^Don J. Brenner; Noel R. Krieg; James T. Staley. George M. Garrity, 编. The Proteobacteria. Bergey's Manual of Systematic Bacteriology 2C 2nd. New York: Springer. July 26, 2005: 1388 [1984(Williams & Wilkins)]. ISBN 978-0-387-24145-6. British Library no. GBA561951.