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Morphology

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Rockhopper penguins measure about 55 centimeters in length and weigh around 2.5 kilograms. These birds stand upright on two short feet. Their legs are set far back on the body. The waterproof coat, composed of feathers that average 2.9 centimeters in length, is white on the underside and bluish-black on the top. The head has bright yellow plumage on the brow; the yellow feathers extend along the sides. The top of the head has spiked black feathers. The wings are strong, stiff, narrow and flipper-like. Rockhopper penguins have tiny eyes.

Range mass: 2000 to 3000 g.

Average length: 55 cm.

Other Physical Features: endothermic ; homoiothermic; bilateral symmetry

Sexual Dimorphism: sexes alike

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Phelan, D. 1999. "Eudyptes chrysocome" (On-line), Animal Diversity Web. Accessed April 27, 2013 at http://animaldiversity.ummz.umich.edu/site/accounts/information/Eudyptes_chrysocome.html
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Life Expectancy

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The average lifespan of a rockhopper penguin is 10 years.

Average lifespan
Status: wild:
10 years.

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Phelan, D. 1999. "Eudyptes chrysocome" (On-line), Animal Diversity Web. Accessed April 27, 2013 at http://animaldiversity.ummz.umich.edu/site/accounts/information/Eudyptes_chrysocome.html
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Habitat

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Rockhopper penguins are found in high grasses called tussocks, where they make burrows and nest. As their name implies, they live on rocky shorelines.

Habitat Regions: temperate ; terrestrial ; saltwater or marine

Terrestrial Biomes: savanna or grassland

Aquatic Biomes: coastal

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Phelan, D. 1999. "Eudyptes chrysocome" (On-line), Animal Diversity Web. Accessed April 27, 2013 at http://animaldiversity.ummz.umich.edu/site/accounts/information/Eudyptes_chrysocome.html
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Distribution

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Rockhopper penguins are found on islands in the southern ocean, such as the Falkland Islands. They occur farther north than many other penguin species.

Biogeographic Regions: neotropical (Native ); atlantic ocean (Native )

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Phelan, D. 1999. "Eudyptes chrysocome" (On-line), Animal Diversity Web. Accessed April 27, 2013 at http://animaldiversity.ummz.umich.edu/site/accounts/information/Eudyptes_chrysocome.html
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Untitled

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Rockhopper penguins keep warm by their well-developed fat layer and system for maintaining heat.

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Phelan, D. 1999. "Eudyptes chrysocome" (On-line), Animal Diversity Web. Accessed April 27, 2013 at http://animaldiversity.ummz.umich.edu/site/accounts/information/Eudyptes_chrysocome.html
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Behavior

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Their loud cry, "ecstatic vocalization", is used to announce their presence, attract a mate, or announce the boundaries of their territory. As well as vocalizing, these birds shake their heads and cause their yellow eyebrows to fly into a "halo" in order to attract a mate.

Communication Channels: visual ; acoustic

Perception Channels: visual ; tactile ; acoustic ; chemical

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Phelan, D. 1999. "Eudyptes chrysocome" (On-line), Animal Diversity Web. Accessed April 27, 2013 at http://animaldiversity.ummz.umich.edu/site/accounts/information/Eudyptes_chrysocome.html
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Conservation Status

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It is estimated that rockhopper penguins have undergone a decline of more than 30% in their total population size over the past 30 years. For this reason, they are classified as vulnerable by the IUCN. If the decline continues, they may be uplisted to endangered in the near future. Threats to rockhopper penguin populations include commercial fishing, which reduces the amount of available prey, and oil spills.

CITES: no special status

IUCN Red List of Threatened Species: vulnerable

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Phelan, D. 1999. "Eudyptes chrysocome" (On-line), Animal Diversity Web. Accessed April 27, 2013 at http://animaldiversity.ummz.umich.edu/site/accounts/information/Eudyptes_chrysocome.html
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Benefits

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Penguins are a tourist attraction, and they are one of the main reasons people travel to the Falkland Islands and other habitats of these penguins.

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Phelan, D. 1999. "Eudyptes chrysocome" (On-line), Animal Diversity Web. Accessed April 27, 2013 at http://animaldiversity.ummz.umich.edu/site/accounts/information/Eudyptes_chrysocome.html
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Trophic Strategy

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Rockhopper penguins eat primarily krill (Euphausiacea). They also eat squid and other crustaceans. They make daily trips to the sea to forage.

