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Image of Orthogonius taghavianae Tian
Creatures » … » Animal » … » Arthropods » … » Hexapods » Insects » Winged Insects » … » Endopterygotes » Beetles » … » Ground Beetles

Orthogonius

Scientific Names

Preferred Names

Orthogonius
Genus recognized by EOL Dynamic Hierarchy April 2022
Reference taxon from Catalogue of Life in Species 2000 & ITIS Catalogue of Life
Orthogonius
Genus recognized by wikipedia NL, wikipedia POL, EOL Dynamic Hierarchy 1.1, EOL Dynamic Hierarchy 1.1, South Africa Species List, Zimbabwe Species List, wikipedia ID, Gulf of Guinea Species List, Indian Ocean Species List, NCBI, wikidata_hierarchy tar gz, wikipedia EN, Wikipedia (inferred records), and Dynamic Hierarchy Version 2.2 - Test
Orthogonius
Recognized by OBIS environmental data records
Orthogonius Macleay 1825
Genus recognized by GBIF classification
Orthogonius W. S. MacLeay 1825
Genus recognized by Catalogue of Life 7 February 2023

Alternative Names

Orthogonius dupuisi Tian & Deuve 2006
Heterotypic synonym according to GBIF classification
Orthogonius rhodesianus Basilewsky 1948
Heterotypic synonym according to GBIF classification
Apsectra Schmidt-Goebel 1846
Synonym according to Catalogue of Life 7 February 2023
Haplopisthius Chaudoir 1850
Synonym according to Catalogue of Life 7 February 2023
Maraga Walker 1858
Synonym according to Catalogue of Life 7 February 2023
Orthogonius
Harvest ancestor according to GBIF national node type records UK, Mozambique Species List, GBIF nat'l node type records Netherlands, and TreatmentBank

Common Names

There are no common names associated with this taxon.

Curated hierarchies for Orthogonius

EOL Dynamic Hierarchy April 2022

Life
Cellular Organisms
Eukaryota
Opisthokonta
Metazoa
Bilateria
Protostomia
Ecdysozoa
Arthropoda
Pancrustacea
Hexapoda
Insecta
Pterygota
Neoptera
Endopterygota
Coleoptera
Adephaga
Carabidae
Orthogonius (this page)
Orthogonius aberlenci Tian & Deuve 2016
Orthogonius acrogonus (Wiedemann 1819)
Orthogonius acutangulus Chaudoir 1878
Orthogonius adoriae Tian & Deuve 2006
Orthogonius aemulus Péringuey 1896
Orthogonius alternans (Wiedemann 1823)
Orthogonius alutaceus Quedenfeldt 1883
Orthogonius andrewesi van Emden 1928
Orthogonius andrewesianus Tian & Deuve 2006
Orthogonius angkor Tian & Deuve 2006
Orthogonius angkorensis Tian & Deuve 2008
Orthogonius angustus Chaudoir 1872
Orthogonius annamensis Jedlicka 1965
Orthogonius annamicola Tian & Deuve 2006
Orthogonius annemariae Tian & Deuve 2006
Orthogonius assamensis Tian & Deuve 2006
Orthogonius assamicola Tian & Deuve 2006
Orthogonius assmuthi Wasmann 1920
Orthogonius baconi Chaudoir 1872
Orthogonius baconioides Tian & Sabu 2009
Orthogonius balthasari Jedlicka 1935
Orthogonius batesi Tian & Deuve 2003
Orthogonius batesianus Tian & Deuve 2007
Orthogonius batesioides Tian & Deuve 2006
Orthogonius baumi Jedlicka 1955
Orthogonius becqueti Burgeon 1937
Orthogonius bellus Tian & Deuve 2016
Orthogonius bicolor Jedlicka 1965
Orthogonius borneoensis Tian & Deuve 2006
Orthogonius brancuccii Tian & Deuve 2006
Orthogonius brevilabris H. Kolbe 1889
Orthogonius brevithorax Dejean 1825
Orthogonius bruggei Tian & Deuve 2006
Orthogonius buquetii Chaudoir 1850
Orthogonius burckhardti Tian & Deuve 2006
Orthogonius burmanensis Tian & Deuve 2006
Orthogonius caffer Boheman 1848
Orthogonius cambodgensis Tian & Deuve 2006
Orthogonius canaraensis Tian & Deuve 2006
Orthogonius canaraicola Tian & Deuve 2006
Orthogonius capucinus Boheman 1848
Orthogonius carbonicolor Tian & Deuve 2006
Orthogonius carinatus Tian & Deuve 2013
Orthogonius celebesicus Tian & Deuve 2006
Orthogonius ceylanicus Tian & Deuve 2016
Orthogonius chaudoiri Tian & Deuve 2001
Orthogonius cheni Tian & Deuve 2001
Orthogonius chiangensis Tian & Deuve 2006
Orthogonius clarkii Murray 1858
Orthogonius confusus Tian & Deuve 2006
Orthogonius constanti Tian & Deuve 2006
Orthogonius constrictus Tian
Orthogonius coomani Tian & Deuve 2006
Orthogonius coomanioides Tian
Orthogonius coracinus H. Kolbe 1896
Orthogonius crassicrus Chaudoir 1872
Orthogonius cruralis Putzeys ex Chaudoir 1872
Orthogonius curvatus Tian & Deuve 2006
Orthogonius cyclothorax Tian & Deuve 2007
Orthogonius davidi Chaudoir 1878
Orthogonius deletus Schmidt-Goebel 1846
Orthogonius deuvei Tian & Kirschenhofer 2013
Orthogonius dispar Bates 1892
Orthogonius dohertyi Tian & Deuve 2006
Orthogonius dongnanya Tian & Deuve 2006
Orthogonius doriae Putzeys ex Chaudoir 1872
Orthogonius drescheri Liebke 1937
Orthogonius drumonti Tian & Deuve 2006
Orthogonius duboisi Tian & Deuve 2006
Orthogonius duplicatus (Wiedemann 1819)
Orthogonius dupuisi Basilewsky 1948
Orthogonius dupuisianus Tian & Deuve 2010
Orthogonius dureli Tian & Deuve 2005
Orthogonius edentatus Tian & Deuve 2006
Orthogonius elisabethanus Burgeon 1937
Orthogonius emarginatus Tian & Deuve 2007
Orthogonius equimarginalis Tian & Deuve 2006
Orthogonius euthyphallus Tian & Deuve 2013
Orthogonius fairmairei Tian & Deuve 2006
Orthogonius feai Tian & Deuve 2007
Orthogonius femoralis Chaudoir 1848
Orthogonius flavipes Deuve 2004
Orthogonius flavus Jedlicka 1964
Orthogonius foveiclypeus Tian & Deuve 2004
Orthogonius fryi Tian & Deuve 2006
Orthogonius fugax Chaudoir 1872
Orthogonius gracililamella Tian
Orthogonius gracilipes Tian & Deuve 2006
Orthogonius gracilis Tian & Deuve 2006
Orthogonius grootaerti Tian & Deuve 2006
Orthogonius hageni Oberthür 1883
Orthogonius himalaya Tian & Deuve 2006
Orthogonius himalayicus Tian & Deuve 2005
Orthogonius hirtulus Tian & Deuve 2006
Orthogonius hirtus Chaudoir 1872
Orthogonius hopei Gray 1832
Orthogonius huananoides Tian & Deuve 2006
Orthogonius huananus Tian & Deuve 2001
Orthogonius hypocrita Chaudoir 1872
Orthogonius hypocritoides Jedlicka 1935
Orthogonius impunctipennis Quedenfeldt 1883
Orthogonius indicus Tian & Deuve 2006
Orthogonius indophilus Tian & Deuve 2006
Orthogonius inexpectatus Tian & Deuve 2006
Orthogonius inops Chaudoir 1872
Orthogonius insularis Chaudoir 1872
Orthogonius intermedius Chaudoir 1872
Orthogonius javanus Tian & Deuve 2006
Orthogonius jianfengling Tian & Deuve 2001
Orthogonius kalimantanensis Tian & Deuve 2006
Orthogonius kapiriensis Burgeon 1937
Orthogonius katangensis Burgeon 1937
Orthogonius kickeli H. Kolbe 1896
Orthogonius kirirom Tian & Deuve 2008
Orthogonius klickai Jedlicka 1935
Orthogonius kubani Tian & Deuve 2006
Orthogonius kumatai Habu 1979
Orthogonius lancangjiang Tian & Deuve 2006
Orthogonius lao Tian & Deuve 2006
Orthogonius latreillei Tian & Deuve 2006
Orthogonius latus Hope 1842
Orthogonius ledouxi Tian & Deuve 2006
