation and Development of Mekong River Dolphin Eco-tourism Zone, and WWF signed the Kratie Declaration on the Conservation of the Mekong River Irrawaddy Dolphin, an agreement binding them to work together, and setting out a roadmap for dolphin conservation in the Mekong River.[48] On 24 August 2012, the Cambodian government announced that 180-kilometre-long (110 mi) stretch of the Mekong River from eastern Kratie province to the border with Laos has been stated as limit fishing zone which uses floating houses, fishing cages and gill nets are disallowed, but simple fishing is allowed.[49] This area is patrolled by a network of river guards, specifically to protect dolphins. Between January and February 2006, a dozen Irrawaddy dolphins were found dead. The discovery of 10 new baby dolphins on the coast of Cambodia was a relief and gave hope that the endangered Irrawaddy dolphins would make a comeback in population. Since the endangerment was evident, 66 guards have been posted on the coast of Cambodia to protect these dolphins, and only two deaths have been reported since these efforts. To make more efforts to keep these animals from going extinct, the use of fishing nets on the coast of Cambodia is banned, as well.[50]

Irrawaddy dolphin in the Sundarbans

India

The Irrawaddy dolphin (under the common name of snubfin dolphin, with the scientific name misspelled as Oreaella brevezastris) is included the Indian Wildlife Protection Act,[51] Schedule I,[52] which bans their killing, transport and sale of products.[7] A major restoration effort to open a new mouth between Chilika Lake and the Bay of Bengal in 2000 was successful in restoring the lake ecology and regulating the salinity gradient in the lake waters, which has resulted in increases in the population of Irrawaddy dolphin due to increase of prey species of fish, prawns, and crabs.[53][54]

Indonesia

A conservation program, entitled Conservation Foundation for the Protection of Rare Aquatic Species of Indonesia, focused on protecting the Irrawaddy dolphin population and their habitat, the Mahakam River. The program not only educates and surveys the public, but also monitors the dolphin population and their habitat. A prime example of this is the establishment of patrols in several villages.[55]

Specimen in Kalimantan

Laos

Some major concern for the population in the Mekong River has arisen due to numerous threats. In the 1970s, many Irrawaddy dolphins were slaughtered for oil, and soon after, intensive fishing practices with explosives and gillnets began. As of now, the dolphins are protected in Cambodia and Laos Democratic Republic. Now, both explosive fishing and use of gillnets is restricted in many of the Irrawaddy dolphin's habitats.[24] Canadian conservationist Ian Baird set up the Lao Community Fisheries and Dolphin Protection Project to study the Irrawaddy dolphins in the Laotian part of the Mekong. Part of this project compensated fishers for the loss of nets damaged to free entangled dolphins. This project was expanded to include Cambodia, after the majority of the dolphin population was determined to have been killed or migrated to Laos' southern neighbor.[56] The Si Phan Don Wetlands Project has successfully encouraged river communities to set aside conservation zones and establish laws to regulate how and when fish are caught.[57] In April, after a 200-kg Irrawaddy dolphin was found dead on the coast of Laos, the death toll for Irrawaddy dolphins is now five in 2015. Usually, Irrawaddy dolphins are found dead with bruises and scars on their body, being killed by illegal poaching, but this Irrawaddy dolphin was found dead because of old age. She is the oldest and largest Irrawaddy dolphin researchers have discovered. The dolphin was 2.5 m (8 ft 2 in) in length and most likely in her late 20s.

Myanmar

Myanmar's Department of Fisheries took charge in December 2005, and instituted a protected region in a 74 km (46 mi) segment of the Ayeyarwady River between Mingun and Kyaukmyaung and created multiple provisions, as well.[29] Protective measures in the area include mandatory release of entangled dolphins, prohibition of the catching or killing of dolphins and trade in whole or parts of them, and the prohibition of electrofishing and gillnets more than 91 metres (300 ft) long, or spaced less than 180 metres (600 ft) apart.[7] Mercury poisoning and habitat loss from gold-mining dredging operations in the river have been eliminated[58]

