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Conservation Status

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Species habitat is threatened by hydroelectric dams, untreated organic waste, and other disturbances
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Cyclicity

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Adults emerge from May to late June.
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Distribution

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In Alberta, this species is known from the Saskatchewan River system. In North America, it ranges from Quebec and Nova Scotia through New England, south to Georgia, and west to Oklahoma, Saskatchewan, and Alberta.
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General Description

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The genus is characterized by the absence of an anal lobe on the hindwings. Males (length = 6.5 mm) have the ninth abdominal segment produced ventrad; this segment is rounded and hairy behind, with the penis lying beneath. Curved chitinous processes lie on each side of the penis, and are joined at their posterior end to form a blunt prong-like tip that may only be apparent in cleared mounts. The male epiproct is rounded. Females (length = 9.0 mm) have the subgenital plate produced over most of the ninth sternum. The subgenital plate is triangular and rounded at the tip. Nymphal hindwing pads are subparallel to the axis of the body and the inner wing pad margins are straight. Mature final-instar nymphs are approximately 7 mm long.
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Habitat

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Larger, warmer rivers, and streams.
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Life Cycle

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Adult emergence occurs in spring, with the time of egg laying and hatching being relatively short. Nymphs can be found in fall (October), and grow rapidly during fall and early winter, but slowly from December to May. Growth resumes in May, but is not closely synchronized because nymphs in various stages of maturity occur in stream benthic samples and emergence is extended over a few weeks.
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Trophic Strategy

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Although Hitchcock (1974) stated that nymphs are herbivorous, Harper and Magnin (1969) contended that nymphs are at least partly carnivorous.
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