Orthogonius legrandi Tian & Deuve 2006
Orthogonius lieftincki Andrewes 1936
Orthogonius limbourgi Tian & Deuve 2016
Orthogonius loeicus Tian & Deuve 2006
Orthogonius loeiensis Tian & Deuve 2006
Orthogonius longicornis Chaudoir 1872
Orthogonius longilamella Tian & Deuve 2008
Orthogonius longipennis Hope 1842
Orthogonius longiphallus Tian & Deuve 2005
Orthogonius lucidus Bates 1891
Orthogonius luzonicus Chaudoir 1872
Orthogonius macrophthalmus Tian & Deuve 2013
Orthogonius maindronianus Tian & Deuve 2006
Orthogonius makiling Tian & Deuve 2006
Orthogonius malacca Tian & Deuve 2006
Orthogonius malaisei Andrewes 1947
Orthogonius malaya Tian & Deuve 2006
Orthogonius malaysiaensis Tian & Deuve 2006
Orthogonius medanensis Tian & Deuve 2006
Orthogonius meghalayaensis Tian & Deuve 2016
Orthogonius mekongensis Tian & Deuve 2006
Orthogonius melanipes Tian & Deuve 2006
Orthogonius mellyi (Chaudoir 1850)
Orthogonius minimus Tian & Deuve 2006
Orthogonius mniszechi Chaudoir 1872
Orthogonius moestus Chaudoir 1872
Orthogonius monolophus Andrewes 1931
Orthogonius montanus Tian & Deuve 2006
Orthogonius morvani Tian & Deuve 2003
Orthogonius morvanianus Tian & Deuve 2016
Orthogonius mouhoti Chaudoir 1872
Orthogonius myanmarensis Tian & Deuve 2006
Orthogonius nahaeo Tian & Deuve 2006
Orthogonius nepalensis Habu 1979
Orthogonius nepalicus Tian & Deuve 2006
Orthogonius niger Jedlicka 1935
Orthogonius nigripes Tian & Deuve 2001
Orthogonius nongfaensis Tian & Kirschenhofer 2013
Orthogonius nyassicus H. Kolbe 1896
Orthogonius obliquepes Tian & Deuve 2006
Orthogonius opacus Schmidt-Goebel 1846
Orthogonius orphnodes Andrewes 1930
Orthogonius ovatulus Tian & Deuve 2003
Orthogonius ovatus Chaudoir 1878
Orthogonius pachlatkoi Tian & Deuve 2006
Orthogonius palangbangensis Tian & Deuve 2006
Orthogonius panghongae Tian & Deuve 2006
Orthogonius pangi Tian & Deuve 2006
Orthogonius parallelus Chaudoir 1872
Orthogonius parastantoni Tian & Deuve 2006
Orthogonius parasuturalis Tian & Deuve 2006
Orthogonius parcepunctatus H. Kolbe 1896
Orthogonius paris Tian & Deuve 2006
Orthogonius parvus Chaudoir 1872
Orthogonius pectinatus Tian & Deuve 2016
Orthogonius perakensis Tian & Deuve 2006
Orthogonius perakicus Tian & Deuve 2007
Orthogonius perpuncticollis Burgeon 1937
Orthogonius petiolaris Tian & Deuve 2006
Orthogonius piceus Chaudoir 1872
Orthogonius picilabris W. S. MacLeay 1825
Orthogonius picipennis Chaudoir 1872
Orthogonius pinguis Murray 1858
Orthogonius pinophilus Tian
Orthogonius planiger (Walker 1858)
Orthogonius plebius Tian & Deuve 2006
Orthogonius poggii Tian & Deuve 2007
Orthogonius politior Tian & Deuve 2016
Orthogonius politus Chaudoir 1872
Orthogonius pollinctor Basilewsky 1948
Orthogonius pradieri Chaudoir 1872
Orthogonius pseudochaudoiri Tian
Orthogonius pseudocheni Tian & Deuve 2006
Orthogonius pseudolongicornis Tian & Deuve 2006
Orthogonius pseudomouhoti Tian & Deuve 2008
Orthogonius punctatellus Tian & Deuve 2006
Orthogonius punctulatus Chaudoir 1872
Orthogonius punctum Tian & Deuve 2016
Orthogonius putzeysi Tian & Deuve 2006
Orthogonius quadrisetosus Tian & Deuve 2006
Orthogonius rhodesianus Csiki 1932
Orthogonius rhodesiensis Lorenz 1998
Orthogonius ribbei Tian & Deuve 2003
Orthogonius rogueti Tian & Deuve 2016
Orthogonius rotundatus Tian & Deuve 2006
Orthogonius rufiventris Bates 1892
Orthogonius rufotestaceus G. Müller 1941
Orthogonius sabahensis Tian & Deuve 2006
Orthogonius sabahicus Tian & Deuve 2016
Orthogonius salvazai Tian & Deuve 2006
Orthogonius sarawakensis Tian & Deuve 2006
Orthogonius saundersi Andrewes 1926
Orthogonius schaumi Chaudoir 1872
Orthogonius schaumioides Tian & Deuve 2006
Orthogonius schauteni Tian & Deuve 2006
Orthogonius schmidtgoebeli Chaudoir 1872
Orthogonius senegalensis Dejean 1831
Orthogonius setosopalpiger Tian
Orthogonius siamensis Tian & Deuve 2006
Orthogonius similaris Tian
Orthogonius simplicatus Tian & Deuve 2006
Orthogonius singaporensis Tian & Deuve 2006
Orthogonius sinuatiphallus Tian & Deuve 2001
Orthogonius smetsi Tian & Deuve 2006
Orthogonius solidicornis Tian & Deuve 2003
Orthogonius solidus Tian & Deuve 2006
Orthogonius spinosus Burgeon 1937
Orthogonius srilankaicus Tian & Deuve 2003
Orthogonius stantoni Tian & Deuve 2006
Orthogonius sterbai Jedlicka 1935
Orthogonius stygius Andrewes 1930
Orthogonius sulawesicus Tian & Deuve 2003
Orthogonius sulcatoides Tian & Deuve 2006
Orthogonius sulcatus Schmidt-Goebel 1846
Orthogonius sumatraicus Tian & Deuve 2006
Orthogonius suturalis Chaudoir 1872
Orthogonius taghavianae Tian
Orthogonius taiwanicus Tian & Deuve 2006
Orthogonius tangsen Tian & Deuve 2006
Orthogonius tenasserimensis Tian & Deuve 2006
Orthogonius termiticola Wasmann 1902
Orthogonius thaicus Tian & Deuve 2003
Orthogonius thaiensis Tian & Deuve 2006
Orthogonius thailandensis Tian & Deuve 2006
Orthogonius thoracicus Gestro 1875
Orthogonius tikekee Tian & Deuve 2006
Orthogonius tonkinensis Tian & Deuve 2006
Orthogonius tonkinicus Tian & Deuve 2006
Orthogonius travancore Tian & Deuve 2006
Orthogonius tricarinatus Burgeon 1937
Orthogonius uncipennis Tian & Deuve 2006
Orthogonius vari Tian
Orthogonius variabilis Tian
Orthogonius virgulatus Andrewes 1931
Orthogonius wallardiensis Tian & Deuve 2006
Orthogonius wrasei Tian & Deuve 2016
Orthogonius xanthomerus L. Redtenbacher 1867
Orthogonius yoga Tian & Deuve 2006
Orthogonius yunnanensis Tian & Deuve 2001
Aepiblemus
Allendia
Allocota
Amaroschema
Amblystomus
Anillopsidius
Anophthalmus
Antennaria
Archipatrobus
Aristochroa
Aristolebia
Assadecma
Atlantomasoreus
Bennigsenium
Bhutanotrechus
Blenniventer
Bradycidus
Cascellius
Chaetauchenium
Chaetocrepis
Chlaeminus
Cratoferonia
Deipyrodes
Demetrias
Diceromerus
Dimorphoreicheia
Dyschirius
Erephognathus
Eripus
Eutogeneius
Habrodera
Hygranillus
Hystricosphodrus
Jeannelius
Kalchdigor
Lachnolebia
Leptognatha
Liagonum
Liocolpodes
Lophyra
Marsyas
Masoreus
Megacephala
Megadromus
Mexisphodrus
Mimotrechus
Minosaphaenops
Monodryxus
Nannoryctes
Neohelluo
Neohyparpalus
Nesarpalus
Notonomus
Ocys
Oedesis
Oxycrepis
Oxygnathopsis
Oxytrechus
Pachycarus
Paracolpodes
Paraliagonum
Pasimachus
Paulianites
Pelecium
Phaeoxantha
Philipis
Physolaesthus
Platypatrobus
Platysphyrus
Pogonoglossus
Pometon
Potamagonum
Procletus
Prothymidia
Protopaussus
Pseudobatenus
Rathymus
Rawlinsius
Rhytiferonia
Ryugadous
Sericoda
Siagonyx
Singilis
Sinocimmerites
Sinometrius
Sloanoglymmius
Smeringocera
Sphaerodinus
Steganomma
Stricklandiana
Tachys
Taridius
Tasmanillus
Trechiamiotes
Trechosia
Trichocarenum
Trichoxycentrus
Typhlochoromus
Xenodochus
Yukihikous
1984 additional siblings truncated for brevity.
See the resource file for a full list.

