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Kodo Millet

Paspalum scrobiculatum L.

Comments

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This polymorphic species complex is thought to comprise an apomictic swarm. Some of its components have been described as separate species, but variation is continuous throughout. Robust forms are sometimes separated as Paspalum auriculatum J. Presl & C. Presl. Three recognizable entities are maintained here at varietal rank.
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Flora of China Vol. 22: 526, 528, 529, 530 in eFloras.org, Missouri Botanical Garden. Accessed Nov 12, 2008.
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Comments

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This is a highly polymorphic species, but with the variation apparently quite continuous; possibly it is a swarm of apomicts (see Clayton 1975).

It is sometimes cultivated as a hot weather crop, but under certain conditions the grain and fodder can be harmful.

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Flora of Pakistan Vol. 0: 213 in eFloras.org, Missouri Botanical Garden. Accessed Nov 12, 2008.
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Description

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Perennials; culm tufted, 50-100 cm tall, erect, decumbent at base. Blade 15-50 cm long, 7-12 mm wide, glabrous, more or less glaucous; sheath glabrous, longer than internode, compressed, keeled; ligule membranaceous, 0.5-1 mm long, with tawny hairs abaxially. Racemes 2-5(-8); rachis 1.5-3 mm wide, margins scabrous. Spikelet solitary, imbricate, glabrous, orbicular to ovate, 2-3 mm long; lower glume absent; upper glume membranaceous, 5 veined; lower lemma 5-7-veined, membranaceous or sometimes indurate; upper lemma dark brown at maturity, coriaceous.
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Gramineae (Poaceae) in Flora of Taiwan Vol. 0 in eFloras.org, Missouri Botanical Garden. Accessed Nov 12, 2008.
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Description

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Perennial or annual. Culms tufted, slender to robust, erect or decumbent and rooting at lower nodes, 30–90(–150) cm tall. Leaf sheaths compressed, keeled, usually glabrous; leaf blades linear or linear-lanceolate, 10–40 × 0.4–1.2 cm, usually glabrous, base subrounded, margins scabrous, apex acuminate; ligule 0.5–1 mm. Inflorescence of 2–5(–8) racemes, subdigitate or on a short axis; racemes 3–10 cm, ascending to widely spreading; spikelets usually single, overlapping in 2 rows, sometimes paired especially in middle of raceme; rachis ribbonlike, 1.5–3 mm wide, margins scabrous. Spikelets green becoming brown, suborbicular, ovate or broadly elliptic, 2–3 mm, glabrous, obtuse to apiculate; upper glume membranous, 3–7-veined; lower lemma membranous or sometimes indurate, 3–5(–7)-veined; upper lemma brown at maturity, subequaling spikelet, coriaceous, finely striate, obtuse. Fl. and fr. May–Nov. 2n = 20, 40 or 60.
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Flora of China Vol. 22: 526, 528, 529, 530 in eFloras.org, Missouri Botanical Garden. Accessed Nov 12, 2008.
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Flora of China @ eFloras.org
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Description

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Perennial; culms 10-150 cm high, 1-6 mm in diameter, 2-17-noded, the nodes commonly exposed, erect, or ascending from a procumbent base and rooting at the lower nodes. Leaf-blades linear, 5-40 cm long, 3-15 mm wide, tapering to a filiform tip. Inflorescence composed of 1-20 racemes, these digitate or borne on an axis up to 8 cm long, the lowest raceme 4-15 cm long, with spikelets borne singly on a ribbon-like rhachis 1-2.5 mm wide. Spikelets broadly elliptic, obovate or suborbicular, 1.4-3 mm long, green, becoming brown; lower glume absent; upper glume papery; lower lemma similar or rarely coriaceous, 3-5-nerved (in the latter case the nerves evenly spaced or the laterals close together); upper lemma finely striate, brown at maturity.
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Flora of Pakistan Vol. 0: 213 in eFloras.org, Missouri Botanical Garden. Accessed Nov 12, 2008.
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Flora of Pakistan @ eFloras.org
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S. I. Ali & M. Qaiser
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Distribution

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S.E. Asia to Polynesia & Australia.
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Annotated Checklist of the Flowering Plants of Nepal Vol. 0 in eFloras.org, Missouri Botanical Garden. Accessed Nov 12, 2008.
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Annotated Checklist of the Flowering Plants of Nepal @ eFloras.org
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Distribution