Animal Foods: fish; aquatic crustaceans

Primary Diet: carnivore (Eats non-insect arthropods)

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Phelan, D. 1999. "Eudyptes chrysocome" (On-line), Animal Diversity Web. Accessed April 27, 2013 at http://animaldiversity.ummz.umich.edu/site/accounts/information/Eudyptes_chrysocome.html
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Reproduction

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Mating calls, which are species specific, are called "ecstatic vocalization." This draws attention to the bird and announces its intentions. Penguins mate with the same partners from previous years.

Mating System: monogamous

Rockhopper penguins typically mate in the early spring or late summer, enabling the young to go to the sea in the mid-summer. They mate in vast colonies and lay up to two eggs, although sometimes pairs "adopt" a third egg. The first egg is usually 20-50% smaller than second one. The small egg is usually lost, although it is capable of maturing into a normal bird. Adopted eggs are also typically lost. After each egg is laid, it is turned over to the male who sits on it and keeps it in his brood pouch for the next four months until it hatches.

Breeding interval: Rockhopper penguins breed once yearly.

Range eggs per season: 1 to 2.

Key Reproductive Features: iteroparous ; seasonal breeding ; gonochoric/gonochoristic/dioecious (sexes separate); sexual ; oviparous

While the male penguin sits on the incubating egg, he is nourished by the female, or else he fasts for the entire period. If the female does not return with food for the chick once it has hatched, the male produces "penguin's milk" from his digestive system and regurgitates it for the baby.

Parental Investment: precocial ; pre-fertilization (Provisioning, Protecting: Female); pre-hatching/birth (Protecting: Male); pre-weaning/fledging (Provisioning: Male, Female, Protecting: Male)

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Biology

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A gregarious species, the southern rockhopper penguin breeds in large colonies that may comprise over a hundred thousand nests. Breeding pairs are monogamous, and usually return to the same nest every year. Egg-laying commences around November, with the female usually producing a clutch of two eggs of unequal size (2). Although, in general, only the chick from the larger egg survives to maturity, populations on the Falkland Islands frequently succeed in raising both (5). Incubation takes around 33 days, with both parent birds taking it in turns to sit on the eggs for extended periods of a time, whilst the other forages for food. Incubation is aided by a bare patch of skin on the lower abdomen (known as a 'brood pouch') that allows greater heat transfer to the egg. Once hatched, the male will remain to brood the chick for the first 25 days, whilst the female regularly brings food back to the nest. After this time, the chick is able to leave the nest, and will congregate with other chicks in small groups known as 'crèches' whilst the parent birds forage (2). In order to maintain its waterproof coat, the southern rockhopper penguin engages in frequent grooming, which helps to flatten the feathers and to spread a waxy substance that is secreted just below the tail. Grooming is also an important social bond between pairs. After breeding the southern rockhopper penguin forages extensively in order to build up fat reserves in preparation for its annual moult. It takes around 25 days for the penguin's coat to be fully replaced, at which point it leaves the land and spends the winter months foraging at sea, before returning to shore to breed in the following spring (2). The diet of the southern rockhopper penguin is composed of a variety of oceanic species, such as crustaceans, squid, octopus and fish (4). Groups may often feed together and dives may be to depths of up to 100 metres (2).
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Conservation

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Many islands that house breeding colonies have been designated as reserves and the populations in the Falklands, Marion, Campbell Islands are regularly monitored and studied (4). Greater investigation of population demographics and of potential threats is required. Following the starvation of over 100,000 rockhopper penguins in the Falkland Islands, the Spheniscus Penguin Conservation Work Group published a report recommending that commercial fishing be excluded within 30 miles of penguin breeding sites (2). These measures have been adopted around southern Chile and Argentina, and these sites are healthy and increasing as a result. The adjacent Falklands have refused to introduce such protection, and populations continue to decline (6).
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Description

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Previously classed as a single species, the rockhopper penguin has now been split into a northern (Eudyptes moseleyi) and southern species (Eudyptes chrysocome) (3). Although both species are similar in appearance, the distinctive yellowish plumes extending from the yellow line above the eye are significantly shorter and less dense in the southern rockhopper penguin (2) (3). The body is small but robust, with slate-grey upperparts and white underparts, the bill is short and reddish-brown and the eyes are red. Juveniles can be identified by the lack of adult yellow markings (2).
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Habitat