Catalogue of Life 7 February 2023

Biota
Animalia
Arthropoda
Insecta
Coleoptera
Caraboidea
Carabidae Latreille 1802
Orthogoniinae Schaum 1857
Orthogoniini Schaum 1857
Orthogoniina Schaum 1857
Orthogonius W. S. MacLeay 1825 (this page)
Orthogonius aberlenci Tian & Deuve 2016
Orthogonius acrogonus (Wiedemann 1819)
Orthogonius acutangulus Chaudoir 1878
Orthogonius adoriae Tian & Deuve 2006
Orthogonius aemulus Péringuey 1896
Orthogonius alternans (Wiedemann 1823)
Orthogonius alutaceus Quedenfeldt 1883
Orthogonius andrewesi Emden 1928
Orthogonius andrewesianus Tian & Deuve 2006
Orthogonius angkor Tian & Deuve 2006
Orthogonius angkorensis Tian & Deuve 2008
Orthogonius angustus Chaudoir 1872
Orthogonius annamensis Jedlicka 1965
Orthogonius annamicola Tian & Deuve 2006
Orthogonius annemariae Tian & Deuve 2006
Orthogonius assamensis Tian & Deuve 2006
Orthogonius assamicola Tian & Deuve 2006
Orthogonius assmuthi Wasmann 1920
Orthogonius baconi Chaudoir 1872
Orthogonius baconioides Tian & Sabu 2009
Orthogonius balthasari Jedlicka 1935
Orthogonius batesi Tian & Deuve 2003
Orthogonius batesianus Tian & Deuve 2007
Orthogonius batesioides Tian & Deuve 2006
Orthogonius baumi Jedlicka 1955
Orthogonius becqueti Burgeon 1937
Orthogonius bellus Tian & Deuve 2016
Orthogonius bicolor Jedlicka 1965
Orthogonius borneoensis Tian & Deuve 2006
Orthogonius brancuccii Tian & Deuve 2006
Orthogonius brevilabris Kolbe 1889
Orthogonius brevithorax Dejean 1825
Orthogonius bruggei Tian & Deuve 2006
Orthogonius buquetii Chaudoir 1850
Orthogonius burckhardti Tian & Deuve 2006
Orthogonius burmanensis Tian & Deuve 2006
Orthogonius caffer Boheman 1848
Orthogonius cambodgensis Tian & Deuve 2006
Orthogonius canaraensis Tian & Deuve 2006
Orthogonius canaraicola Tian & Deuve 2006
Orthogonius capucinus Boheman 1848
Orthogonius carbonicolor Tian & Deuve 2006
Orthogonius carinatus Tian & Deuve 2013
Orthogonius celebesicus Tian & Deuve 2006
Orthogonius ceylanicus Tian & Deuve 2016
Orthogonius chaudoiri Tian & Deuve 2001
Orthogonius cheni Tian & Deuve 2001
Orthogonius chiangensis Tian & Deuve 2006
Orthogonius clarkii Murray 1858
Orthogonius confusus Tian & Deuve 2006
Orthogonius constanti Tian & Deuve 2006
Orthogonius constrictus Tian
Orthogonius coomani Tian & Deuve 2006
Orthogonius coomanioides Tian
Orthogonius coracinus Kolbe 1895
Orthogonius crassicrus Chaudoir 1872
Orthogonius cruralis Putzeys ex Chaudoir 1872
Orthogonius curvatus Tian & Deuve 2006
Orthogonius cyclothorax Tian & Deuve 2007
Orthogonius davidi Chaudoir 1878
Orthogonius deletus Schmidt-Goebel 1846
Orthogonius deuvei Tian & Kirschenhofer 2013
Orthogonius dispar Bates 1892
Orthogonius dohertyi Tian & Deuve 2006
Orthogonius dongnanya Tian & Deuve 2006
Orthogonius doriae Putzeys ex Chaudoir 1872
Orthogonius drescheri Liebke 1937
Orthogonius drumonti Tian & Deuve 2006
Orthogonius duboisi Tian & Deuve 2006
Orthogonius duplicatus (Wiedemann 1819)
Orthogonius dupuisi Basilewsky 1948
Orthogonius dupuisianus Tian & Deuve 2010
Orthogonius dureli Tian & Deuve 2005
Orthogonius edentatus Tian & Deuve 2006
Orthogonius elisabethanus Burgeon 1937
Orthogonius emarginatus Tian & Deuve 2007
Orthogonius equimarginalis Tian & Deuve 2006
Orthogonius euthyphallus Tian & Deuve 2013
Orthogonius fairmairei Tian & Deuve 2006
Orthogonius feai Tian & Deuve 2007
Orthogonius femoralis Chaudoir 1848
Orthogonius flavipes Deuve 2004
Orthogonius flavus Jedlicka 1964
Orthogonius foveiclypeus Tian & Deuve 2004
Orthogonius fryi Tian & Deuve 2006
Orthogonius fugax Chaudoir 1872
Orthogonius gracililamella Tian
Orthogonius gracilipes Tian & Deuve 2006
Orthogonius gracilis Tian & Deuve 2006
Orthogonius grootaerti Tian & Deuve 2006
Orthogonius hageni Oberthür 1883
Orthogonius himalaya Tian & Deuve 2006
Orthogonius himalayicus Tian & Deuve 2005
Orthogonius hirtulus Tian & Deuve 2006
Orthogonius hirtus Chaudoir 1872
Orthogonius hopei Gray 1832
Orthogonius huananoides Tian & Deuve 2006
Orthogonius huananus Tian & Deuve 2001
Orthogonius hypocrita Chaudoir 1872
Orthogonius hypocritoides Jedlicka 1935
Orthogonius impunctipennis Quedenfeldt 1883
Orthogonius indicus Tian & Deuve 2006
Orthogonius indophilus Tian & Deuve 2006
Orthogonius inexpectatus Tian & Deuve 2006
Orthogonius inops Chaudoir 1872
Orthogonius insularis Chaudoir 1872
Orthogonius intermedius Chaudoir 1872
Orthogonius javanus Tian & Deuve 2006
Orthogonius jianfengling Tian & Deuve 2001
Orthogonius kalimantanensis Tian & Deuve 2006
Orthogonius kapiriensis Burgeon 1937
Orthogonius katangensis Burgeon 1937
Orthogonius kickeli Kolbe 1895
Orthogonius kirirom Tian & Deuve 2008
Orthogonius klickai Jedlicka 1935
Orthogonius kubani Tian & Deuve 2006
Orthogonius kumatai Habu 1979
Orthogonius lancangjiang Tian & Deuve 2006
Orthogonius lao Tian & Deuve 2006
Orthogonius latreillei Tian & Deuve 2006
Orthogonius latus Hope 1842
Orthogonius ledouxi Tian & Deuve 2006
Orthogonius legrandi Tian & Deuve 2006
Orthogonius lieftincki Andrewes 1936
Orthogonius limbourgi Tian & Deuve 2016
Orthogonius loeicus Tian & Deuve 2006
Orthogonius loeiensis Tian & Deuve 2006
Orthogonius longicornis Chaudoir 1872
Orthogonius longilamella Tian & Deuve 2008
Orthogonius longipennis Hope 1842
Orthogonius longiphallus Tian & Deuve 2005
Orthogonius lucidus Bates 1891
Orthogonius luzonicus Chaudoir 1872
Orthogonius macrophthalmus Tian & Deuve 2013
Orthogonius maindronianus Tian & Deuve 2006
Orthogonius makiling Tian & Deuve 2006
Orthogonius malacca Tian & Deuve 2006
Orthogonius malaisei Andrewes 1947
Orthogonius malaya Tian & Deuve 2006
Orthogonius malaysiaensis Tian & Deuve 2006
Orthogonius medanensis Tian & Deuve 2006
Orthogonius meghalayaensis Tian & Deuve 2016
Orthogonius mekongensis Tian & Deuve 2006
Orthogonius melanipes Tian & Deuve 2006
Orthogonius mellyi (Chaudoir 1850)
Orthogonius minimus Tian & Deuve 2006
Orthogonius mniszechi Chaudoir 1872
Orthogonius moestus Chaudoir 1872
Orthogonius monolophus Andrewes 1931
Orthogonius montanus Tian & Deuve 2006
Orthogonius morvani Tian & Deuve 2003
Orthogonius morvanianus Tian & Deuve 2016
Orthogonius mouhoti Chaudoir 1872
Orthogonius myanmarensis Tian & Deuve 2006
Orthogonius nahaeo Tian & Deuve 2006
Orthogonius nepalensis Habu 1979
Orthogonius nepalicus Tian & Deuve 2006
Orthogonius niger Jedlicka 1935
Orthogonius nigripes Tian & Deuve 2001
Orthogonius nongfaensis Tian & Kirschenhofer 2013
Orthogonius nyassicus Kolbe 1895
Orthogonius obliquepes Tian & Deuve 2006
Orthogonius opacus Schmidt-Goebel 1846
Orthogonius orphnodes Andrewes 1930
Orthogonius ovatulus Tian & Deuve 2003
Orthogonius ovatus Chaudoir 1878
Orthogonius pachlatkoi Tian & Deuve 2006
Orthogonius palangbangensis Tian & Deuve 2006
Orthogonius panghongae Tian & Deuve 2006
Orthogonius pangi Tian & Deuve 2006
Orthogonius parallelus Chaudoir 1872
Orthogonius parastantoni Tian & Deuve 2006
Orthogonius parasuturalis Tian & Deuve 2006
Orthogonius parcepunctatus Kolbe 1895
Orthogonius paris Tian & Deuve 2006
Orthogonius parvus Chaudoir 1872
Orthogonius pectinatus Tian & Deuve 2016
Orthogonius perakensis Tian & Deuve 2006
Orthogonius perakicus Tian & Deuve 2007
Orthogonius perpuncticollis Burgeon 1937
Orthogonius petiolaris Tian & Deuve 2006
Orthogonius piceus Chaudoir 1872
Orthogonius picilabris (W. S. MacLeay 1825)
Orthogonius picipennis Chaudoir 1872
Orthogonius pinguis Murray 1858
Orthogonius pinophilus Tian
Orthogonius planiger (Walker 1858)
Orthogonius plebius Tian & Deuve 2006
Orthogonius poggii Tian & Deuve 2007
Orthogonius politior Tian & Deuve 2016
Orthogonius politus Chaudoir 1872
Orthogonius pollinctor Basilewsky 1948
Orthogonius pradieri Chaudoir 1872
Orthogonius pseudochaudoiri Tian
Orthogonius pseudocheni Tian & Deuve 2006
Orthogonius pseudolongicornis Tian & Deuve 2006
Orthogonius pseudomouhoti Tian & Deuve 2008
Orthogonius punctatellus Tian & Deuve 2006
Orthogonius punctulatus Chaudoir 1872
Orthogonius punctum Tian & Deuve 2016
Orthogonius putzeysi Tian & Deuve 2006
Orthogonius quadrisetosus Tian & Deuve 2006
Orthogonius rhodesianus Csiki 1932
Orthogonius rhodesiensis Lorenz 1998
Orthogonius ribbei Tian & Deuve 2003
Orthogonius rogueti Tian & Deuve 2016
Orthogonius rotundatus Tian & Deuve 2006
Orthogonius rufiventris Bates 1892
Orthogonius rufotestaceus G. Müller 1941
Orthogonius sabahensis Tian & Deuve 2006
Orthogonius sabahicus Tian & Deuve 2016
Orthogonius salvazai Tian & Deuve 2006
Orthogonius sarawakensis Tian & Deuve 2006
Orthogonius saundersi Andrewes 1926
Orthogonius schaumi Chaudoir 1872
Orthogonius schaumioides Tian & Deuve 2006
Orthogonius schauteni Tian & Deuve 2006
Orthogonius schmidtgoebeli Chaudoir 1872
Orthogonius senegalensis Dejean 1831
Orthogonius setosopalpiger Tian
Orthogonius siamensis Tian & Deuve 2006
Orthogonius similaris Tian
Orthogonius simplicatus Tian & Deuve 2006
Orthogonius singaporensis Tian & Deuve 2006
Orthogonius sinuatiphallus Tian & Deuve 2001
Orthogonius smetsi Tian & Deuve 2006
Orthogonius solidicornis Tian & Deuve 2003
Orthogonius solidus Tian & Deuve 2006
Orthogonius spinosus Burgeon 1937
Orthogonius srilankaicus Tian & Deuve 2003
Orthogonius stantoni Tian & Deuve 2006
Orthogonius sterbai Jedlicka 1935
Orthogonius stygius Andrewes 1930
Orthogonius sulawesicus Tian & Deuve 2003
Orthogonius sulcatoides Tian & Deuve 2006
Orthogonius sulcatus Schmidt-Goebel 1846
Orthogonius sumatraicus Tian & Deuve 2006
Orthogonius suturalis Chaudoir 1872
Orthogonius taghavianae Tian
Orthogonius taiwanicus Tian & Deuve 2006
Orthogonius tangsen Tian & Deuve 2006
Orthogonius tenasserimensis Tian & Deuve 2006
Orthogonius termiticola Wasmann 1902
Orthogonius thaicus Tian & Deuve 2003
Orthogonius thaiensis Tian & Deuve 2006
Orthogonius thailandensis Tian & Deuve 2006
Orthogonius thoracicus Gestro 1875
Orthogonius tikekee Tian & Deuve 2006
Orthogonius tonkinensis Tian & Deuve 2006
Orthogonius tonkinicus Tian & Deuve 2006
Orthogonius travancore Tian & Deuve 2006
Orthogonius tricarinatus Burgeon 1937
Orthogonius uncipennis Tian & Deuve 2006
Orthogonius vari Tian
Orthogonius variabilis Tian
Orthogonius virgulatus Andrewes 1931
Orthogonius wallardiensis Tian & Deuve 2006
Orthogonius wrasei Tian & Deuve 2016
Orthogonius xanthomerus L. Redtenbacher 1868
Orthogonius yoga Tian & Deuve 2006
Orthogonius yunnanensis Tian & Deuve 2001
Actenoncus Chaudoir 1872
Anoncopeucus Chaudoir 1872
Hexachaetus Chaudoir 1872
Neoorthogonius Tian & Deuve 2006
Nepalorthogonius Habu 1979