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Tropical Asia and Pacific islands. Taiwan, common in tea plantation, roadsides, dry fields, grasslands and waste places.
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Gramineae (Poaceae) in Flora of Taiwan Vol. 0 in eFloras.org, Missouri Botanical Garden. Accessed Nov 12, 2008.
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Distribution

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Distribution: Pakistan (Sind); throughout the Old World tropics.
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Flora of Pakistan Vol. 0: 213 in eFloras.org, Missouri Botanical Garden. Accessed Nov 12, 2008.
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Flora of Pakistan @ eFloras.org
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S. I. Ali & M. Qaiser
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Elevation Range

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200-2200 m
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Annotated Checklist of the Flowering Plants of Nepal Vol. 0 in eFloras.org, Missouri Botanical Garden. Accessed Nov 12, 2008.
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Annotated Checklist of the Flowering Plants of Nepal @ eFloras.org
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Flower/Fruit

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Fl. & Fr. Per.: April-May and again October-December.
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Flora of Pakistan Vol. 0: 213 in eFloras.org, Missouri Botanical Garden. Accessed Nov 12, 2008.
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Flora of Pakistan @ eFloras.org
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S. I. Ali & M. Qaiser
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Habitat & Distribution

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Roadsides, weedy places, often on damp soils. Fujian, Guangdong, Guangxi, Guizhou, Hainan, Hubei, Jiangsu, Jiangxi, Sichuan, Taiwan, Yunnan, Zhejiang [tropics and subtropics of the Old World; introduced in America].
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Missouri Botanical Garden, 4344 Shaw Boulevard, St. Louis, MO, 63110 USA
bibliographic citation
Flora of China Vol. 22: 526, 528, 529, 530 in eFloras.org, Missouri Botanical Garden. Accessed Nov 12, 2008.
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Flora of China @ eFloras.org
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Wu Zhengyi, Peter H. Raven & Hong Deyuan
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Derivation of specific name

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scrobiculatum: minutely pitted
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Mark Hyde, Bart Wursten and Petra Ballings
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Hyde, M.A., Wursten, B.T. and Ballings, P. (2002-2014). Paspalum scrobiculatum L. Flora of Zimbabwe website. Accessed 28 August 2014 at http://www.zimbabweflora.co.zw/speciesdata/species.php?species_id=107130
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Mark Hyde
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Bart Wursten
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Petra Ballings
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Description

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Perennial tufted grass, up to 1.5 m, but usually much smaller. Leaves light green, often tinged with purple. Inflorescence mostly digitate with 2 or 3 racemes, but occasionally more. Spikelets almost round, glabrous, in 2 overlapping rows along the thickened rhachis
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Mark Hyde, Bart Wursten and Petra Ballings
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Hyde, M.A., Wursten, B.T. and Ballings, P. (2002-2014). Paspalum scrobiculatum L. Flora of Zimbabwe website. Accessed 28 August 2014 at http://www.zimbabweflora.co.zw/speciesdata/species.php?species_id=107130
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Mark Hyde
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Bart Wursten
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Frequency

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Common
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Mark Hyde, Bart Wursten and Petra Ballings
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Hyde, M.A., Wursten, B.T. and Ballings, P. (2002-2014). Paspalum scrobiculatum L. Flora of Zimbabwe website. Accessed 28 August 2014 at http://www.zimbabweflora.co.zw/speciesdata/species.php?species_id=107130
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Comprehensive Description

