More info for the terms:
cone,
layering,
litter,
seed,
tree,
tussockCone and seed production: Cone production begins at about 15 years of
age for open-grown trees and 35 to 40 years of age for trees in
well-stocked stands [
11]. Large quantities of seed are usually not
produced until trees are at least 40 years old. Fifty- to 150-year-old
open-grown trees produce the best cone crops, with individual trees
sometimes producing as many as 20,000 cones in a good year [
11]. Good
seed crops are produced every 3 to 6 years, with some seed produced in
intervening years. Tamarack seeds are about 0.12 inch (3 mm) long and
have a wing about 0.25 inch (6 mm) long [
26].
Dispersal: Seeds are dispersed in the fall over a relatively short time
period. In a Minnesota study, dispersal began about September 1, and by
October 31 about 98 percent of seed had fallen [
11]. The remaining seed
fell throughout the winter. In interior Alaska, 95 percent of tamarack
seeds are shed by November [
4]. Tamarack seeds are primarily wind
dispersed, but red squirrels disperse some seed. Most wind-dispersed
seeds fall within a distance of two tree heights, but a small percentage
travels greater distances [
4,
11].
Seed destruction and predation: In Minnesota, small mammals, presumably
mice, voles, and shrews, consume large quantities of tamarack seed off
the ground and can destroy up to one-half of a tamarack seed crop [
11].
Also, seeds on the ground are susceptible to infections from bacteria
and fungi. Consequently, only about 4 or 5 percent of tamarack seed
that reaches the ground germinates [
26]. In tamarack stands in New
Brunswick, insects destroyed between 25 and 88 percent of seed produced.
Larvae of the spruce budworm and the cone maggot were responsible for
greatest loss [
53].
Viability: Tamarack seed remains viable for only about 1 year after
dispersal [
11]. Typically a large percentage of tamarack seed is
unfilled. In Minnesota about one-third, and in northern Ontario about
one-half of seed had undeveloped embryo and endosperm [
11,
15]. At the
northern portion of the species range in the Northwest Territories,
tamarack produced a limited amount of seed, but none was viable [
12].
Germination and establishment: Germinative capacity ranges from about
30 to 60 percent [
11]. Neither light nor pH appear to influence
germination appreciably [
11,
15]. Tamarack seeds require a moist but
unsaturated substrate for germination. The best seedbed is warm, moist
mineral or organic soil free from competing vegetation [
26].
Slow-growing sphagnum mosses also provide a good seedbed, as they have a
tendency to remain moist. In open swampy habitats, tamarack seedlings
are often found on sphagnum mosses [
1,
7]. Feather mosses are usually
poor seedbeds because they tend to dry out, but if they remain moist,
they can provide a favorable seedbed [
4]. On poorly drained river
terraces in interior Alaska, tamarack seedlings are more abundant on
sphagnum and feather mosses than on sedge tussock tops, troughs between
the tussocks, or litter-covered sites; on well-drained river terraces,
seedlings are primarily restricted to mineral soil [
4]. Tamarack
seedlings are intolerant of shade and flooding. Seedlings may survive a
few years in shade, but most will die unless released. Partial water
submersion for 1 to 3 weeks kills 1st-year seedlings [
11]. In full
sunlight seedlings grow relatively rapidly, reaching heights of 7 to 9
inches (18-23 cm) after one growing season, and 18 to 25 inches (46-51
cm) tall after three [
26]. Roots of seedlings growing in nearly full
sunlight may reach depths of 2.5 to 3.5 inches (5-11 cm) after one
growing season, while over the same time period roots of seedlings
growing in shade reach depths of only about 1 inch (2.5 cm) [
11].
Vegetative reproduction: Layering is a dominant mode of reproduction at
the northern limit of the species' range. In the southern part of its
range, layering is uncommon but does occur when lower branches become
covered with litter or fast-growing mosses [
26]. Curtis [
7] reported
that tamarack has the unusual ability to produce root sprouts up to 30
feet (9 m) away from a mother tree.