Dynamic Hierarchy Version 2.2 - Test

Life
Cellular Organisms
Eukaryota
Opisthokonta
Metazoa
Bilateria
Protostomia
Ecdysozoa
Arthropoda
Pancrustacea
Hexapoda
Insecta
Pterygota
Neoptera
Endopterygota
Coleoptera
Adephaga
Carabidae
Orthogonius (this page)
Orthogonius aberlenci Tian & Deuve 2016
Orthogonius acrogonus (Wiedemann 1819)
Orthogonius acutangulus Chaudoir 1878
Orthogonius adoriae Tian & Deuve 2006
Orthogonius aemulus Péringuey 1896
Orthogonius alternans (Wiedemann 1823)
Orthogonius alutaceus Quedenfeldt 1883
Orthogonius andrewesi van Emden 1928
Orthogonius andrewesianus Tian & Deuve 2006
Orthogonius angkor Tian & Deuve 2006
Orthogonius angkorensis Tian & Deuve 2008
Orthogonius angustus Chaudoir 1872
Orthogonius annamensis Jedlicka 1965
Orthogonius annamicola Tian & Deuve 2006
Orthogonius annemariae Tian & Deuve 2006
Orthogonius assamensis Tian & Deuve 2006
Orthogonius assamicola Tian & Deuve 2006
Orthogonius assmuthi Wasmann 1920
Orthogonius baconi Chaudoir 1872
Orthogonius baconioides Tian & Sabu 2009
Orthogonius balthasari Jedlicka 1935
Orthogonius batesi Tian & Deuve 2003
Orthogonius batesianus Tian & Deuve 2007
Orthogonius batesioides Tian & Deuve 2006
Orthogonius baumi Jedlicka 1955
Orthogonius becqueti Burgeon 1937
Orthogonius bellus Tian & Deuve 2016
Orthogonius bicolor Jedlicka 1965
Orthogonius borneoensis Tian & Deuve 2006
Orthogonius brancuccii Tian & Deuve 2006
Orthogonius brevilabris H. Kolbe 1889
Orthogonius brevithorax Dejean 1825
Orthogonius bruggei Tian & Deuve 2006
Orthogonius buquetii Chaudoir 1850
Orthogonius burckhardti Tian & Deuve 2006
Orthogonius burmanensis Tian & Deuve 2006
Orthogonius caffer Boheman 1848
Orthogonius cambodgensis Tian & Deuve 2006
Orthogonius canaraensis Tian & Deuve 2006
Orthogonius canaraicola Tian & Deuve 2006
Orthogonius capucinus Boheman 1848
Orthogonius carbonicolor Tian & Deuve 2006
Orthogonius carinatus Tian & Deuve 2013
Orthogonius celebesicus Tian & Deuve 2006
Orthogonius ceylanicus Tian & Deuve 2016
Orthogonius chaudoiri Tian & Deuve 2001
Orthogonius cheni Tian & Deuve 2001
Orthogonius chiangensis Tian & Deuve 2006
Orthogonius clarkii Murray 1858
Orthogonius confusus Tian & Deuve 2006
Orthogonius constanti Tian & Deuve 2006
Orthogonius constrictus Tian
Orthogonius coomani Tian & Deuve 2006
Orthogonius coomanioides Tian
Orthogonius coracinus H. Kolbe 1896
Orthogonius crassicrus Chaudoir 1872
Orthogonius cruralis Putzeys ex Chaudoir 1872
Orthogonius curvatus Tian & Deuve 2006
Orthogonius cyclothorax Tian & Deuve 2007
Orthogonius davidi Chaudoir 1878
Orthogonius deletus Schmidt-Goebel 1846
Orthogonius deuvei Tian & Kirschenhofer 2013
Orthogonius dispar Bates 1892
Orthogonius dohertyi Tian & Deuve 2006
Orthogonius dongnanya Tian & Deuve 2006
Orthogonius doriae Putzeys ex Chaudoir 1872
Orthogonius drescheri Liebke 1937
Orthogonius drumonti Tian & Deuve 2006
Orthogonius duboisi Tian & Deuve 2006
Orthogonius duplicatus (Wiedemann 1819)
Orthogonius dupuisi Basilewsky 1948
Orthogonius dupuisianus Tian & Deuve 2010
Orthogonius dureli Tian & Deuve 2005
Orthogonius edentatus Tian & Deuve 2006
Orthogonius elisabethanus Burgeon 1937
Orthogonius emarginatus Tian & Deuve 2007
Orthogonius equimarginalis Tian & Deuve 2006
Orthogonius euthyphallus Tian & Deuve 2013
Orthogonius fairmairei Tian & Deuve 2006
Orthogonius feai Tian & Deuve 2007
Orthogonius femoralis Chaudoir 1848
Orthogonius flavipes Deuve 2004
Orthogonius flavus Jedlicka 1964
Orthogonius foveiclypeus Tian & Deuve 2004
Orthogonius fryi Tian & Deuve 2006
Orthogonius fugax Chaudoir 1872
Orthogonius gracililamella Tian
Orthogonius gracilipes Tian & Deuve 2006
Orthogonius gracilis Tian & Deuve 2006
Orthogonius grootaerti Tian & Deuve 2006
Orthogonius hageni Oberthür 1883
Orthogonius himalaya Tian & Deuve 2006
Orthogonius himalayicus Tian & Deuve 2005
Orthogonius hirtulus Tian & Deuve 2006
Orthogonius hirtus Chaudoir 1872
Orthogonius hopei Gray 1832
Orthogonius huananoides Tian & Deuve 2006
Orthogonius huananus Tian & Deuve 2001
Orthogonius hypocrita Chaudoir 1872
Orthogonius hypocritoides Jedlicka 1935
Orthogonius impunctipennis Quedenfeldt 1883
Orthogonius indicus Tian & Deuve 2006
Orthogonius indophilus Tian & Deuve 2006
Orthogonius inexpectatus Tian & Deuve 2006
Orthogonius inops Chaudoir 1872
Orthogonius insularis Chaudoir 1872
Orthogonius intermedius Chaudoir 1872
Orthogonius javanus Tian & Deuve 2006
Orthogonius jianfengling Tian & Deuve 2001
Orthogonius kalimantanensis Tian & Deuve 2006
Orthogonius kapiriensis Burgeon 1937
Orthogonius katangensis Burgeon 1937
Orthogonius kickeli H. Kolbe 1896
Orthogonius kirirom Tian & Deuve 2008
Orthogonius klickai Jedlicka 1935
Orthogonius kubani Tian & Deuve 2006
Orthogonius kumatai Habu 1979
Orthogonius lancangjiang Tian & Deuve 2006
Orthogonius lao Tian & Deuve 2006
Orthogonius latreillei Tian & Deuve 2006
Orthogonius latus Hope 1842
Orthogonius ledouxi Tian & Deuve 2006
Orthogonius legrandi Tian & Deuve 2006
Orthogonius lieftincki Andrewes 1936
Orthogonius limbourgi Tian & Deuve 2016
Orthogonius loeicus Tian & Deuve 2006
Orthogonius loeiensis Tian & Deuve 2006
Orthogonius longicornis Chaudoir 1872
Orthogonius longilamella Tian & Deuve 2008
Orthogonius longipennis Hope 1842
Orthogonius longiphallus Tian & Deuve 2005
Orthogonius lucidus Bates 1891
Orthogonius luzonicus Chaudoir 1872
Orthogonius macrophthalmus Tian & Deuve 2013
Orthogonius maindronianus Tian & Deuve 2006
Orthogonius makiling Tian & Deuve 2006
Orthogonius malacca Tian & Deuve 2006
Orthogonius malaisei Andrewes 1947
Orthogonius malaya Tian & Deuve 2006
Orthogonius malaysiaensis Tian & Deuve 2006
Orthogonius medanensis Tian & Deuve 2006
Orthogonius meghalayaensis Tian & Deuve 2016
Orthogonius mekongensis Tian & Deuve 2006
Orthogonius melanipes Tian & Deuve 2006
Orthogonius mellyi (Chaudoir 1850)
Orthogonius minimus Tian & Deuve 2006
Orthogonius mniszechi Chaudoir 1872
Orthogonius moestus Chaudoir 1872
Orthogonius monolophus Andrewes 1931
Orthogonius montanus Tian & Deuve 2006
Orthogonius morvani Tian & Deuve 2003
Orthogonius morvanianus Tian & Deuve 2016
Orthogonius mouhoti Chaudoir 1872
Orthogonius myanmarensis Tian & Deuve 2006
Orthogonius nahaeo Tian & Deuve 2006
Orthogonius nepalensis Habu 1979
Orthogonius nepalicus Tian & Deuve 2006
Orthogonius niger Jedlicka 1935
Orthogonius nigripes Tian & Deuve 2001
Orthogonius nongfaensis Tian & Kirschenhofer 2013
Orthogonius nyassicus H. Kolbe 1896
Orthogonius obliquepes Tian & Deuve 2006
Orthogonius opacus Schmidt-Goebel 1846
Orthogonius orphnodes Andrewes 1930
Orthogonius ovatulus Tian & Deuve 2003
Orthogonius ovatus Chaudoir 1878
Orthogonius pachlatkoi Tian & Deuve 2006
Orthogonius palangbangensis Tian & Deuve 2006
Orthogonius panghongae Tian & Deuve 2006
Orthogonius pangi Tian & Deuve 2006
Orthogonius parallelus Chaudoir 1872
Orthogonius parastantoni Tian & Deuve 2006
Orthogonius parasuturalis Tian & Deuve 2006
Orthogonius parcepunctatus H. Kolbe 1896
Orthogonius paris Tian & Deuve 2006
Orthogonius parvus Chaudoir 1872
Orthogonius pectinatus Tian & Deuve 2016
Orthogonius perakensis Tian & Deuve 2006
Orthogonius perakicus Tian & Deuve 2007
Orthogonius perpuncticollis Burgeon 1937
Orthogonius petiolaris Tian & Deuve 2006
Orthogonius piceus Chaudoir 1872
Orthogonius picilabris W. S. MacLeay 1825
Orthogonius picipennis Chaudoir 1872
Orthogonius pinguis Murray 1858
Orthogonius pinophilus Tian
Orthogonius planiger (Walker 1858)
Orthogonius plebius Tian & Deuve 2006
Orthogonius poggii Tian & Deuve 2007
Orthogonius politior Tian & Deuve 2016
Orthogonius politus Chaudoir 1872
Orthogonius pollinctor Basilewsky 1948
Orthogonius pradieri Chaudoir 1872
Orthogonius pseudochaudoiri Tian
Orthogonius pseudocheni Tian & Deuve 2006
Orthogonius pseudolongicornis Tian & Deuve 2006
Orthogonius pseudomouhoti Tian & Deuve 2008
Orthogonius punctatellus Tian & Deuve 2006
Orthogonius punctulatus Chaudoir 1872
Orthogonius punctum Tian & Deuve 2016
Orthogonius putzeysi Tian & Deuve 2006
Orthogonius quadrisetosus Tian & Deuve 2006
Orthogonius rhodesianus Csiki 1932
Orthogonius rhodesiensis Lorenz 1998
Orthogonius ribbei Tian & Deuve 2003
Orthogonius rogueti Tian & Deuve 2016
Orthogonius rotundatus Tian & Deuve 2006
Orthogonius rufiventris Bates 1892
Orthogonius rufotestaceus G. Müller 1941
Orthogonius sabahensis Tian & Deuve 2006
Orthogonius sabahicus Tian & Deuve 2016
Orthogonius salvazai Tian & Deuve 2006
Orthogonius sarawakensis Tian & Deuve 2006
Orthogonius saundersi Andrewes 1926
Orthogonius schaumi Chaudoir 1872
Orthogonius schaumioides Tian & Deuve 2006
Orthogonius schauteni Tian & Deuve 2006
Orthogonius schmidtgoebeli Chaudoir 1872
Orthogonius senegalensis Dejean 1831
Orthogonius setosopalpiger Tian
Orthogonius siamensis Tian & Deuve 2006
Orthogonius similaris Tian
Orthogonius simplicatus Tian & Deuve 2006
Orthogonius singaporensis Tian & Deuve 2006
Orthogonius sinuatiphallus Tian & Deuve 2001
Orthogonius smetsi Tian & Deuve 2006
Orthogonius solidicornis Tian & Deuve 2003
Orthogonius solidus Tian & Deuve 2006
Orthogonius spinosus Burgeon 1937
Orthogonius srilankaicus Tian & Deuve 2003
Orthogonius stantoni Tian & Deuve 2006
Orthogonius sterbai Jedlicka 1935
Orthogonius stygius Andrewes 1930
Orthogonius sulawesicus Tian & Deuve 2003
Orthogonius sulcatoides Tian & Deuve 2006
Orthogonius sulcatus Schmidt-Goebel 1846
Orthogonius sumatraicus Tian & Deuve 2006
Orthogonius suturalis Chaudoir 1872
Orthogonius taghavianae Tian
Orthogonius taiwanicus Tian & Deuve 2006
Orthogonius tangsen Tian & Deuve 2006
Orthogonius tenasserimensis Tian & Deuve 2006
Orthogonius termiticola Wasmann 1902
Orthogonius thaicus Tian & Deuve 2003
Orthogonius thaiensis Tian & Deuve 2006
Orthogonius thailandensis Tian & Deuve 2006
Orthogonius thoracicus Gestro 1875
Orthogonius tikekee Tian & Deuve 2006
Orthogonius tonkinensis Tian & Deuve 2006
Orthogonius tonkinicus Tian & Deuve 2006
Orthogonius travancore Tian & Deuve 2006
Orthogonius tricarinatus Burgeon 1937
Orthogonius uncipennis Tian & Deuve 2006
Orthogonius vari Tian
Orthogonius variabilis Tian
Orthogonius virgulatus Andrewes 1931
Orthogonius wallardiensis Tian & Deuve 2006
Orthogonius wrasei Tian & Deuve 2016
Orthogonius xanthomerus L. Redtenbacher 1867
Orthogonius yoga Tian & Deuve 2006
Orthogonius yunnanensis Tian & Deuve 2001
Aepiblemus
Allendia
Allocota
Amaroschema
Amblystomus
Anillopsidius
Anophthalmus
Antennaria
Archipatrobus
Aristochroa
Aristolebia
Assadecma
Atlantomasoreus
Bennigsenium
Bhutanotrechus
Blenniventer
Bradycidus
Cascellius
Chaetauchenium
Chaetocrepis
Chlaeminus
Cratoferonia
Deipyrodes
Demetrias
Diceromerus
Dimorphoreicheia
Dyschirius
Erephognathus
Eripus
Eutogeneius
Habrodera
Hygranillus
Hystricosphodrus
Jeannelius
Kalchdigor
Lachnolebia
Leptognatha
Liagonum
Liocolpodes
Lophyra
Marsyas
Masoreus
Megacephala
Megadromus
Mexisphodrus
Mimotrechus
Minosaphaenops
Monodryxus
Nannoryctes
Neohelluo
Neohyparpalus
Nesarpalus
Notonomus
Ocys
Oedesis
Oxycrepis
Oxygnathopsis
Oxytrechus
Pachycarus
Paracolpodes
Paraliagonum
Pasimachus
Paulianites
Pelecium
Phaeoxantha
Philipis
Physolaesthus
Platypatrobus
Platysphyrus
Pogonoglossus
Pometon
Potamagonum
Procletus
Prothymidia
Protopaussus
Pseudobatenus
Rathymus
Rawlinsius
Rhytiferonia
Ryugadous
Sericoda
Siagonyx
Singilis
Sinocimmerites
Sinometrius
Sloanoglymmius
Smeringocera
Sphaerodinus
Steganomma
Stricklandiana
Tachys
Taridius
Tasmanillus
Trechiamiotes
Trechosia
Trichocarenum
Trichoxycentrus
Typhlochoromus
Xenodochus
Yukihikous
1984 additional siblings truncated for brevity.
See the resource file for a full list.