provided by North American Flora
Paspalum boscianum Fliigge, Gram. Monog. 170. 1810
Paspalum virgatum Walt. Fl. Car. 75. 1788. Not P. virgatum L. 1759. Paspalum brunneum Eosc; Fliigge, Gram. Monog. 171, as synonym. 1810. Paspalum purpurascens Ell. Bot. S. C. & Ga. 1 : 108. 1816. ? Paspalum compressum Raf . Fl. I^udov. 15. 1817. Paspalum conferlum Le Conte, Jour, de Phys. 91 : 285. 1820. A rather stout glabrous perennial with compressed stems, which often root at the lower nodes, flat leaf-blades and glabrous spikelets. Stems often branched, 5-12 dm. long; leafsheaths compressed, glabrous or the basal ones papillose-hirsute; blades 3 dm. long or less, 310 mm. wide, linear, flat, of medium texture, smooth or roughish, papillose-hirsute above near the base, otherwise glabrous; racemes 2-13, spreading or ascending, 4-9 cm. long, the rachis straight, 2-2.5 mm. wide; spikelets in pairs and often so crowded as to appear in 4 rows, redbrown, 2-2.3 mm. long, 1.5-1.8 mm. wide, broadly obovoid, glabrous, the first scale wanting, the second 5-nerved, the third 3-nerved, the fruiting scale deep-brown at maturity.
Type locality : [South] Carolina.
Distribution : Virginia and Tennessee to Florida and Texas ; Porto Rico.
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bibliographic citation
George Valentine Nash. 1912. (POALES); POACEAE (pars). North American flora. vol 17(2). New York Botanical Garden, New York, NY
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Comprehensive Description

provided by North American Flora
Paspalum scrobiculatum L. Mant. 29. 1767
A tufted smooth and glabrous perennial, with fiat leaf-blades and glabrous spikelets. Stems 2-6 dm. tall, sometimes branched; leaf -sheaths compressed; blades 2 dm. long or less, 2-8 mm. wide, linear, erect, or nearly so, flat, of medium texture; racemes 2-5, 2-7 cm. long, the rachis 1.5-2 mm. wide; spikelets singly disposed, 2-2.5 mm. long and 1.5-1.75 mm. wide, oval, glabrous, the first scale wanting, the second and third scales 5-nerved, the lateral nerves approximate and distant from the midnerve, the fruiting scale deep seal-brown at maturity.
Type locality : Kast Indies.
Distribution ; Florida ; tropics of the Old World.
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George Valentine Nash. 1912. (POALES); POACEAE (pars). North American flora. vol 17(2). New York Botanical Garden, New York, NY
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Physical Description

provided by USDA PLANTS text
Annuals, Aquatic, leaves emergent, Terrestrial, not aquatic, Stems trailing, spreading or prostrate, Stems nodes swollen or brittle, Stems erect or ascending, Stems geniculate, decumbent, or lax, sometimes rooting at nodes, Stems terete, round in cross section, or polygonal, Stem internodes solid or spongy, Stem internodes hollow, Stems with inflorescence less than 1 m tall, Stems, culms, or scapes exceeding basal leaves, Leaves mostly cauline, Leaves conspicuously 2-ranked, distichous, Leaves sheathing at base, Leaf sheath mostly open, or loose, Leaf sheath smooth, glabrous, Leaf sheath and blade differentiated, Leaf blades linear, Leaf blades 2-10 mm wide, Leaf blades 1-2 cm wide, Leaf blades mostly flat, Leaf blades mostly glabrous, Leaf blades more or less hairy, Leaf blades scabrous, roughened, or wrinkled, Ligule present, Lig ule an unfringed eciliate membrane, Inflorescence terminal, Inflorescence solitary, with 1 spike, fascicle, glomerule, head, or cluster per stem or culm, Inflorescence a panicle with narrowly racemose or spicate branches, Inflorescence with 2-10 branches, Inflorescence branches 1-sided, Rachis dilated, flat, central axis to which spikelets are attached, Flowers bisexual, Spikelets pedicellate, Spikelets dorsally compressed or terete, Spikelet less than 3 mm wide, Spikelets with 1 fertile floret, Spikelets with 2 florets, Spikelet with 1 fertile floret and 1-2 sterile florets, Spikelets paired at rachis nodes, Spikelets all alike and fertille, Spikelets bisexual, Spikelets disarticulating below the glumes, Spikelets secund, in rows on one side of rachis, Rachilla or pedicel glabrous, Glumes present, empty bracts, Glumes 2 clearly present, Glumes distinctly unequal, Glumes equal to or longer than adjacent lemma, Glume equal to or longer than spikelet, Glumes 4-7 nerved, Lemma similar in texture to glumes, Lemma 3 nerved, Lemma glabrous, Lemma apex truncate, rounded, or obtuse, Lemma awnless, Lemma straight, Palea present, well developed, Palea shorter than lemma, Stamens 3, Styles 2-fid, deeply 2-branched, Stigmas 2, Fruit - caryopsis, Caryopsis white.
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Paspalum scrobiculatum