Philippines

In 2000, Malampaya Sound was proclaimed a protected seascape. This is the lowest possible prioritization given to a protected area.[41] Malampaya Sound Ecological Studies Project was initiated by the WWF. With technical support provided by the project, the municipality of Taytay and the Malampaya park management developed fishery policies to minimize the threats to the Irrawaddy dolphin from bycatch capture. Gear studies and gear modification to conserve the dolphin species were implemented. The project was completed in 2007.[59] In 2007, the Coral Triangle Initiative, a new multilateral partnership to help safeguard the marine and coastal resources of the Coral Triangle, including the Irrawaddy dolphin subpopulation in Malampaya Sound, was launched.[60][61] In 2006, a new population was discovered in Guimaras island in the Visayas. In 2015, another new population was discovered in Bago in Negros Occidental, part of Negros island in the Visayas.[62]

Irrawaddy dolphin on Mekong River at Kratié, Cambodia

Thailand

In 2002, the Marine and Coastal Resources Department was assigned to protect rare aquatic animals such as dolphins, whales, and turtles in Thai territorial waters. To protect the dolphins, patrol vessels ensure boats stay at least 30 m (98 ft) away from dolphins and no chasing of or running through schools of dolphins occurs. Many fishermen on the Bang Pakong River, Prachinburi Province, have been persuaded by authorities to stop shrimp fishing in a certain area and 30 to 40 fishing boats have been modified so they can offer dolphin sightseeing tours.[63] A total of 65 Irrawaddi dolphins has been found dead along the coast of Trat Province in the past three years.[64] The local fishing industry is blamed for the deaths of the dolphins.[65] In January 2013, over a dozen dead Irrawaddy dolphins were found on the coast of Thailand. These dolphins were said to be dead because of a lack of oxygen. Dolphins are mammals, and unlike other animals that live in the sea, they must come to the surface for air. Many of the dolphins are found dead in the water, and others were washed ashore, said to have been dead for a few days. Also, in the first week of February 2013, as many as four Irrawaddy dolphins were found dead.[66] The Natural Resources and Environment Ministry is studying artificial breeding options in 2022 to try to save the last herd of Irrawaddy dolphins in Songkhla Lake, which is on the verge of extinction after only 14 were found in the most recent survey.[67] They also intend to list dolphins as a protected species, Thailand's highest level of animal protection.[68]

Malaysia

In 2008, the Department of Forestry and Sarawak Forestry Cooperative in Sarawak established a protected area for Irrawaddy dolphins in Santubong and Damai (Kuching Wetland).[7] Furthermore, they plan to establish more beaches in Miri as protected areas for them. The protection measures in the area include prohibition of catching or killing of dolphins and trade in whole or parts of them, and prohibiting the use of gillnets. The government may also start small- and medium-scale research of this species at Sarawak Malaysia University with sponsorship from Sarawak Shell.

Vietnam

In 2012 in Vietnam, a group of scientists took in four Irrawaddy dolphins and provided them with medical care to see how they would survive. However, they found this to be the first case they saw of Irrawaddy dolphins having bacterial infections. The bacterial infection, chorioamnionitis, is common in many marine animals, but when these few dolphins were taken in, the scientists discovered this same bacterial infection for the first time in this group of dolphins. This disease mostly affects animals that are pregnant because the infection occurs through the umbilical cord and goes into the maternal bloodstream. One of the dolphins was pregnant and before her death was found circling around the bottom of the pool and was found dead early the next morning. This bacterial infection affects many organs in the body of the animal.[69]

See also

References

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Irrawaddy dolphin: Brief Summary

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The Irrawaddy dolphin (Orcaella brevirostris) is a euryhaline species of oceanic dolphin found in scattered subpopulations near sea coasts and in estuaries and rivers in parts of the Bay of Bengal and Southeast Asia. It closely resembles the Australian snubfin dolphin (of the same genus, Orcaella), which was not described as a separate species until 2005. It has a slate blue to a slate gray color. Although found in much of the riverine and marine zones of South and Southeast Asia, the only concentrated lagoon populations are found in Chilika Lake in Odisha, India and Songkhla Lake in southern Thailand.

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Habitat

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in coastal, brackish and fresh waters, tropical and subtropical

Reference

van der Land, J. (ed). (2008). UNESCO-IOC Register of Marine Organisms (URMO).

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IUCN Red List Category

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Vulnerable (VU)

Reference

IUCN (2008) Cetacean update of the 2008 IUCN Red List of Threatened Species.

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Perrin, William [email]