EOL Dynamic Hierarchy 1.1

Life
Cellular Organisms
Eukaryota
Opisthokonta
Metazoa
Bilateria
Protostomia
Ecdysozoa
Panarthropoda
Arthropoda
Pancrustacea
Hexapoda
Insecta
Pterygota
Neoptera
Endopterygota
Coleoptera
Adephaga
Carabidae
Orthogonius (this page)
Orthogonius aberlenci Tian & Deuve 2016
Orthogonius acrogonus (Wiedemann 1819)
Orthogonius acutangulus Chaudoir 1878
Orthogonius adoriae Tian & Deuve 2006
Orthogonius aemulus Péringuey 1896
Orthogonius alternans (Wiedemann 1823)
Orthogonius alutaceus Quedenfeldt 1883
Orthogonius andrewesi van Emden 1928
Orthogonius andrewesianus Tian & Deuve 2006
Orthogonius angkor Tian & Deuve 2006
Orthogonius angkorensis Tian & Deuve 2008
Orthogonius angustus Chaudoir 1872
Orthogonius annamensis Jedlicka 1965
Orthogonius annamicola Tian & Deuve 2006
Orthogonius annemariae Tian & Deuve 2006
Orthogonius assamensis Tian & Deuve 2006
Orthogonius assamicola Tian & Deuve 2006
Orthogonius assmuthi Wasmann 1920
Orthogonius baconi Chaudoir 1872
Orthogonius baconioides Tian & Sabu 2009
Orthogonius balthasari Jedlicka 1935
Orthogonius batesi Tian & Deuve 2003
Orthogonius batesianus Tian & Deuve 2007
Orthogonius batesioides Tian & Deuve 2006
Orthogonius baumi Jedlicka 1955
Orthogonius becqueti Burgeon 1937
Orthogonius bellus Tian & Deuve 2016
Orthogonius bicolor Jedlicka 1965
Orthogonius borneoensis Tian & Deuve 2006
Orthogonius brancuccii Tian & Deuve 2006
Orthogonius brevilabris H. Kolbe 1889
Orthogonius brevithorax Dejean 1825
Orthogonius bruggei Tian & Deuve 2006
Orthogonius buquetii Chaudoir 1850
Orthogonius burckhardti Tian & Deuve 2006
Orthogonius burmanensis Tian & Deuve 2006
Orthogonius caffer Boheman 1848
Orthogonius cambodgensis Tian & Deuve 2006
Orthogonius canaraensis Tian & Deuve 2006
Orthogonius canaraicola Tian & Deuve 2006
Orthogonius capucinus Boheman 1848
Orthogonius carbonicolor Tian & Deuve 2006
Orthogonius carinatus Tian & Deuve 2013
Orthogonius celebesicus Tian & Deuve 2006
Orthogonius ceylanicus Tian & Deuve 2016
Orthogonius chaudoiri Tian & Deuve 2001
Orthogonius cheni Tian & Deuve 2001
Orthogonius chiangensis Tian & Deuve 2006
Orthogonius clarkii Murray 1858
Orthogonius confusus Tian & Deuve 2006
Orthogonius constanti Tian & Deuve 2006
Orthogonius constrictus Tian
Orthogonius coomani Tian & Deuve 2006
Orthogonius coomanioides Tian
Orthogonius coracinus H. Kolbe 1896
Orthogonius crassicrus Chaudoir 1872
Orthogonius cruralis Putzeys ex Chaudoir 1872
Orthogonius curvatus Tian & Deuve 2006
Orthogonius cyclothorax Tian & Deuve 2007
Orthogonius davidi Chaudoir 1878
Orthogonius deletus Schmidt-Goebel 1846
Orthogonius deuvei Tian & Kirschenhofer 2013
Orthogonius dispar Bates 1892
Orthogonius dohertyi Tian & Deuve 2006
Orthogonius dongnanya Tian & Deuve 2006
Orthogonius doriae Putzeys ex Chaudoir 1872
Orthogonius drescheri Liebke 1937
Orthogonius drumonti Tian & Deuve 2006
Orthogonius duboisi Tian & Deuve 2006
Orthogonius duplicatus (Wiedemann 1819)
Orthogonius dupuisi Basilewsky 1948
Orthogonius dupuisianus Tian & Deuve 2010
Orthogonius dureli Tian & Deuve 2005
Orthogonius edentatus Tian & Deuve 2006
Orthogonius elisabethanus Burgeon 1937
Orthogonius emarginatus Tian & Deuve 2007
Orthogonius equimarginalis Tian & Deuve 2006
Orthogonius euthyphallus Tian & Deuve 2013
Orthogonius fairmairei Tian & Deuve 2006
Orthogonius feai Tian & Deuve 2007
Orthogonius femoralis Chaudoir 1848
Orthogonius flavipes Deuve 2004
Orthogonius flavus Jedlicka 1964
Orthogonius foveiclypeus Tian & Deuve 2004
Orthogonius fryi Tian & Deuve 2006
Orthogonius fugax Chaudoir 1872
Orthogonius gracililamella Tian
Orthogonius gracilipes Tian & Deuve 2006
Orthogonius gracilis Tian & Deuve 2006
Orthogonius grootaerti Tian & Deuve 2006
Orthogonius hageni Oberthür 1883
Orthogonius himalaya Tian & Deuve 2006
Orthogonius himalayicus Tian & Deuve 2005
Orthogonius hirtulus Tian & Deuve 2006
Orthogonius hirtus Chaudoir 1872
Orthogonius hopei Gray 1832
Orthogonius huananoides Tian & Deuve 2006
Orthogonius huananus Tian & Deuve 2001
Orthogonius hypocrita Chaudoir 1872
Orthogonius hypocritoides Jedlicka 1935
Orthogonius impunctipennis Quedenfeldt 1883
Orthogonius indicus Tian & Deuve 2006
Orthogonius indophilus Tian & Deuve 2006
Orthogonius inexpectatus Tian & Deuve 2006
Orthogonius inops Chaudoir 1872
Orthogonius insularis Chaudoir 1872
Orthogonius intermedius Chaudoir 1872
Orthogonius javanus Tian & Deuve 2006
Orthogonius jianfengling Tian & Deuve 2001
Orthogonius kalimantanensis Tian & Deuve 2006
Orthogonius kapiriensis Burgeon 1937
Orthogonius katangensis Burgeon 1937
Orthogonius kickeli H. Kolbe 1896
Orthogonius kirirom Tian & Deuve 2008
Orthogonius klickai Jedlicka 1935
Orthogonius kubani Tian & Deuve 2006
Orthogonius kumatai Habu 1979
Orthogonius lancangjiang Tian & Deuve 2006
Orthogonius lao Tian & Deuve 2006
Orthogonius latreillei Tian & Deuve 2006
Orthogonius latus Hope 1842
Orthogonius ledouxi Tian & Deuve 2006
Orthogonius legrandi Tian & Deuve 2006
Orthogonius lieftincki Andrewes 1936
Orthogonius limbourgi Tian & Deuve 2016
Orthogonius loeicus Tian & Deuve 2006
Orthogonius loeiensis Tian & Deuve 2006
Orthogonius longicornis Chaudoir 1872
Orthogonius longilamella Tian & Deuve 2008
Orthogonius longipennis Hope 1842
Orthogonius longiphallus Tian & Deuve 2005
Orthogonius lucidus Bates 1891
Orthogonius luzonicus Chaudoir 1872
Orthogonius macrophthalmus Tian & Deuve 2013
Orthogonius maindronianus Tian & Deuve 2006
Orthogonius makiling Tian & Deuve 2006
Orthogonius malacca Tian & Deuve 2006
Orthogonius malaisei Andrewes 1947
Orthogonius malaya Tian & Deuve 2006
Orthogonius malaysiaensis Tian & Deuve 2006
Orthogonius medanensis Tian & Deuve 2006
Orthogonius meghalayaensis Tian & Deuve 2016
Orthogonius mekongensis Tian & Deuve 2006
Orthogonius melanipes Tian & Deuve 2006
Orthogonius mellyi (Chaudoir 1850)
Orthogonius minimus Tian & Deuve 2006
Orthogonius mniszechi Chaudoir 1872
Orthogonius moestus Chaudoir 1872
Orthogonius monolophus Andrewes 1931
Orthogonius montanus Tian & Deuve 2006
Orthogonius morvani Tian & Deuve 2003
Orthogonius morvanianus Tian & Deuve 2016
Orthogonius mouhoti Chaudoir 1872
Orthogonius myanmarensis Tian & Deuve 2006
Orthogonius nahaeo Tian & Deuve 2006
Orthogonius nepalensis Habu 1979
Orthogonius nepalicus Tian & Deuve 2006
Orthogonius niger Jedlicka 1935
Orthogonius nigripes Tian & Deuve 2001
Orthogonius nongfaensis Tian & Kirschenhofer 2013
Orthogonius nyassicus H. Kolbe 1896
Orthogonius obliquepes Tian & Deuve 2006
Orthogonius opacus Schmidt-Goebel 1846
Orthogonius orphnodes Andrewes 1930
Orthogonius ovatulus Tian & Deuve 2003
Orthogonius ovatus Chaudoir 1878
Orthogonius pachlatkoi Tian & Deuve 2006
Orthogonius palangbangensis Tian & Deuve 2006
Orthogonius panghongae Tian & Deuve 2006
Orthogonius pangi Tian & Deuve 2006
Orthogonius parallelus Chaudoir 1872
Orthogonius parastantoni Tian & Deuve 2006
Orthogonius parasuturalis Tian & Deuve 2006
Orthogonius parcepunctatus H. Kolbe 1896
Orthogonius paris Tian & Deuve 2006
Orthogonius parvus Chaudoir 1872
Orthogonius pectinatus Tian & Deuve 2016
Orthogonius perakensis Tian & Deuve 2006
Orthogonius perakicus Tian & Deuve 2007
Orthogonius perpuncticollis Burgeon 1937
Orthogonius petiolaris Tian & Deuve 2006
Orthogonius piceus Chaudoir 1872
Orthogonius picilabris W. S. Mac Leay 1825
Orthogonius picipennis Chaudoir 1872
Orthogonius pinguis Murray 1858
Orthogonius pinophilus Tian
Orthogonius planiger (Walker 1858)
Orthogonius plebius Tian & Deuve 2006
Orthogonius poggii Tian & Deuve 2007
Orthogonius politior Tian & Deuve 2016
Orthogonius politus Chaudoir 1872
Orthogonius pollinctor Basilewsky 1948
Orthogonius pradieri Chaudoir 1872
Orthogonius pseudochaudoiri Tian
Orthogonius pseudocheni Tian & Deuve 2006
Orthogonius pseudolongicornis Tian & Deuve 2006
Orthogonius pseudomouhoti Tian & Deuve 2008
Orthogonius punctatellus Tian & Deuve 2006
Orthogonius punctulatus Chaudoir 1872
Orthogonius punctum Tian & Deuve 2016
Orthogonius putzeysi Tian & Deuve 2006
Orthogonius quadrisetosus Tian & Deuve 2006
Orthogonius rhodesianus Csiki 1932
Orthogonius rhodesiensis Lorenz 1998
Orthogonius ribbei Tian & Deuve 2003
Orthogonius rogueti Tian & Deuve 2016
Orthogonius rotundatus Tian & Deuve 2006
Orthogonius rufiventris Bates 1892
Orthogonius rufotestaceus G. Müller 1941
Orthogonius sabahensis Tian & Deuve 2006
Orthogonius sabahicus Tian & Deuve 2016
Orthogonius salvazai Tian & Deuve 2006
Orthogonius sarawakensis Tian & Deuve 2006
Orthogonius saundersi Andrewes 1926
Orthogonius schaumi Chaudoir 1872
Orthogonius schaumioides Tian & Deuve 2006
Orthogonius schauteni Tian & Deuve 2006
Orthogonius schmidtgoebeli Chaudoir 1872
Orthogonius senegalensis Dejean 1831
Orthogonius setosopalpiger Tian
Orthogonius siamensis Tian & Deuve 2006
Orthogonius similaris Tian
Orthogonius simplicatus Tian & Deuve 2006
Orthogonius singaporensis Tian & Deuve 2006
Orthogonius sinuatiphallus Tian & Deuve 2001
Orthogonius smetsi Tian & Deuve 2006
Orthogonius solidicornis Tian & Deuve 2003
Orthogonius solidus Tian & Deuve 2006
Orthogonius spinosus Burgeon 1937
Orthogonius srilankaicus Tian & Deuve 2003
Orthogonius stantoni Tian & Deuve 2006
Orthogonius sterbai Jedlicka 1935
Orthogonius stygius Andrewes 1930
Orthogonius sulawesicus Tian & Deuve 2003
Orthogonius sulcatoides Tian & Deuve 2006
Orthogonius sulcatus Schmidt-Goebel 1846
Orthogonius sumatraicus Tian & Deuve 2006
Orthogonius suturalis Chaudoir 1872
Orthogonius taghavianae Tian
Orthogonius taiwanicus Tian & Deuve 2006
Orthogonius tangsen Tian & Deuve 2006
Orthogonius tenasserimensis Tian & Deuve 2006
Orthogonius termiticola Wasmann 1902
Orthogonius thaicus Tian & Deuve 2003
Orthogonius thaiensis Tian & Deuve 2006
Orthogonius thailandensis Tian & Deuve 2006
Orthogonius thoracicus Gestro 1875
Orthogonius tikekee Tian & Deuve 2006
Orthogonius tonkinensis Tian & Deuve 2006
Orthogonius tonkinicus Tian & Deuve 2006
Orthogonius travancore Tian & Deuve 2006
Orthogonius tricarinatus Burgeon 1937
Orthogonius uncipennis Tian & Deuve 2006
Orthogonius vari Tian
Orthogonius variabilis Tian
Orthogonius virgulatus Andrewes 1931
Orthogonius wallardiensis Tian & Deuve 2006
Orthogonius wrasei Tian & Deuve 2016
Orthogonius xanthomerus L. Redtenbacher 1867
Orthogonius yoga Tian & Deuve 2006
Orthogonius yunnanensis Tian & Deuve 2001
Aepiblemus
Allendia
Allocota
Amaroschema
Amblystomus
Amphoxyne
Anillopsidius
Anophthalmus
Antennaria
Archipatrobus
Aristochroa
Aristolebia
Assadecma
Atlantomasoreus
Beipiaocarabus
Bhutanotrechus
Blenniventer
Bradycidus
Cascellius
Chaetauchenium
Chaetocrepis
Chlaeminus
Cratoferonia
Deipyrodes
Demetrias
Diceromerus
Dimorphoreicheia
Dyschirius
Erephognathus
Eripus
Eutogeneius
Habrodera
Hygranillus
Hystricosphodrus
Jeannelius
Kalchdigor
Lachnolebia
Liagonum
Liocolpodes
Lophyra
Marsyas
Masoreus
Megacephala
Megadromus
Mexisphodrus
Mimotrechus
Minosaphaenops
Monodryxus
Nannoryctes
Neohelluo
Neohyparpalus
Nesarpalus
Notonomus
Ocys
Oedesis
Oxycrepis
Oxygnathopsis
Oxytrechus
Pachycarus
Paracolpodes
Paraliagonum
Pasimachus
Paulianites
Pelecium
Phaeoxantha
Philipis
Physolaesthus
Platypatrobus
Platysphyrus
Pogonoglossus
Pometon
Potamagonum
Procletus
Prothymidia
Protopaussus
Pseudobatenus
Rathymus
Rawlinsius
Rhytiferonia
Ryugadous
Siagonyx
Singilis
Sinocalosoma
Sinocimmerites
Sinometrius
Sloanoglymmius
Smeringocera
Sphaerodinus
Steganomma
Stricklandiana
Tachys
Taridius
Tasmanillus
Trechiamiotes
Trechosia
Trichocarenum
Trichoxycentrus
Typhlochoromus
Xenodochus
Yukihikous
2068 additional siblings truncated for brevity.
See the resource file for a full list.