provided by wikipedia EN

Paspalum scrobiculatum, commonly called Kodo millet or Koda millet,[1][2][3] is an annual grain that is grown primarily in Nepal (not to be confused with Ragi (Finger millet, Eleusine coracana))[4][5] and also in India, Philippines, Indonesia, Vietnam, Thailand, and in West Africa from where it originated. It is grown as a minor crop in most of these areas, with the exception of the Deccan plateau in India where it is grown as a major food source.[6] It is a very hardy crop that is drought tolerant and can survive on marginal soils where other crops may not survive, and can supply 450–900 kg of grain per hectare.[7] Kodo millet has large potential to provide nourishing food to subsistence farmers in Africa and elsewhere.

The plant is called Arikelu in the Telugu language, Varagu in Tamil, Varak (വരക്) in Malayalam, Arka in Kannada,Kodo in Hindi and Kodra in Punjabi

Description

Kodo millet is a monocot and an annual grass that grows to heights of approximately four feet.[8] It has an inflorescence that produces 4-6 racemes that are 4–9 cm long. Its slender, light green leaves grow to be 20 to 40 centimeters in length. The seeds it produces are very small and ellipsoidal, being approximately 1.5 mm in width and 2 mm in length; they vary in colour from being light brown to a dark grey. Kodo millet has a shallow root system which may be ideal for intercropping.[7]

History, geography, and ethnography

Infloresences

Paspalum scrobiculatum var. scrobiculatum is grown in India as an important crop, while Paspalum scrobiculatum var. commersonii is the wild variety indigenous to Africa.[7] The kodo millet, also known as cow grass, rice grass, ditch millet, Native Paspalum, or Indian Crown Grass originates in tropical Africa, and it is estimated to have been domesticated in India 3000 years ago.[9] The domestication process is still ongoing. In southern India, it is called varaku or koovaraku. Kodo is probably a corrupt form of kodra, a Hindi name of the plant. It is grown as an annual. It is a minor food crop eaten in many Asian countries, primarily in India where in some regions it is extremely important. It grows wild as a perennial in the west of Africa, where it is eaten as a famine food.[10] Often it grows as a weed in rice fields. Many farmers do not mind it, as it can be harvested as an alternative crop if their primary crop fails.[10] In the Southern United States and Hawaii, it is considered to be a noxious weed.[11]

Growing conditions

Kodo millet is propagated from seed, ideally in row planting instead of broadcast sowing. Its preferred soil type is a very fertile, clay-based soil. Var. scrobiculatum is better suited to dried conditions than its wild counterpart, which requires approximately 800–1200 mm of water annually and is well suited to sub-humid aridity conditions.[12][7] With very low competition from other plants or weeds for nutrients, it can grow well in poor-nutrient soils. However, it does best in soils supplemented with a general fertilizer.[7] The recommended dose for optimal growth is 40 kg of nitrogen plus 20 kg of phosphorus per hectare. A case study in India's Rewa district in 1997 showed a 72% increase in kodo millet grain yields as opposed to no fertilizer. Lodging issues may accompany this. (Please see section "Other farming issues").[13] Kodo millet prefers full light for optimal growth, but can tolerate some partial shading. Its ideal temperature for growth is 25-27 °C. It requires four months until maturity and harvesting.[7]

Other farming issues

The kodo millet is prone to lodging at maturity, causing loss of grain.[13] To prevent this, limited fertilization is recommended. While plenty of fertilizer dramatically improves yields, there is the risk of lodging accompanying vigorous growth. A good balance is applying 14–22 kg of nitrogen. Lodging also occurs due to heavy rains.[14] Kodo millet is harvested by cutting the stalk of the grass and allowing it to dry in the sun for a day or two. It is then ground to remove the husk. Weather dependency is a major issue related to proper harvesting and storage. Additionally, threshing on roads damages the grains, and husking is a very time-consuming process. Kodo millets are believed by farmers to be the toughest grain to de-husk.[15]