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Nesting occurs on cliffs and rocky gullies, and chosen sites are usually situated near to freshwater, either natural springs or puddles (2).
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Range

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The southern rockhopper penguin breeds on a number of Southern Ocean islands. Two subspecies are currently recognised, Eudyptes chrysocome chrysocome, which is found in the Falkland Islands, Isla Pinguino, Staten Island, and islands off southern Chile and Argentina, and Eudyptes chrysocome fiholi, which is found on several subantarctic islands to the south of New Zealand and South Africa (4).
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Status

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Classified as Vulnerable (VU) on the IUCN Red List (1).
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Threats

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Some southern rockhopper penguin nesting colonies have recently shown dramatic falls in numbers of breeding pairs. The Falkland Islands once housed the stronghold for southern rockhopper penguins, but over the last 60 years, numbers have declined by 90% (4). The reasons for these declines range from increasing disturbance and pollution, to declining fish stocks as a result of over fishing, failure to provide no-fishing zones around penguin colonies (6) and global warming (4).
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Southern rockhopper penguin

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The southern rockhopper penguin group[3] (Eudyptes chrysocome), is a species of rockhopper penguin, that is sometimes considered distinct from the northern rockhopper penguin. It occurs in subantarctic waters of the western Pacific and Indian Oceans, as well as around the southern coasts of South America.

Taxonomy

In 1743 the English naturalist George Edwards included an illustration and a description of the southern rockhopper penguin in the first volume of his A Natural History of Uncommon Birds. Edwards based his hand-coloured etching on a preserved specimen owned by Peter Collinson.[4] When in 1758 the Swedish naturalist Carl Linnaeus updated his Systema Naturae for the tenth edition, he placed the southern rockhopper penguin with the red-billed tropicbird in the genus Phaethon. Linnaeus included a brief description, coined the binomial name Phaethon demersus and cited Edwards' work.[5] The use of Linnaeus' binomial name was not adopted by later ornithologists, perhaps because he had already used the specific demersa for the African penguin which he placed with the wandering albatross in the genus Diomedea.[6][7]

The southern rockhopper penguin was formally described in 1781 by the German naturalist Johann Reinhold Forster under the binomial name Aptenodytes chrysocome.[8] The species is now placed in the genus Eudyptes that was introduced by the French ornithologist Louis Jean Pierre Vieillot in 1816.[9][10] The genus name combines the Ancient Greek eu meaning "fine" with dyptes meaning "diver". The specific epithet chrysocome is from the Ancient Greek khrusokomos meaning "golden-haired" (from khrusos meaning "gold" and komē meaning "hair").[11]

Two subspecies are recognised:[10]

The rockhopper penguin complex is confusing. Many taxonomists consider all three rockhopper penguin forms subspecies. Some split the northern subspecies (moseleyi) from the southern forms (chrysocome and filholi). Still others consider all three distinct. The subspecies recognized for the southern rockhopper penguin complex are:[2]

  • Eudyptes chrysocome chrysocome, the western rockhopper penguin or American southern rockhopper penguin – breeds around the southern tip of South America.
  • Eudyptes chrysocome filholi, the eastern rockhopper penguin or Indopacific southern rockhopper penguin – breeds on subantarctic islands of the Indian and western Pacific oceans.

The northern rockhopper penguin lives in a different water mass than the western and eastern rockhopper penguin, separated by the Subtropical Front, and they are genetically different. Therefore, northern birds are sometimes separated as E. moseleyi. The rockhopper penguins are closely related to the macaroni penguin (E. chrysolophus) and the royal penguin (E. schlegeli), which may just be a colour morph of the macaroni penguin.