GBIF classification

Animalia
Arthropoda
Insecta
Coleoptera
Carabidae
Orthogonius Macleay 1825 (this page)
Orthogonius aberlenci Tian & Deuve 2016
Orthogonius acrogonus (Wiedemann 1819)
Orthogonius acutangulus Chaudoir 1878
Orthogonius adoriae Tian & Deuve 2006
Orthogonius aemulus Péringuey 1896
Orthogonius alternans (Wiedemann 1823)
Orthogonius alutaceus Quedenfeldt 1883
Orthogonius andrewesi van Emden 1928
Orthogonius andrewesianus Tian & Deuve 2006
Orthogonius angkor Tian & Deuve 2006
Orthogonius angkorensis Tian & Deuve 2008
Orthogonius angustus Chaudoir 1872
Orthogonius annamensis Jedlicka 1965
Orthogonius annamicola Tian & Deuve 2006
Orthogonius annemariae Tian & Deuve 2006
Orthogonius assamensis Tian & Deuve 2006
Orthogonius assamicola Tian & Deuve 2006
Orthogonius assmuthi Wasmann 1920
Orthogonius baconi Chaudoir 1872
Orthogonius baconioides Tian & Sabu 2009
Orthogonius balthasari Jedlicka 1935
Orthogonius batesi Tian & Deuve 2003
Orthogonius batesianus Tian & Deuve 2007
Orthogonius batesioides Tian & Deuve 2006
Orthogonius baumi Jedlicka 1955
Orthogonius becqueti Burgeon 1937
Orthogonius bellus Tian & Deuve 2016
Orthogonius bicolor Jedlicka 1965
Orthogonius borneoensis Tian & Deuve 2006
Orthogonius brancuccii Tian & Deuve 2006
Orthogonius brevilabris H. Kolbe 1889
Orthogonius brevimandibularis Tian, Deuve & Presumed Unpublished
Orthogonius brevithorax Dejean 1825
Orthogonius bruggei Tian & Deuve 2006
Orthogonius buquetii Chaudoir 1850
Orthogonius burckhardti Tian & Deuve 2006
Orthogonius burmanensis Tian & Deuve 2006
Orthogonius caffer Boheman 1848
Orthogonius cambodgensis Tian & Deuve 2006
Orthogonius canaraensis Tian & Deuve 2006
Orthogonius canaraicola Tian & Deuve 2006
Orthogonius capucinus Boheman 1848
Orthogonius carbonicolor Tian & Deuve 2006
Orthogonius carinatus Tian & Deuve 2013
Orthogonius celebesicus Tian & Deuve 2006
Orthogonius ceylanicus Tian & Deuve 2016
Orthogonius chaudoiri Tian & Deuve 2001
Orthogonius cheni Tian & Deuve 2001
Orthogonius chiangensis Tian & Deuve 2006
Orthogonius clarkii Murray 1858
Orthogonius confusus Tian & Deuve 2006
Orthogonius constanti Tian & Deuve 2006
Orthogonius constrictus Tian, Deuve & Felix 2012
Orthogonius coomani Tian & Deuve 2006
Orthogonius coomanioides Tian, Deuve & Felix 2012
Orthogonius coracinus H. Kolbe 1896
Orthogonius crassicrus Chaudoir 1872
Orthogonius cruralis Putzeys 1872
Orthogonius curvatus Tian & Deuve 2006
Orthogonius cyclothorax Tian & Deuve 2007
Orthogonius davidi Chaudoir 1878
Orthogonius deletus Schmidt-Goebel 1846
Orthogonius deuvei Tian & Kirschenhofer 2013
Orthogonius dispar Bates 1892
Orthogonius dohertyi Tian & Deuve 2006
Orthogonius dongnanya Tian & Deuve 2006
Orthogonius doriae Putzeys 1872
Orthogonius drescheri Liebke 1937
Orthogonius drumonti Tian & Deuve 2006
Orthogonius duboisi Tian & Deuve 2006
Orthogonius duplicatus (Wiedemann 1819)
Orthogonius dupuisi Basilewsky 1948
Orthogonius dupuisianus Tian & Deuve 2010
Orthogonius dureli Tian & Deuve 2005
Orthogonius edentatus Tian & Deuve 2006
Orthogonius elisabethanus Burgeon 1937
Orthogonius emarginatus Tian & Deuve 2007
Orthogonius equimarginalis Tian & Deuve 2006
Orthogonius euthyphallus Tian & Deuve 2013
Orthogonius fairmairei Tian & Deuve 2006
Orthogonius feai Tian & Deuve 2007
Orthogonius femoralis Chaudoir 1848
Orthogonius flavipes Deuve 2004
Orthogonius flavus Jedlicka 1964
Orthogonius foveiclypeatus Tian, Deuve & Presumed Unpublished
Orthogonius foveiclypeus Tian & Deuve 2004
Orthogonius fryi Tian & Deuve 2006
Orthogonius fugax Chaudoir 1872
Orthogonius gracililamella Tian, Deuve & Felix 2012
Orthogonius gracilipes Tian & Deuve 2006
Orthogonius gracilis Tian & Deuve 2006
Orthogonius grootaerti Tian & Deuve 2006
Orthogonius hageni Oberthür 1883
Orthogonius himalaya Tian & Deuve 2006
Orthogonius himalayicus Tian & Deuve 2005
Orthogonius hirtulus Tian & Deuve 2006
Orthogonius hirtus Chaudoir 1872
Orthogonius hopei Gray 1832
Orthogonius huananoides Tian & Deuve 2006
Orthogonius huananus Tian & Deuve 2001
Orthogonius hypocrita Chaudoir 1872
Orthogonius hypocritoides Jedlicka 1935
Orthogonius impunctipennis Quedenfeldt 1883
Orthogonius indicus Tian & Deuve 2006
Orthogonius indophilus Tian & Deuve 2006
Orthogonius inexpectatus Tian & Deuve 2006
Orthogonius inops Chaudoir 1872
Orthogonius insularis Chaudoir 1872
Orthogonius intermedius Chaudoir 1872
Orthogonius javanus Tian & Deuve 2006
Orthogonius jianfengling Tian & Deuve 2001
Orthogonius kalimantanensis Tian & Deuve 2006
Orthogonius katangensis Burgeon 1937
Orthogonius kickeli H. Kolbe 1896
Orthogonius kirirom Tian & Deuve 2008
Orthogonius klickai Jedlicka 1935
Orthogonius kubani Tian & Deuve 2006
Orthogonius kumatai Habu 1979
Orthogonius lancangjiang Tian & Deuve 2006
Orthogonius lao Tian & Deuve 2006
Orthogonius latreillei Tian & Deuve 2006
Orthogonius latus Hope 1842
Orthogonius ledouxi Tian & Deuve 2006
Orthogonius legrandi Tian & Deuve 2006
Orthogonius leoeinsis Tian & Deuve 2006
Orthogonius lieftincki Andrewes 1936
Orthogonius limbourgi Tian & Deuve 2016
Orthogonius loeicus Tian & Deuve 2006
Orthogonius loeiensis Tian & Deuve 2006
Orthogonius longicornis Chaudoir 1872
Orthogonius longilamella Tian & Deuve 2008
Orthogonius longipennis Hope 1842
Orthogonius longiphallus Tian & Deuve 2005
Orthogonius lucidus Bates 1891
Orthogonius luzonicus Chaudoir 1872
Orthogonius macrophthalmus Tian & Deuve 2013
Orthogonius maindronianus Tian & Deuve 2006
Orthogonius makiling Tian & Deuve 2006
Orthogonius malacca Tian & Deuve 2006
Orthogonius malaisei Andrewes 1947
Orthogonius malaya Tian & Deuve 2006
Orthogonius malaysiaensis Tian & Deuve 2006
Orthogonius medanensis Tian & Deuve 2006
Orthogonius meghalayaensis Tian & Deuve 2016
Orthogonius mekongensis Tian & Deuve 2006
Orthogonius melanipes Tian & Deuve 2006
Orthogonius mellyi (Chaudoir 1850)
Orthogonius minimus Tian & Deuve 2006
Orthogonius mniszechi Chaudoir 1872
Orthogonius moestus Chaudoir 1872
Orthogonius monolophus Andrewes 1931
Orthogonius montanus Tian & Deuve 2006
Orthogonius morvani Tian & Deuve 2003
Orthogonius morvanianus Tian & Deuve 2016
Orthogonius mouhoti Chaudoir 1872
Orthogonius myanmarensis Tian & Deuve 2006
Orthogonius nahaeo Tian & Deuve 2006
Orthogonius nepalensis Habu 1979
Orthogonius nepalicus Tian & Deuve 2006
Orthogonius niger Jedlicka 1935
Orthogonius nigripes Tian & Deuve 2001
Orthogonius nongfaensis Tian & Kirschenhofer 2013
Orthogonius nyassicus H. Kolbe 1896
Orthogonius obliquepes Tian & Deuve 2006