Stress tolerance

The kodo millet can survive well on marginal soils; var. scrobiculatum requires very little water in order to grow, and thus has very good drought tolerance.[7] It can be cultivated without an irrigation system. Farmyard manures provide adequate nutrients in terms of adding fertilizer, but kodo millets can still survive on low-nutrient soils. The wild variety is better suited to wetter conditions, and can tolerate flooded areas and swampy ground.[7]

Major weeds, pests and diseases

Paspalum ergot is a fungal disease to which kodo millet is susceptible. Hardened masses of this fungus, called sclerotia, will grow in place of the millet grain.[7] These compact fungi growths contain a chemical compound that is poisonous to humans and livestock if consumed, and potentially fatal. It causes damage to the central nervous system, causing excitability in animals and eventually loss of muscle control. If the symptoms are caught early and the animals are removed from the infected food, they have a good chance of recovery. Cleaning the seeds by winnowing them before storage may remove the fungal spores.[7]

Insect pests include:[16]

Shoot and stem feeders
Leaf feeders
Sucking pests
Panicle pests

Consumption and uses

In India, kodo millet is ground into flour and used to make pudding.[7] In Africa it is cooked like rice. It is also a good choice for animal fodder for cattle, goats, pigs, sheep, and poultry.[11] In Hawaii, var. scrobiculatum is found to grow well on hillside slopes where other grasses do not flourish. It has the potential to be grown as a food source on hillside farms.[11] It may also have potential to be used as grass ties on hillside plots to prevent soil erosion, while also providing a famine food as a secondary purpose. It has been noted that it makes a good cover crop.[7]

Nutritional information

Kodo millet is a nutritious grain and a good substitute to rice or wheat. The grain is composed of 11% of protein, providing 9 grams/100 g consumed.[18] It is an excellent source of fibre at 10 grams (37-38%), as opposed to rice, which provides 0.2/100 g, and wheat, which provides 1.2/100 g. An adequate fibre source helps combat the feeling of hunger. Kodo millet contains 66.6 g of carbohydrates and 353 kcal per 100 g of grain, comparable to other millets. It also contains 3.6 g of fat per 100 g. It provides minimal amounts of iron, at 0.5/100 mg, and minimal amounts of calcium, and 27/100 mg.[18] Kodo millets also contain high amounts of polyphenols, an antioxidant compound.[19]

Practical information

If fertilizer is available, supplemented nitrogen and phosphorus in limited amounts can increase yields dramatically. Before applying fertilizer, soil pH tests should be used to make sure it ideal for proper nutrient uptake. While pH levels can vary across fields, a few may be able to appropriate a rough guess. If pH levels are not suitable, fertilizer cannot be taken up by plants and will be wasted. Planting kodo millet in rows, instead of broadcasting the seeds, will increase yields and make weeding easier. Kodo millet will grow on marginal soils, but only if it has little competition from weeds.[19] Finally, proper cleaning of the seeds by wind winnowing will help prevent the sclerotia of fungal diseases being consumed accidentally.[7] Kodo millet seeds can be obtained from the International Crop Research Institute for the Semi-Arid Tropics [20]