Interbreeding with the macaroni penguin has been reported at Heard and Marion Islands, with three hybrids recorded there by a 1987–88 Australian National Antarctic Research Expedition.[12]

Description

Southern rockhopper penguin(Eudyptes chrysocome) displaying its distinctive crest

This is the smallest yellow-crested, black-and-white penguin in the genus Eudyptes. It reaches a length of 45–58 cm (18–23 in) and typically weighs 2–3.4 kg (4.4–7.5 lb), although there are records of exceptionally large rockhoppers weighing 4.5 kg (9.9 lb).[13] It has slate-grey upper parts and has straight, bright yellow eyebrows ending in long yellowish plumes projecting sideways behind a red eye.[13]

Ecology

Eudyptes chrysocome chrysocome colony on Saunders Island, Falkland Islands

The southern rockhopper penguin group has a global population of roughly 1 million pairs. About two-thirds of the global population belongs to E. c. chrysocome which breeds on the Falkland Islands and on islands off Patagonia.[14] These include most significantly Isla de los Estados, the Ildefonso Islands, the Diego Ramírez Islands and Isla Noir. E. c. filholi breeds on the Prince Edward Islands, the Crozet Islands, the Kerguelen Islands, Heard Island, Macquarie Island, Campbell Island, the Auckland Islands and the Antipodes Islands. Outside the breeding season, the birds can be found roaming the waters offshore their colonies.[15]

These penguins feed on krill, squid, octopus, lantern fish, mollusks, plankton, cuttlefish, and mainly crustaceans.

A rockhopper penguin, named Rocky, in Bergen Aquarium in Norway, lived to 29 years 4 months. It died in October 2003. This stands as the age record for rockhopper penguins, and possibly it was the oldest penguin known.[16]

Behaviour

E. c. chrysocome on Saunders Island, Falkland Islands hopping over a crack

Their common name refers to the fact that, unlike many other penguins which get around obstacles by sliding on their bellies or by awkward climbing using their flipper-like wings as aid, rockhoppers will try to jump over boulders and across cracks.[13]

This behaviour is by no means unique to this species however – at least the other "crested" penguins of the genus Eudyptes hop around rocks too. But the rockhopper's congeners occur on remote islands in the New Zealand region, whereas the rockhopper penguins are found in places that were visited by explorers and whalers since the Early Modern era. Hence, it is this particular species in which this behaviour was first noted.

Egg
A chick on the Saunders Island in the Falkland Islands

Their breeding colonies are located from sea level to cliff-tops and sometimes inland. Their breeding season starts in September and ends in November.[13] Two eggs are laid but only one is usually incubated.[13] Incubation lasts 35 days and their chicks are brooded for 26 days.

Variation in foraging behaviour

Rockhopper penguins are known to have complex foraging behaviors. Influenced by factors such as sea ice abundance, prey availability, breeding stage, and seasonality, rockhopper penguins must be able to adapt their behavior to fit the current conditions.[17] Rockhopper penguins employ different strategies according to their conditions. When making foraging trips, rockhoppers typically leave and return to their colonies in groups. One study showed they are known for going up to 157 km away from their colonies when foraging.[18] Females typically forage during the day in 11-12h trips consisting of many dives, but they will occasionally forage at night.[17] Night dives are typically much shallower than day dives. Dives typically last around 12h, but can be up to 15hrs, with penguins leaving the colony around dawn (04:00) and returning at dusk (19:00).[19]

Rockhopper penguin skeleton in Manchester Museum

Rockhopper penguins employ different strategies and foraging behaviors depending on the climate and environment. A main factor is food location. Subantarctic penguins must dive for longer periods of time and much deeper in search of food than do species in warmer waters where food is more easily accessible.[17]

Benthic and pelagic dives

Rockhopper penguins are known to employ two different types of dives when foraging, pelagic and benthic dives. Pelagic dives are typically short and relatively shallow and used very frequently. Benthic dives are much deeper dives near the seafloor (up to 100 m deep)[19] that typically last longer and have longer bottom time. Penguins performing benthic dives typically only perform a few depth wiggles (changes in depth profile) at their maximum depth.[20] at an average speed of range of 6.9–8.1 kilometres per hour (6,900–8,100 m/h).[18] Although deeper dives tend to be a bit longer than shallow dives, foraging rockhoppers will minimize their travel time when performing benthic dives to gain maximum efficiency. Benthic dives in particular show a stronger correlation to full stomachs than pelagic dives. Emperor penguins, gentoo penguins, yellow-eyed penguins and king penguins also use this deep-dive technique to obtain food.[20]

Prey availability is dependent on many factors, such as current climate and conditions of the area. Typically, females will bring back a majority of crustaceans and occasionally some fish for their young. The female's foraging success directly affects chick growth.[17] If food is scarce, females are able to fast for very long periods of time and sometimes will only forage for the chick's benefit.[19]