Orthogonius opacus Schmidt-Goebel 1846
Orthogonius orphnodes Andrewes 1930
Orthogonius ovatulus Tian & Deuve 2003
Orthogonius ovatus Chaudoir 1878
Orthogonius pachlatkoi Tian & Deuve 2006
Orthogonius palangbangensis Tian & Deuve 2006
Orthogonius panghongae Tian & Deuve 2006
Orthogonius pangi Tian & Deuve 2006
Orthogonius parallelus Chaudoir 1872
Orthogonius parastantoni Tian & Deuve 2006
Orthogonius parasuturalis Tian & Deuve 2006
Orthogonius parcepunctatus H. Kolbe 1896
Orthogonius paris Tian & Deuve 2006
Orthogonius parvus Chaudoir 1872
Orthogonius pectinatus Tian & Deuve 2016
Orthogonius perakensis Tian & Deuve 2006
Orthogonius perakicus Tian & Deuve 2007
Orthogonius perpuncticollis Burgeon 1937
Orthogonius petiolaris Tian & Deuve 2006
Orthogonius piceus Chaudoir 1872
Orthogonius picilabris W. S. Mac Leay 1825
Orthogonius picipennis Chaudoir 1872
Orthogonius pinguis Murray 1858
Orthogonius pinophilus Tian, Deuve & Felix 2012
Orthogonius planiger (Walker 1858)
Orthogonius plebius Tian & Deuve 2006
Orthogonius poggii Tian & Deuve 2007
Orthogonius politior Tian & Deuve 2016
Orthogonius politus Chaudoir 1872
Orthogonius pollinctor Basilewsky 1948
Orthogonius pradieri Chaudoir 1872
Orthogonius pseudochaudoiri Tian, Deuve & Felix 2012
Orthogonius pseudocheni Tian & Deuve 2006
Orthogonius pseudolongicornis Tian & Deuve 2006
Orthogonius pseudomouhoti Tian & Deuve 2008
Orthogonius punctatellus Tian & Deuve 2006
Orthogonius punctulatus Chaudoir 1872
Orthogonius punctum Tian & Deuve 2016
Orthogonius putzeysi Tian & Deuve 2006
Orthogonius quadrisetosus Tian & Deuve 2006
Orthogonius rhodesianus Csiki 1932
Orthogonius rhodesiensis Lorenz 1998
Orthogonius ribbei Tian & Deuve 2003
Orthogonius rogueti Tian & Deuve 2016
Orthogonius rotundatus Tian & Deuve 2006
Orthogonius rufiventris Bates 1892
Orthogonius rufotestaceus G. Müller 1941
Orthogonius sabahensis Tian & Deuve 2006
Orthogonius sabahicus Tian & Deuve 2016
Orthogonius salvazai Tian & Deuve 2006
Orthogonius sarawakensis Tian & Deuve 2006
Orthogonius saundersi Andrewes 1926
Orthogonius schaumi Chaudoir 1872
Orthogonius schaumioides Tian & Deuve 2006
Orthogonius schauteni Tian & Deuve 2006
Orthogonius schmidtgoebeli Chaudoir 1872
Orthogonius senegalensis Dejean 1831
Orthogonius setosopalpiger Tian, Deuve & Felix 2012
Orthogonius siamensis Tian & Deuve 2006
Orthogonius similaris Tian, Deuve & Felix 2012
Orthogonius simplicatus Tian & Deuve 2006
Orthogonius singaporensis Tian & Deuve 2006
Orthogonius sinuatiphallus Tian & Deuve 2001
Orthogonius smetsi Tian & Deuve 2006
Orthogonius solidicornis Tian & Deuve 2003
Orthogonius solidus Tian & Deuve 2006
Orthogonius spinosus Burgeon 1937
Orthogonius srilankaicus Tian & Deuve 2003
Orthogonius stantoni Tian & Deuve 2006
Orthogonius sterbai Jedlicka 1935
Orthogonius stygius Andrewes 1930
Orthogonius sulawesicus Tian & Deuve 2003
Orthogonius sulcatoides Tian & Deuve 2006
Orthogonius sulcatus Schmidt-Goebel 1846
Orthogonius sumatraicus Tian & Deuve 2006
Orthogonius suturalis Chaudoir 1872
Orthogonius taghavianae Tian, Deuve & Felix 2012
Orthogonius taiwanicus Tian & Deuve 2006
Orthogonius tangsen Tian & Deuve 2006
Orthogonius tenasserimensis Tian & Deuve 2006
Orthogonius termiticola Wasmann 1902
Orthogonius thaicus Tian & Deuve 2003
Orthogonius thaiensis Tian & Deuve 2006
Orthogonius thailandensis Tian & Deuve 2006
Orthogonius thoracicus Gestro 1875
Orthogonius tikekee Tian & Deuve 2006
Orthogonius tonkinensis Tian & Deuve 2006
Orthogonius tonkinicus Tian & Deuve 2006
Orthogonius travancore Tian & Deuve 2006
Orthogonius tricarinatus Burgeon 1937
Orthogonius uncipennis Tian & Deuve 2006
Orthogonius vari Tian, Deuve & Felix 2012
Orthogonius variabilis Tian, Deuve & Felix 2012
Orthogonius virgulatus Andrewes 1931
Orthogonius wallardiensis Tian & Deuve 2006
Orthogonius wrasei Tian & Deuve 2016
Orthogonius xanthomerus L. Redtenbacher 1867
Orthogonius yoga Tian & Deuve 2006
Orthogonius yunnanensis Tian & Deuve 2001
Abacidus Leconte 1863
Amara Stephens 1827
Amerinus Casey 1884
Ameroduvalius Valentine 1952
Amphasia Newman 1838
Anillaspis Casey 1918
Anillinus Casey 1918
Anisodactylus Dejean 1829
Apenes Le Conte 1851
Apristus Chaudoir 1846
Ardistomis Putzeys 1846
Asaphidion Gozis 1886
Aspidoglossa Putzeys 1846
Atranus Le Conte 1847
Axinopalpus Le Conte 1846
Bembidion Latreille 1802
Blackburnia Sharp 1878
Brachinus Weber 1801
Bradycellus Erichson 1837
Broscodera Lindroth 1961
Carabus Linnaeus 1758
Cicindela Linnaeus 1758
Clivina Latreille 1802
Colliuris De Geer 1774
Cratacanthus Dejean 1829
Cychrus Fabricius 1794
Cymindis Latreille 1805
Darlingtonea Valentine 1952
Dercylinus Chaudoir 1883
Diacheila Motschulsky 1844
Dicheirus Mannerheim 1843
Diplochaetus Chaudoir 1872
Diplous Motschulsky 1850
Discoderus Le Conte 1853
Dromius Bonelli 1810
Drypta Latreille 1796
Ega Laporte 1834
Elaphrus Fabricius 1775
Endynomena Chaudoir 1872
Euphorticus G. Horn 1881
Euryderus Le Conte 1846
Evolenes Le Conte 1853
Geopinus Le Conte 1847
Gnathaphanus W. Mac Leay 1825
Halocoryza Alluaud 1919
Harpalobrachys Tschitschérine 1899
Harpalus Latreille 1802
Hartonymus Casey 1914
Himalotrechus Belousov, Kabak & Schmidt 2019
Horologion Valentine 1932
Indocarterus
Lachnophorus Dejean 1831
Lebia Latreille 1802
Loricera Latreille 1802
Loxandrus Le Conte 1852
Masoreus Dejean 1821
Megacephala Latreille 1802
Metrius Eschscholtz 1829
Micrixys Le Conte 1854
Microlestes Schmidt-Gébel 1846
Mioptachys Bates 1882
Miscodera Eschscholtz 1830
Neaphaenops Jeannel 1920
Nebria Latreille 1802
Nemotarsus Le Conte 1853
Nomius Laporte 1835
Opisthius Kirby 1837
Oxycrepis Reiche 1843
Pachyteles Perty 1830
Panagaeus Latreille 1802
Patrobus Dejean 1821
Pentagonica Schmidt-Gébel 1846
Perigona Laporte 1835
Perileptus Schaum 1860
Phloeoxena Chaudoir 1870
Phrypeus Casey 1924
Pinacodera Schaum 1857
Platidiolus Chaudoir 1878
Platypatrobus Darlington 1938
Pogonistes Chaudoir 1871
Polyderis Motschulsky 1862
Pseudanophthalmus Jeannel 1920
Psydrus Le Conte 1846
Rhadine Le Conte 1846
Scarites Fabricius 1775
Schizogenius Putzeys 1846
Selenophorus Dejean 1829
Sericoda Kirby 1837
Somotrichus Seidlitz 1887
Stenocrepis Chaudoir 1857
Tachyta Kirby 1837
Tefflus Leach 1819
Tetragonoderus Dejean 1829
Trechoblemus Ganglbauer 1891
Trichotichnus Morawitz 1863
Typhlonesiotes Jeannel 1937
Xenotrechus Barr & Krekeler 1967
Xestonotus Le Conte 1853
Zacotus Le Conte 1869
Zuphium Latreille 1805
2122 additional siblings truncated for brevity.
See the resource file for a full list.