References

  1. ^ A. E. Grant (1898), "Poisonous Koda millet". Letter to Nature, volume 57, page 271.
  2. ^ Harry Nelson Vinall(1917), Foxtail Millet: Its Culture and Utilization in the United States. Issue 793 of Farmers' bulletin, U.S. Department of Agriculture. 28 pages.
  3. ^ Sabelli, Paolo A.; Larkins, Brian A. (2009). "The Development of Endosperm in Grasses". Plant Physiology. American Society of Plant Biologists (ASPB). 149 (1): 14–26. doi:10.1104/pp.108.129437. ISSN 0032-0889. PMC 2613697. PMID 19126691.
  4. ^ Bastola, Biswash Raj; Pandey, M.P.; Ojha, B.R.; Ghimire, S.K.; Baral, K. (2015-06-25). "Phenotypic Diversity of Nepalese Finger Millet (Eleusine coracana (L.) Gaertn.) Accessions at IAAS, Rampur, Nepal". International Journal of Applied Sciences and Biotechnology. 3 (2): 285–290. doi:10.3126/ijasbt.v3i2.12413. ISSN 2091-2609.
  5. ^ LI-BIRD (2017). "Released and promising crop varieties for mountain agriculture in Nepal" (PDF).
  6. ^ |"Millets". Earth360. (2010-13). http://earth360.in/web/Millets.html
  7. ^ a b c d e f g h i j k l m Heuzé V., Tran G., Giger-Reverdin S., 2015. Scrobic (Paspalum scrobiculatum) forage and grain. Feedipedia, a programme by INRA, CIRAD, AFZ and FAO. https://www.feedipedia.org/node/401 Last updated on October 6, 2015, 12:07
  8. ^ "Kodomillet". United States Department of Agriculture. (No date given, accessed November 11, 2013). http://plants.usda.gov/core/profile?symbol=Pasc6
  9. ^ "Kodo millet". International Crop Research Institute for the Semi-Arid Tropics. (December 4, 2013). http://www.icrisat.org/crop-kodomillet.htm Archived 2013-12-11 at the Wayback Machine
  10. ^ a b Board on Science and Technology for International Development, Office of International Affairs, National Research Council. "Kodo Millet". Lost Crops of Africa; Volume 1: Grains. (1996). http://books.nap.edu/openbook.php?record_id=2305&page=249
  11. ^ a b c "Paspalum scrobiculatum (grass)." Global Invasive Species Database. (2010). http://www.issg.org/database/species/ecology.asp?si=1423&lang=EN
  12. ^ "Agroclimatic Zones". Production Estimates and Crop Assessment Division Foreign Agricultural Service. (2013). http://www.fas.usda.gov/pecad2/highlights/2002/10/ethiopia/baseline/Eth_Agroeco_Zones.htm Archived 2013-12-14 at the Wayback Machine
  13. ^ a b J. M. J. de Wet, K. E. Prasada Rao, M. H. Mengesha and D. E. Brink. "Diversity in Kodo Millet". New York Botanical Garden Press. (1983). JSTOR 4254476
  14. ^ Johns, M. "Millet for Forage Use: Frequently asked Questions". Alberta Agriculture and Rural Development. (2007). http://www1.agric.gov.ab.ca/$department/deptdocs.nsf/all/faq8355
  15. ^ "Report on Survey of Post-Harvest Technology and Constraints Faced by Women Farmers related to Small Millets and Associated Crops". DHAN Foundation. (2011). http://www.dhan.org/smallmillets/docs/report/PHT_final_report.pdf
  16. ^ Kalaisekar, A (2017). Insect pests of millets: systematics, bionomics, and management. London: Elsevier. ISBN 978-0-12-804243-4. OCLC 967265246.
  17. ^ Kalaisekar, A.; Padmaja, P.G.; Bhagwat, V.R.; Patil, J.V. (2017). Insect Pests of Millets: Systematics, Bionomics, and Management. Academic Press. ISBN 978-0-12-804243-4.
  18. ^ a b "Millets: Future of Food & Farming". Millet Network of India. (No date given, accessed November 13th 2013.) http://www.swaraj.org/shikshantar/millets.pdf
  19. ^ a b Hedge, P.S.; Chandra, T.S. (2005). "ESR spectroscopic study reveals higher free radical quenching potential in kodo millet (Paspalum scrobiculatum) compared to other millets". Food Chemistry. 92: 177–182. doi:10.1016/j.foodchem.2004.08.002.
  20. ^ "|| ICRISAT || Genebank Home". Archived from the original on 2013-12-11. Retrieved 2013-12-04.

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Paspalum scrobiculatum: Brief Summary

provided by wikipedia EN

Paspalum scrobiculatum, commonly called Kodo millet or Koda millet, is an annual grain that is grown primarily in Nepal (not to be confused with Ragi (Finger millet, Eleusine coracana)) and also in India, Philippines, Indonesia, Vietnam, Thailand, and in West Africa from where it originated. It is grown as a minor crop in most of these areas, with the exception of the Deccan plateau in India where it is grown as a major food source. It is a very hardy crop that is drought tolerant and can survive on marginal soils where other crops may not survive, and can supply 450–900 kg of grain per hectare. Kodo millet has large potential to provide nourishing food to subsistence farmers in Africa and elsewhere.

The plant is called Arikelu in the Telugu language, Varagu in Tamil, Varak (വരക്) in Malayalam, Arka in Kannada,Kodo in Hindi and Kodra in Punjabi

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