Dive limitations

Because foraging conditions and outcomes are so variable, several factors can limit foraging practices. The timing of breeding, incubation and brooding periods greatly affect foraging time, as females are unable to leave broods for long periods of time.[18] Females during the brooding period will follow a much more fixed foraging schedule, leaving and returning to the colony at roughly the same time each day. When not in breeding season, females have much more variability in the length of foraging trips. If females have low energy levels because they are fasting while provisioning chicks, they may make several short foraging trips instead of one longer one.[17]

While benthic dives are efficient and favorable for rockhoppers, they present physiological limitations such as limits in lung capacity, which affects duration of dives. The longest aerobic dive rockhoppers can perform is about 110 seconds long,[20] but dives can last upwards of 180–190 seconds.[17][18]

Status and conservation

The southern rockhopper penguin group is classified as vulnerable by the IUCN.[2] Its population has declined by about one-third in the last thirty years.[15][21] This decline has earned them the classification of a vulnerable species by the IUCN. Threats to their population include commercial fishing and oil spills.[22]

With the approval of the European Association of Zoos and Aquaria (EAZA), Drusillas Park in East Sussex holds the studbook for rockhopper penguins in Europe. Zoo manager Sue Woodgate has specialist knowledge of the species, so the zoo is responsible for co-ordinating the movements of penguins within zoos in Europe to take part in breeding programmes and offer their advice and information about the species.[23]

Relationship with humans

Adult E. c. chrysocome in the New Island (Falkland Islands) rookery

Rockhopper penguins are the most familiar of the crested penguins to the general public. Their breeding colonies, namely those around South America, today attract many tourists who enjoy watching the birds' antics. Historically, the same islands were popular stopover and replenishing sites for whalers and other seafarers since at least the early 18th century. Almost all crested penguins depicted in movies, books and other media are ultimately based on Eudyptes chrysocome chrysocome.