NCBI

root
cellular organisms
Eukaryota
Opisthokonta
Metazoa
Eumetazoa
Bilateria
Protostomia
Ecdysozoa
Panarthropoda
Arthropoda
Mandibulata
Pancrustacea
Hexapoda
Insecta
Dicondylia
Pterygota
Neoptera
Holometabola
Coleoptera
Adephaga
Caraboidea
Carabidae
Harpalinae
Orthogoniini
Orthogonius (this page)
Orthogonius fugax
unclassified Orthogonius

wikidata_hierarchy tar gz

Biota
Eukaryota
Unikonta
Opisthokonta
Holozoa
Filozoa
Apoikozoa
Animalia
Eumetazoa
Bilateria
Protostomia
Ecdysozoa
Arthropoda
Hexapoda
Insecta
Dicondylia
Pterygota
Metapterygota
Neoptera
Eumetabola
Endopterygota
Coleoptera
Adephaga
Caraboidea
Carabidae
Harpalinae
Orthogoniini
Orthogonius (this page)
Orthogonius aberlenci
Orthogonius acrogonus
Orthogonius acutangulus
Orthogonius adoriae
Orthogonius aemulus
Orthogonius alternans
Orthogonius alutaceus
Orthogonius andrewesi
Orthogonius andrewesianus
Orthogonius angkor
Orthogonius angkorensis
Orthogonius angustus
Orthogonius annamensis
Orthogonius annamicola
Orthogonius annemariae
Orthogonius assamensis
Orthogonius assamicola
Orthogonius assmuthi
Orthogonius baconi
Orthogonius baconioides
Orthogonius balthasari
Orthogonius batesi
Orthogonius batesianus
Orthogonius batesioides
Orthogonius baumi
Orthogonius becqueti
Orthogonius bellus
Orthogonius bicolor
Orthogonius borneoensis
Orthogonius brancuccii
Orthogonius brevilabris
Orthogonius brevithorax
Orthogonius bruggei
Orthogonius buqueti
Orthogonius buquetii
Orthogonius burckhardti
Orthogonius burmanensis
Orthogonius caffer
Orthogonius cambodgensis
Orthogonius canaraensis
Orthogonius canaraicola
Orthogonius capucinus
Orthogonius carbonicolor
Orthogonius carinatus
Orthogonius celebesicus
Orthogonius ceylanicus
Orthogonius chaudoiri
Orthogonius cheni
Orthogonius chiangensis
Orthogonius clarkei
Orthogonius clarkii
Orthogonius confusus
Orthogonius constanti
Orthogonius constrictus
Orthogonius coomani
Orthogonius coomanioides
Orthogonius coracinus
Orthogonius crassicrus
Orthogonius cruralis
Orthogonius curvatus
Orthogonius cyclothorax
Orthogonius davidi
Orthogonius deletus
Orthogonius deuvei
Orthogonius dispar
Orthogonius dohertyi
Orthogonius dongnanya
Orthogonius doriae
Orthogonius drescheri
Orthogonius drumonti
Orthogonius duboisi
Orthogonius duplicatus
Orthogonius dupuisi
Orthogonius dupuisianus
Orthogonius dureli
Orthogonius edentatus
Orthogonius elisabethanus
Orthogonius emarginatus
Orthogonius equimarginalis
Orthogonius euthyphallus
Orthogonius fairmairei
Orthogonius feai
Orthogonius femoralis
Orthogonius flavipes
Orthogonius flavus
Orthogonius foersteri
Orthogonius foveiclypeus
Orthogonius fryi
Orthogonius fugax
Orthogonius gracililamella
Orthogonius gracilipes
Orthogonius gracilis
Orthogonius grootaerti
Orthogonius hageni
Orthogonius himalaya
Orthogonius himalayicus
Orthogonius hirtulus
Orthogonius hirtus
Orthogonius hopei
Orthogonius huananoides
Orthogonius huananus
Orthogonius hypocrita
Orthogonius hypocritoides
Orthogonius impunctipennis
Orthogonius indicus
Orthogonius indophilus
Orthogonius inexpectatus
Orthogonius inops
Orthogonius insularis
Orthogonius intermedius
Orthogonius javanus
Orthogonius jianfengling
Orthogonius kalimantanensis
Orthogonius kapiriensis
Orthogonius katangensis
Orthogonius kickeli
Orthogonius kirirom
Orthogonius klickai
Orthogonius kubani
Orthogonius kumatai
Orthogonius lancangjiang
Orthogonius lao
Orthogonius latreillei
Orthogonius latus
Orthogonius ledouxi
Orthogonius legrandi
Orthogonius lieftincki
Orthogonius limbourgi
Orthogonius loeicus
Orthogonius loeiensis
Orthogonius longicornis
Orthogonius longilamella
Orthogonius longipennis
Orthogonius longiphallus
Orthogonius lucidus
Orthogonius luzonicus
Orthogonius macrophthalmus
Orthogonius maindronianus
Orthogonius makiling
Orthogonius malacca
Orthogonius malaisei
Orthogonius malaya
Orthogonius malaysiaensis
Orthogonius medanensis
Orthogonius meghalayaensis
Orthogonius mekongensis
Orthogonius melanipes
Orthogonius mellyi
Orthogonius minimus
Orthogonius mniszechi
Orthogonius moestus
Orthogonius monolophus
Orthogonius montanus
Orthogonius morvani
Orthogonius morvanianus
Orthogonius mouhoti
Orthogonius myanmarensis
Orthogonius nahaeo
Orthogonius nepalensis
Orthogonius nepalicus
Orthogonius niger
Orthogonius nigripes
Orthogonius nongfaensis
Orthogonius nyassicus
Orthogonius obliquepes
Orthogonius opacus
Orthogonius orphnodes
Orthogonius ovatulus
Orthogonius ovatus
Orthogonius pachlatkoi
Orthogonius palangbangensis
Orthogonius panghongae
Orthogonius pangi
Orthogonius parallelus
Orthogonius parastantoni
Orthogonius parasuturalis
Orthogonius parcepunctatus
Orthogonius paris
Orthogonius parvus
Orthogonius pectinatus
Orthogonius perakensis
Orthogonius perakicus
Orthogonius perpuncticollis
Orthogonius petiolaris
Orthogonius piceus
Orthogonius picilabris
Orthogonius picipennis
Orthogonius pinguis
Orthogonius pinophilus
Orthogonius planiger
Orthogonius plebius
Orthogonius poggii
Orthogonius politior
Orthogonius politus
Orthogonius pollinctor
Orthogonius pradieri
Orthogonius pseudochaudoiri
Orthogonius pseudocheni
Orthogonius pseudolongicornis
Orthogonius pseudomouhoti
Orthogonius punctatellus
Orthogonius punctulatus
Orthogonius punctum
Orthogonius putzeysi
Orthogonius quadrisetosus
Orthogonius rhodesianus
Orthogonius rhodesiensis
Orthogonius ribbei
Orthogonius rogueti
Orthogonius rotundatus
Orthogonius rufiventris
Orthogonius rufotestaceus
Orthogonius sabahensis
Orthogonius sabahicus
Orthogonius salvazai
Orthogonius sarawakensis
Orthogonius saundersi
Orthogonius schaumi
Orthogonius schaumioides
Orthogonius schauteni
Orthogonius schmidtgoebeli
Orthogonius senegalensis
Orthogonius setosopalpiger
Orthogonius siamensis
Orthogonius similaris
Orthogonius simplicatus
Orthogonius singaporensis
Orthogonius sinuatiphallus
Orthogonius smetsi
Orthogonius solidicornis
Orthogonius solidus
Orthogonius srilankaicus
Orthogonius stantoni
Orthogonius sterbai
Orthogonius stygius
Orthogonius sulawesicus
Orthogonius sulcatoides
Orthogonius sulcatus
Orthogonius sumatraicus
Orthogonius suturalis
Orthogonius taghavianae
Orthogonius taiwanicus
Orthogonius tangsen
Orthogonius tenasserimensis
Orthogonius termiticola
Orthogonius thaicus
Orthogonius thaiensis
Orthogonius thailandensis
Orthogonius thoracicus
Orthogonius tikekee
Orthogonius tonkinensis
Orthogonius tonkinicus
Orthogonius travancore
Orthogonius tricarinatus
Orthogonius uncipennis
Orthogonius vari
Orthogonius variabilis
Orthogonius virgulatus
Orthogonius wallardiensis
Orthogonius wrasei
Orthogonius xanthomerus
Orthogonius yoga
Orthogonius yunnanensis
Hexachaetus
Orthogoniina
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