References

  1. ^ "Fossilworks: Eudyptes chrysocome".
  2. ^ a b c BirdLife International (2020). "Eudyptes chrysocome". IUCN Red List of Threatened Species. 2020: e.T22735250A182762377. doi:10.2305/IUCN.UK.2020-3.RLTS.T22735250A182762377.en. Retrieved February 20, 2022.
  3. ^ English name updates – IOC Version 2.9 (July 10, 2011) Archived November 7, 2011, at the Wayback Machine, IOC World Bird List
  4. ^ Edwards, George (1751). A Natural History of Uncommon Birds. Vol. Part 1. London: Printed for the author at the College of Physicians. p. 49, Plate 49.
  5. ^ Linnaeus, Carl (1758). Systema Naturae per regna tria naturae, secundum classes, ordines, genera, species, cum characteribus, differentiis, synonymis, locis (in Latin). Vol. 1 (10th ed.). Holmiae (Stockholm): Laurentii Salvii. p. 135.
  6. ^ Linnaeus, Carl (1758). Systema Naturae per regna tria naturae, secundum classes, ordines, genera, species, cum characteribus, differentiis, synonymis, locis (in Latin). Vol. 1 (10th ed.). Holmiae (Stockholm): Laurentii Salvii. p. 132.
  7. ^ Allen, J.A. (1904). "The case of Megalestris vs. Catharacta". The Auk. 21 (3): 345–348 [346, Footnote 1]. doi:10.2307/4070197. hdl:2027/hvd.32044107327124. JSTOR 4070197.
  8. ^ Forster, Johann Reinhold (1780). "Historia Aptenodytae. Generis Avium orbi Australi proprii". Commentationes Societatis Regiae Scientiarum Gottingensis (in Latin). 3: 121–148 [133, 135]. Although the volume is dated 1780, the article was not published until 1781.
  9. ^ Vieillot, Louis Jean Pierre (1816). Analyse d'une Nouvelle Ornithologie Élémentaire (in French). Paris: Deterville/self. pp. 67, 70. The genus name is misspelled as Endyptes on page 67.
  10. ^ a b Gill, Frank; Donsker, David; Rasmussen, Pamela, eds. (July 2021). "Kagu, Sunbittern, tropicbirds, loons, penguins". IOC World Bird List Version 11.2. International Ornithologists' Union. Retrieved November 1, 2021.
  11. ^ Jobling, James A. (2010). The Helm Dictionary of Scientific Bird Names. London: Christopher Helm. pp. 152, 105. ISBN 978-1-4081-2501-4.
  12. ^ Woehler, E. J.; Gilbert, C. A. (1990). "Hybrid Rockhopper-Macaroni Penguins, interbreeding and mixed-species pairs at Heard and Marion Islands". Emu. 90 (3): 198–210. doi:10.1071/MU9900198.
  13. ^ a b c d e Trewby, Mary (2002). Antarctica: an encyclopedia from Abbot Ice Shelf to Zooplankton. Auckland, New Zealand: Firefly Books. p. 152. ISBN 978-1-55297-590-9.
  14. ^ "Animals at Drusillas Zoological Park - Best Small UK Zoo for Kids". www.drusillas.co.uk. Retrieved February 18, 2023.
  15. ^ a b BirdLife International (2008). [2008 IUCN Redlist status changes]. Retrieved May 23, 2008.
  16. ^ Glenday, Craig (ed.) (2008). Guinness World Records 2008. Guinness Media, Inc. ISBN 1-904994-19-9
  17. ^ a b c d e f Tremblay, Yann; Cherel, Yves (2003). "Geographic variation in the foraging behaviour, diet and chick growth of rockhopper penguins". Marine Ecology Progress Series. 251: 279–297. Bibcode:2003MEPS..251..279T. doi:10.3354/meps251279.
  18. ^ a b c d Brown, Christopher (1987). "Traveling Speed and Foraging Range of Macaroni and Rockhopper Penguins at Marion Island (Velocidad de Movimiento y Extensión de las Áreas de Forrajeo de los Pingüinos Eudyptes chrysolophus y e. Chrysocome)". Journal of Field Ornithology. 58 (2): 118–125. JSTOR 4513209.
  19. ^ a b c Putz, Klemens (November 29, 2005). "Diving characteristics of southern rockhopper penguins (Eudyptes c. chrysocome) in the southwest Atlantic". Marine Biology. 149 (2): 125–137. doi:10.1007/s00227-005-0179-y. S2CID 84393587.
  20. ^ a b c Tremblay, Yann; Cherel, Yves (2000). "Benthic and pelagic dives: a new foraging behaviour in rockhopper penguins". Marine Ecology Progress Series. 204: 257–267. Bibcode:2000MEPS..204..257T. doi:10.3354/meps204257.
  21. ^ BirdLife International (2008) Southern Rockhopper Penguin Species Factsheet Archived December 11, 2008, at the Wayback Machine. Retrieved May 27, 2008.
  22. ^ Devon Phelan. "Eudyptes chrysocome rockhopper penguin". Animal Diversity Web. Retrieved July 9, 2013.
  23. ^ "Conservation at Drusillas Zoo Park – Kids Days Out With Animals". www.drusillas.co.uk. Retrieved February 18, 2023.

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Southern rockhopper penguin: Brief Summary

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The southern rockhopper penguin group (Eudyptes chrysocome), is a species of rockhopper penguin, that is sometimes considered distinct from the northern rockhopper penguin. It occurs in subantarctic waters of the western Pacific and Indian Oceans, as well as around the southern coasts of South America.

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Breeding Category

provided by World Register of Marine Species
Vagrant

Reference

Woehler E.J. (compiler) 2006. Species list prepared for SCAR/IUCN/BirdLife International Workshop on Antarctic Regional Seabird Populations, March 2005, Cambridge, UK.

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Charbonneau, Danny [email]

Description

provided by World Register of Marine Species
Length: 66-76 cm. Plumage: adult with head, throat, back, wings, tail and undertail coverts black, breast and belly white. Yellow line above eye not meeting on forehead, but ending in long, pale yellow plumes behind each eye. Immature with a line of cream feathers above eye and replacing plumes behind eye. Bare parts: iris red; bill pink to orange-red; feet dull pink with darker webs and black soles.

Reference

Brown, L.H., E.K. Urban & K. Newman. (1982). The Birds of Africa, Volume I. Academic Press, London.

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Edward Vanden Berghe [email]

IUCN Red List Category

provided by World Register of Marine Species
Vulnerable

Reference

Woehler E.J. (compiler) 2006. Species list prepared for SCAR/IUCN/BirdLife International Workshop on Antarctic Regional Seabird Populations, March 2005, Cambridge, UK.

license
cc-by-4.0
copyright
WoRMS Editorial Board
contributor
Charbonneau, Danny [email]